277 lines
35 KiB
XML
277 lines
35 KiB
XML
<document id="0B6F4F6EA591095C3B38B3E5E73C2003" ID-DOI="10.11646/zootaxa.5481.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="12742760" ID-ZooBank="9ECB8E6A-5B6F-451F-AA95-7084B79AFE6B0" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721033546162" checkinUser="plazi" docAuthor="Hájek, Jiří & Sheth, Sayali D." docDate="2024" docId="03D187BD06254919FF27244FAC7DDAD7" docLanguage="en" docName="zootaxa.5481.1.7.pdf" docOrigin="Zootaxa 5481 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5481.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Copelatus kopetzi Hájek & Sheth, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="126" masterDocId="FFE8FFC50623491EFFB0213DA937DC4E" masterDocTitle="A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="130" masterPageNumber="119" pageNumber="125" updateTime="1721034136821" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="6C1DCC74BD2705F14861002BFE8FDD93">
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<mods:title id="1EBBCF4387ED0429EF528B0297352B37">A review of the Oriental Copelatus latipes complex, with the description of two new species (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:name id="5264466CBF642AB5D4CD05E542F3F6EB" type="personal">
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<mods:roleTerm id="F3A85C73AE9BD7EADAC01AE8A329FA56">Author</mods:roleTerm>
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<mods:namePart id="21B1F88DA30A33B29658889E425F2116">Hájek, Jiří</mods:namePart>
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<mods:affiliation id="28CB6948CED12B6949B87C9BD3A321C8">Department of Entomology, National Museum, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Poc ̌ ernice, Czech Republic</mods:affiliation>
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</mods:name>
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<mods:name id="E21AA52AFFFE667FA8D041E2739B6A9A" type="personal">
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<mods:roleTerm id="C03C46D238BD078852CBCF8F6629BC04">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="4DB9A8D09C23702FFE5D924B6BE67B72">Sheth, Sayali D.</mods:namePart>
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<mods:affiliation id="9375FA69E05F352D20A5AC4FA24DB01E">Department of Zoology, Prof. Ramkrishna More Arts, Commerce & Science College, Akurdi, Pune - 411044, Maharashtra, India</mods:affiliation>
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<mods:typeOfResource id="36DDDBCDC600AA0D33540A9C2C7A63CB">text</mods:typeOfResource>
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<mods:title id="9C9C475530A2B1766BC6F0F2BB181F71">Zootaxa</mods:title>
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<mods:part id="BB72A152A83654BB73CBE1EE62653C86">
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<mods:date id="2381C2BBA12B806F2BD3EF4EC4DE3A54">2024</mods:date>
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<mods:number id="36B60410CF86E6CF1B7E15C3724AB92A">2024-07-15</mods:number>
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<mods:number id="51C61A1A1FDD6F01C5C99E6879A6D0A4">5481</mods:number>
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<mods:start id="D8BE259994B419B776310665D4A9EFF4">119</mods:start>
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<mods:url id="64A20BF42D71217E8C9A81FC24759E84">http://dx.doi.org/10.11646/zootaxa.5481.1.7</mods:url>
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<mods:classification id="734C6EAD48F192462B070D2D3211AA71">journal article</mods:classification>
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<mods:identifier id="6A95875973210E49339677CF0B7EC55E" type="DOI">10.11646/zootaxa.5481.1.7</mods:identifier>
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<mods:identifier id="A13AB2F63AD004F7D3B9343C506C1B89" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D187BD06254919FF27244FAC7DDAD7" ID-DOI="http://doi.org/10.5281/zenodo.12742801" ID-Zenodo-Dep="12742801" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7" httpUri="http://treatment.plazi.org/id/03D187BD06254919FF27244FAC7DDAD7" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125">
