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<mods:title id="E0F58A1DE206E3E1093A54BB96421B06">Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)</mods:title>
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<mods:affiliation id="81B25B877389C81D19B35C4AFBFAC201">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<mods:namePart id="23567F154B64BE31A2D46ABAA14E208D">Mathis, Wayne N.</mods:namePart>
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<mods:affiliation id="778CAC8FA7B70A3C5AFB8176D6A5D504">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143 &amp; Department of Entomology, PO BOX 37012, MRC 169, Smithsonian Institution, Washington, DC 20013 - 7012, USA mathisw @ si. edu; https: // orcid. org / 0000 - 0002 - 5012 - 1762 &amp; Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<emphasis id="B90CEAB33A1A7B1AFF4EFB59558FFB2D" bold="true" box="[151,741,1213,1240]" pageId="73" pageNumber="74">
19.
<taxonomicName id="4C784D223A1A7B1AFF1AFB59558FFB2D" ID-CoL="4V43Q" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" box="[195,741,1213,1240]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A1A7B1AFF1AFB5956DAFB2D" bold="true" box="[195,432,1213,1240]" italics="true" pageId="73" pageNumber="74">Scatophila conifera</emphasis>
Sturtevant and Wheeler
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<paragraph id="8BC736A13A1A7B1AFF4EFB0556BBFB01" blockId="73.[151,741,1213,1276]" box="[151,465,1249,1276]" pageId="73" pageNumber="74">
<figureCitation id="13432A243