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<subSubSection id="C362652006254918FF27244FA8E6D9C3" box="[151,465,1394,1421]" pageId="6" pageNumber="125" type="nomenclature">
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<paragraph id="8BC736AB06254918FF27244FA8E6D9C3" blockId="6.[151,465,1394,1457]" box="[151,465,1394,1421]" pageId="6" pageNumber="125">
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<heading id="D08F81C706254918FF27244FA8E6D9C3" bold="true" box="[151,465,1394,1421]" fontSize="11" level="1" pageId="6" pageNumber="125" reason="1">
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<emphasis id="B90CEAB906254918FF27244FA8E6D9C3" bold="true" box="[151,465,1394,1421]" pageId="6" pageNumber="125">
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<taxonomicName id="4C784D2806254918FF27244FA85BD9C3" authorityName="Hájek & Sheth" authorityYear="2024" box="[151,364,1394,1421]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
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<emphasis id="B90CEAB906254918FF27244FA85BD9C3" bold="true" box="[151,364,1394,1421]" italics="true" pageId="6" pageNumber="125">Copelatus kopetzi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23F57C206254918FEC3244EA8E6D9C3" box="[371,465,1395,1421]" pageId="6" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C362652006254918FF2724AAA874D9FF" box="[151,323,1431,1457]" pageId="6" pageNumber="125" type="description">
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<paragraph id="8BC736AB06254918FF2724AAA874D9FF" blockId="6.[151,465,1394,1457]" box="[151,323,1431,1457]" pageId="6" pageNumber="125">
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(
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||
<figureCitation id="13432A2E06254918FF2F24AAA9DFD9FF" box="[159,232,1431,1457]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="6" pageNumber="125">Figs 4</figureCitation>
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,
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<figureCitation id="13432A2E06254918FF4424AAA80CD9FF" box="[244,315,1431,1457]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="6" pageNumber="125">13–14</figureCitation>
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)
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C362652006254918FF2724E2ADD7DAC7" pageId="6" pageNumber="125" type="materials_examined">
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<paragraph id="8BC736AB06254918FF2724E2AA22DA53" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
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<materialsCitation id="3B103CF606254918FF2724E2AA26DA53" country="Nepal" county="Dsauda river" latitude="28.8975" location="Sudurpashchim Province" longLatPrecision="20" longitude="80.2275" municipality="Shuklaphanta Nature Reserve" pageId="6" pageNumber="125" specimenCount="1" typeStatus="holotype">
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<emphasis id="B90CEAB906254918FF2724E2A80ED9B7" bold="true" box="[151,313,1503,1529]" pageId="6" pageNumber="125">
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<typeStatus id="54C3880906254918FF2724E2A9E5D9B7" box="[151,210,1503,1529]" pageId="6" pageNumber="125">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F36F763B06254918FEF424E2A8BDD9B7" box="[324,394,1503,1529]" name="Nepal" pageId="6" pageNumber="125">Nepal</collectingCountry>
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,
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<location id="8EA7607006254918FE2824E2AB80D9B7" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FE2824E2AB80D9B7" box="[408,695,1503,1529]" country="Nepal" county="Dsauda river" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Shuklaphanta Nature Reserve" name="Sudurpashchim Province" pageId="6" pageNumber="125">Sudurpashchim Province</location>
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,
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<location id="8EA7607006254918FD7624E2AA87D9B7" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FD7624E2AA87D9B7" box="[710,944,1503,1529]" country="Nepal" county="Dsauda river" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Shuklaphanta Nature Reserve" name="Kanchanpur District" pageId="6" pageNumber="125">Kanchanpur District</location>
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, ca.
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<locationDeviation id="9A752F8906254918FC5924E2ADC1D9B7" box="[1001,1270,1503,1529]" location="Bhimdatta" pageId="6" pageNumber="125">
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<quantity id="4C809B4E06254918FC5924E2AD11D9B7" box="[1001,1062,1503,1529]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="125" unit="km" value="9.0">9 km</quantity>
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SE of Bhimdatta
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</locationDeviation>
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,
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<collectingMunicipality id="6BA3ACD106254918FAB424E2A870DA53" pageId="6" pageNumber="125">Shuklaphanta Nature Reserve</collectingMunicipality>
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,
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<collectingCounty id="62A64E2706254918FEE3273EA8D2DA53" box="[339,485,1539,1565]" pageId="6" pageNumber="125">Dsauda river</collectingCounty>
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,
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<geoCoordinate id="EE4C506C06254918FE5F273EAB4ADA50" box="[495,637,1539,1566]" degrees="28" direction="north" minutes="53" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="51" value="28.8975">28°53’51’’N</geoCoordinate>
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<geoCoordinate id="EE4C506C06254918FD34273EAA26DA53" box="[644,785,1539,1565]" degrees="80" direction="east" minutes="13" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="39" value="80.2275">80°13’39’’E</geoCoordinate>
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="8BC736AB06254918FF77271AADD7DAC7" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
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<emphasis id="B90CEAB906254918FF77271AA8B6DA0F" bold="true" box="[199,385,1575,1601]" pageId="6" pageNumber="125">Type material.</emphasis>
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<materialsCitation id="3B103CF606254918FE24271AADD7DAC7" collectingDate="2017-07-01" collectionCode="NMPC" collectorName="A. Kopetz" county="Kanchanpur" determinerName="Sheth" elevation="160" latitude="28.8975" location="Shuklaphanta" longLatPrecision="20" longitude="80.2275" municipality="Mahandranagar" pageId="6" pageNumber="125" specimenCount="2" specimenCount-male="2" stateProvince="Mahakali" typeStatus="holotype">
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<typeStatus id="54C3880906254918FE24271AA8CBDA0F" box="[404,508,1575,1601]" pageId="6" pageNumber="125" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D7EFD2206254918FDBF271AAB13DA0F" box="[527,548,1575,1601]" count="1" pageId="6" pageNumber="125" type="male">♂</specimenCount>
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(
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<collectionCode id="ED69AE6E06254918FD8F271AABA3DA0F" box="[575,660,1575,1601]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="6" pageNumber="125">NMPC</collectionCode>
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), labelled: “NEP:
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<collectingRegion id="49BCF84906254918FC32271AAADCDA0F" box="[898,1003,1575,1601]" country="Nepal" name="Mahakali" pageId="6" pageNumber="125">Mahakali</collectingRegion>
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/
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<collectingCounty id="62A64E2706254918FC44271AAD4ADA0F" box="[1012,1149,1575,1601]" pageId="6" pageNumber="125">Kanchanpur</collectingCounty>
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vic. /
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<collectingMunicipality id="6BA3ACD106254918FB57271AACAEDA0F" box="[1255,1433,1575,1601]" pageId="6" pageNumber="125">Mahandranagar</collectingMunicipality>
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,
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<location id="8EA7607006254918FF272776A818DA2B" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FF272776A818DA2B" box="[151,303,1611,1637]" county="Kanchanpur" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Mahandranagar" name="Shuklaphanta" pageId="6" pageNumber="125" stateProvince="Mahakali">Shuklaphanta</location>
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/ Nature Res.,
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<location id="8EA7607006254918FE792776AB68DA2B" LSID="urn:lsid:plazi:treatment:03D187BD06254919FF27244FAC7DDAD7:8EA7607006254918FE792776AB68DA2B" box="[457,607,1611,1637]" county="Kanchanpur" latitude="28.8975" longLatPrecision="20" longitude="80.2275" municipality="Mahandranagar" name="Dsauda river" pageId="6" pageNumber="125" stateProvince="Mahakali">Dsauda river</location>
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, LFF /
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<geoCoordinate id="EE4C506C06254918FD1D2776AA0EDA28" box="[685,825,1611,1638]" degrees="28" direction="north" minutes="53" orientation="latitude" pageId="6" pageNumber="125" precision="15" seconds="51" value="28.8975">N28°53’51”</geoCoordinate>
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,
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<geoCoordinate id="EE4C506C06254918FCF42776AAFBDA2B" box="[836,972,1611,1637]" degrees="80" direction="east" minutes="13" orientation="longitude" pageId="6" pageNumber="125" precision="15" seconds="39" value="80.2275">E80°13’39”</geoCoordinate>
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,
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<quantity id="4C809B4E06254918FC672776AD20DA2B" box="[983,1047,1611,1637]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="125" unit="m" value="160.0">
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<elevation id="0055D19806254918FC672776AD20DA2B" box="[983,1047,1611,1637]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="6" pageNumber="125" unit="m" value="160.0">160m</elevation>
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</quantity>
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||
/
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<date id="FFC6106B06254918FB9B2776AD95DA2B" box="[1067,1186,1611,1637]" pageId="6" pageNumber="125" value="2017-07-01">
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<collectingDate id="EF82E98306254918FB9B2776AD95DA2B" box="[1067,1186,1611,1637]" pageId="6" pageNumber="125" value="2017-07-01">1.VII.2017</collectingDate>
|
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</date>
|
||
,leg.
|
||
<collectorName id="268D537D06254918FB7F2776AC74DA2B" box="[1231,1347,1611,1637]" pageId="6" pageNumber="125">A.Kopetz</collectorName>
|
||
#17–19 [p] //
|
||
<typeStatus id="54C3880906254918FF652752A84CDAC7" box="[213,379,1647,1673]" pageId="6" pageNumber="125" type="holotype">HOLOTYPUS</typeStatus>
|
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<specimenCount id="9D7EFD2206254918FE332752A8AFDAC7" box="[387,408,1647,1673]" count="1" pageId="6" pageNumber="125" type="male">♂</specimenCount>
|
||
/
|
||
<taxonomicName id="4C784D2806254918FE1E2752AB7EDAC7" authorityName="Erichson" authorityYear="1832" box="[430,585,1647,1673]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B90CEAB906254918FE1E2752AB7EDAC7" box="[430,585,1647,1673]" italics="true" pageId="6" pageNumber="125">COPELATUS</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C784D2806254918FDEE2752AB9ADAC7" authorityName="Hájek & Sheth" authorityYear="2024" box="[606,685,1647,1673]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="125" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
|
||
<emphasis id="B90CEAB906254918FDEE2752AB9ADAC7" box="[606,685,1647,1673]" italics="true" pageId="6" pageNumber="125">kopetzi</emphasis>
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||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23F57C206254918FD042752AA3CDAC7" box="[692,779,1647,1673]" pageId="6" pageNumber="125" rank="species">sp. nov.</taxonomicNameLabel>
|
||
/ Hájek &
|
||
<determinerName id="7E1BDA6F06254918FC372752AAF0DAC7" box="[903,967,1647,1673]" pageId="6" pageNumber="125">Sheth</determinerName>
|
||
det. 2024 [red label, p]”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362652006254919FF7727AEAA6DD91B" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125" type="description">
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<paragraph id="8BC736AB06254918FF7727AEAB6CDA9F" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
|
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<emphasis id="B90CEAB906254918FF7727AEAB61DAE3" bold="true" box="[199,598,1682,1709]" pageId="6" pageNumber="125">
|
||
Description of
|
||
<typeStatus id="54C3880906254918FE3127AEA8DFDAE3" box="[385,488,1683,1709]" pageId="6" pageNumber="125" type="holotype">holotype</typeStatus>
|
||
.
|
||
<emphasis id="B90CEAB906254918FE4A27AFAB61DAE3" bold="true" box="[506,598,1682,1709]" italics="true" pageId="6" pageNumber="125">Habitus</emphasis>
|
||
</emphasis>
|
||
oblong-oval, with continuous outline, broadest at mid-length of elytra, slightly convex. Dorsal surface submatt.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06254918FF7727E7AA13DBE7" blockId="6.[151,1437,1503,2033]" pageId="6" pageNumber="125">
|
||
<emphasis id="B90CEAB906254918FF7727E7A86DDABB" bold="true" box="[199,346,1754,1781]" italics="true" pageId="6" pageNumber="125">Colouration.</emphasis>
|
||
Head reddish brown, blackish around eyes and medially between eyes, lighter on clypeus, labrum and medially on vertex. Pronotum brown blackish, laterally reddish, pale colouration extending anteriorly. Elytra black, with orange apex and sides laterally from elytral stria 6, and orangish pattern consisting of: irregular basal transverse band split into three spots, somewhat extending between striae 1–2 and 4–5; and longitudinal orange stripes between striae 3–4, in posterior fourth between striae 4–5 and in posterior 2/3 between striae 5–6. Ventral surface reddish brown. Appendages orange to red (
|
||
<figureCitation id="13432A2E06254918FD6126B2AA21DBE7" box="[721,790,1935,1961]" captionStart="FIGURES 3–6" captionStartId="3.[152,270,1943,1967]" captionTargetBox="[206,1387,193,1905]" captionTargetId="figure-17@3.[174,1413,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–6. Habitus of Copelatus latipes species complex. 3. C. brivioi (♂; West Bengal, India); 4. C. kopetzi sp. nov. (holotype ♂; Nepal); 5. C. lanxangensis sp. nov. (holotype ♂; Laos); 6. C. latipes (♂; Pahang, Malaysia). Scale bar 2 mm." figureDoi="http://doi.org/10.5281/zenodo.12742764" httpUri="https://zenodo.org/record/12742764/files/figure.png" pageId="6" pageNumber="125">Fig. 4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06254919FF77268FAD68DCB7" blockId="6.[151,1437,1503,2033]" lastBlockId="7.[151,1437,151,1689]" lastPageId="7" lastPageNumber="126" pageId="6" pageNumber="125">
|
||
<emphasis id="B90CEAB906254918FF77268FA83BDB83" bold="true" box="[199,268,1970,1997]" pageId="6" pageNumber="125">
|
||
<emphasis id="B90CEAB906254918FF77268FA832DB83" bold="true" box="[199,261,1970,1997]" italics="true" pageId="6" pageNumber="125">Head</emphasis>
|
||
.
|
||
</emphasis>
|
||
Moderately broad, ca. 0.63× width of pronotum, transversely elliptical. Anterior margin of clypeus slightly concave. Antenna with antennomeres long and slender. Reticulation consisting of fine, well impressed isodiametric polygonal meshes. Punctation double; several large setigerous punctures present in fronto-clypeal depressions, frontal depressions at level of anterior margin of eyes, and in depressions along inner margin of eyes; fine punctures smaller and sparser on clypeus, becoming larger and denser posteriorly.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF77203EAAD1DDE3" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF77203EA874DD53" bold="true" box="[199,323,259,285]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF77203EA80BDD53" bold="true" box="[199,316,259,285]" italics="true" pageId="7" pageNumber="126">Pronotum</emphasis>
|
||
.
|
||
</emphasis>
|
||
Transverse (width/length ratio = 2.9), broadest between posterior angles, lateral margins moderately curved. Lateral sides with beading thin but distinct, except at anterior angles. Reticulation similar to that of head. Punctation similar to that of head; rows of coarse setigerous punctures present along anterior margin, laterally close to sides, several punctures present also in shallow basolateral depressions along basal margin.
|
||
<collectingRegion id="49BCF84906244919FB732052AC39DDC7" box="[1219,1294,367,393]" country="France" name="Centre" pageId="7" pageNumber="126">Centre</collectingRegion>
|
||
of disc with shallowly impressed medial longitudinal smooth line (without reticulation).
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF77208BAB18DEE7" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF77208BA822DD9F" bold="true" box="[199,277,438,465]" italics="true" pageId="7" pageNumber="126">Elytra.</emphasis>
|
||
Elytral striation consisting of six well impressed discal striae and one submarginal stria on each elytron: stria 1 beginning subbasally, striae 2–6 beginning at base of elytra; striae 1 and 4 ending close to apex; striae 2, 3 and 5 shorter than striae 1 and 4—not reaching apex of elytra; stria 6 shortest, ending approximately in apical fifth of elytral length; submarginal stria long, starting at about elytral mid-length and terminating shortly before apex, at same level as dorsal striae 2, 3 and 5. Reticulation similar to that of head and pronotum, but less impressed. Punctation consisting of coarse setigerous punctures present along elytral striae and lateral margins of elytra, and scattered very fine sparse punctures.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF77238EABB2DF5B" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF77238EA832DE83" bold="true" box="[199,261,691,717]" italics="true" pageId="7" pageNumber="126">Legs.</emphasis>
|
||
Protibia modified, angled near base, distinctly broadened anteriorly, club shaped. Pro- and mesotarsomeres 1–3 distinctly broadened, with four rows of adhesive setae on their ventral side; claws simple. Longer spur on metatibia indistinctly sinuate in apical third.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF772223A83BD833" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF772223A860DF77" bold="true" box="[199,343,798,825]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF772223A867DF77" bold="true" box="[199,336,798,825]" italics="true" pageId="7" pageNumber="126">Ventral side</emphasis>
|
||
.
|
||
</emphasis>
|
||
Prosternum sinuate anteriorly, obtusely keeled medially. Prosternal process shortly lanceolate, in cross-section convex, apex obtuse; process distinctly bordered laterally; reticulation not perceptible. Metaventrite with microsculpture consisting of polygonal meshes; lateral parts of metaventrite (‘metasternal wings’) tongueshaped, slender. Metacoxal lines incomplete, absent in ca. anterior fifth of metacoxal length. Metacoxal plates covered with long oblique strioles; reticulation consisting of elongate, oblique, polygonal meshes. Metacoxal processes rounded at posterior margin. Abdominal ventrites I–II with longitudinal strioles; ventrites III–IV with oblique strioles laterally. Tuft of setae present medially on ventrites III–V; ventrite VI with setigerous punctures laterally on either side. Abdominal reticulation consisting of elongate polygonal meshes, longitudinal on ventrites I–II, oblique on ventrite III and transverse on ventrites IV–VI. Punctation consisting of fine, sparsely distributed punctures.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF7725BBAAD1D943" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF7725BBA842D8EE" box="[199,373,1158,1185]" italics="true" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF7725BBA859D8EF" bold="true" box="[199,366,1158,1185]" italics="true" pageId="7" pageNumber="126">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Median lobe of aedeagus in lateral view (
|
||
<figureCitation id="13432A2E06244919FCF525BAAAA2D8EF" box="[837,917,1159,1185]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Fig. 13</figureCitation>
|
||
) broad in basal 2/3; apical third sickle-shaped, evenly curved, subapically narrowed to pointed apex; at 2/3 of its length with two distinct ‚teeth‘ on ventral side— broad blunt proximal one, and bilobed distal one. Parameres moderately broad, ‚D‘ shaped, with distinct lobe on inner margin, apex short and broad; apical lobe long, club-shaped (
|
||
<figureCitation id="13432A2E06244919FC3525CEAAEFD943" box="[901,984,1267,1293]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF77242AA895D97F" blockId="7.[151,1437,151,1689]" box="[199,418,1303,1329]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF77242AA812D97F" bold="true" box="[199,293,1303,1329]" pageId="7" pageNumber="126">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
<paragraph id="8BC736AB06244919FF772406AA6DD91B" blockId="7.[151,1437,151,1689]" box="[199,858,1338,1365]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF772406A849D91B" bold="true" box="[199,382,1339,1365]" pageId="7" pageNumber="126">Measurements.</emphasis>
|
||
TL:
|
||
<quantity id="4C809B4E06244919FE052407AB39D91B" box="[437,526,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="126" unit="mm" value="5.7">5.7 mm</quantity>
|
||
; Tl-h:
|
||
<quantity id="4C809B4E06244919FDE82407AB86D91B" box="[600,689,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="7" pageNumber="126" unit="mm" value="5.1">5.1 mm</quantity>
|
||
; MW:
|
||
<quantity id="4C809B4E06244919FD4D2406AA61D91B" box="[765,854,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="7" pageNumber="126" unit="mm" value="2.9">2.9 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362652006244919FF772463AD3EDA63" pageId="7" pageNumber="126" type="diagnosis">
|
||
<paragraph id="8BC736AB06244919FF772463AD3EDA63" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF772463A8E7D936" bold="true" box="[199,464,1374,1400]" pageId="7" pageNumber="126">Differential diagnosis.</emphasis>
|
||
Based on the presence of six dorsal striae and a submarginal elytral stria,
|
||
<taxonomicName id="4C784D2806244919FA9B2462A9D1D9D3" authorityName="Hájek & Sheth" authorityYear="2024" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="kopetzi" status="sp. nov.">
|
||
<emphasis id="B90CEAB906244919FA9B2462A9D1D9D3" italics="true" pageId="7" pageNumber="126">Copelatus kopetzi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90CEAB906244919FF5D24BEA870D9D3" bold="true" box="[237,327,1411,1437]" pageId="7" pageNumber="126">
|
||
<taxonomicNameLabel id="A23F57C206244919FF5D24BEA870D9D3" box="[237,327,1411,1437]" pageId="7" pageNumber="126" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be classified within the
|
||
<taxonomicName id="4C784D2806244919FD3524BEAA08D9D2" authorityName="Regimbart" authorityYear="1899" box="[645,831,1411,1437]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="irinus">
|
||
<emphasis id="B90CEAB906244919FD3524BEAA08D9D2" box="[645,831,1411,1437]" italics="true" pageId="7" pageNumber="126">Copelatus irinus</emphasis>
|
||
</taxonomicName>
|
||
group
|
||
<emphasis id="B90CEAB906244919FC3E24BEAAFBD9D2" box="[910,972,1411,1436]" italics="true" pageId="7" pageNumber="126">sensu</emphasis>
|
||
<bibRefCitation id="EFE94B5A06244919FC6324BEADB3D9D3" author="Guignot, F." box="[979,1156,1411,1437]" pageId="7" pageNumber="126" pagination="659 - 995" refId="ref6814" refString="Guignot, F. (1961) Revision des hydrocanthares d'Afrique (Coleoptera Dityscoidea [sic!]). Troisieme partie. Annales du Musee Royal du Congo Belge, Serie 8 vo (Sciences Zoologiques), 90, 659 - 995." type="journal article" year="1961">Guignot (1961)</bibRefCitation>
|
||
. The characteristic shape of the median lobe with two ‘teeth’ on the ventral side (
|
||
<figureCitation id="13432A2E06244919FCA9249AAA5CD98F" box="[793,875,1447,1473]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Fig. 13</figureCitation>
|
||
) places the new species into the newly proposed
|
||
<taxonomicName id="4C784D2806244919FF2724F6A833D9AB" authorityName="Sharp" authorityYear="1882" box="[151,260,1483,1509]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="latipes">
|
||
<emphasis id="B90CEAB906244919FF2724F6A833D9AB" box="[151,260,1483,1509]" italics="true" pageId="7" pageNumber="126">C. latipes</emphasis>
|
||
</taxonomicName>
|
||
complex. Within this species complex, the new species is most similar to the sympatric
|
||
<taxonomicName id="4C784D2806244919FB5E24F6AC6CD9AB" authorityName="Rocchi" authorityYear="1976" box="[1262,1371,1483,1509]" class="Insecta" family="Dytiscidae" genus="Copelatus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="brivioi">
|
||
<emphasis id="B90CEAB906244919FB5E24F6AC6CD9AB" box="[1262,1371,1483,1509]" italics="true" pageId="7" pageNumber="126">C. brivioi</emphasis>
|
||
</taxonomicName>
|
||
; both species can be distinguished from each other by the shape of the median lobe of aedeagus—especially by the shape of the apex and the shape of processes on the ventral side (see
|
||
<figureCitation id="13432A2E06244919FCE4272EAACCDA63" box="[852,1019,1555,1581]" captionStart="FIGURES 7–18" captionStartId="5.[152,270,1908,1932]" captionTargetBox="[195,1393,190,1873]" captionTargetId="figure-17@5.[182,1406,181,1884]" captionTargetPageId="5" captionText="FIGURES 7–18. Male genitalia of Copelatus latipes species complex. 7, 9, 11, 13, 15, 17. median lobe in lateral (a) and ventro-lateral (b) view; 8, 10, 12, 14, 16, 18. paramere. 7–8. C. bezdeki (holotype; Maharashtra, India); 9–10. C. schereri (Sri Lanka); 11–12. C. brivioi (paratype; Bangladesh); 13–14. C. kopetzi sp. nov. (holotype; Nepal); 15–16. C. lanxangensis sp. nov. (holotype; Laos); 17–18. C. latipes (Pahang, Malaysia). Scale bar 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.12742766" httpUri="https://zenodo.org/record/12742766/files/figure.png" pageId="7" pageNumber="126">Figs 11 and 13</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362652006244919FF77270AADDCDA3B" pageId="7" pageNumber="126" type="etymology">
|
||
<paragraph id="8BC736AB06244919FF77270AADDCDA3B" blockId="7.[151,1437,151,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF77270AA879DA1F" bold="true" box="[199,334,1591,1617]" pageId="7" pageNumber="126">Etymology.</emphasis>
|
||
The new species is dedicated to its collector, Andreas Kopetz (Amt Wachsenburg,
|
||
<collectingCountry id="F36F763B06244919FB5D270AAC6DDA1F" box="[1261,1370,1591,1617]" name="Germany" pageId="7" pageNumber="126">Germany</collectingCountry>
|
||
), well known specialist on soldier beetles (
|
||
<taxonomicName id="4C784D2806244919FD9C2766AB8DDA3B" authorityName="Imhoff" authorityYear="1856" box="[556,698,1627,1653]" class="Insecta" family="Cantharidae" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="family">Cantharidae</taxonomicName>
|
||
). The specific name is a noun in the genitive case.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C362652006244919FF772743AC7DDAD7" box="[199,1354,1662,1689]" pageId="7" pageNumber="126" type="distribution">
|
||
<paragraph id="8BC736AB06244919FF772743AC7DDAD7" blockId="7.[151,1437,151,1689]" box="[199,1354,1662,1689]" pageId="7" pageNumber="126">
|
||
<emphasis id="B90CEAB906244919FF772743A856DAD6" bold="true" box="[199,353,1662,1688]" pageId="7" pageNumber="126">Distribution.</emphasis>
|
||
The species is so far only known from the
|
||
<typeStatus id="54C3880906244919FCF42742AA43DAD7" box="[836,884,1663,1689]" pageId="7" pageNumber="126">type</typeStatus>
|
||
locality in south-western
|
||
<collectingCountry id="F36F763B06244919FB272742ADEDDAD7" box="[1175,1242,1663,1689]" name="Nepal" pageId="7" pageNumber="126">Nepal</collectingCountry>
|
||
(
|
||
<figureCitation id="13432A2E06244919FB592742AC0BDAD7" box="[1257,1340,1663,1689]" captionStart="FIGURE 19" captionStartId="6.[152,255,1288,1312]" captionTargetBox="[156,1437,273,1261]" captionTargetId="figure-66@6.[151,1437,273,1265]" captionTargetPageId="6" captionText="FIGURE 19. Map of distribution of Copelatus latipes species complex." figureDoi="http://doi.org/10.5281/zenodo.12742768" httpUri="https://zenodo.org/record/12742768/files/figure.png" pageId="7" pageNumber="126">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |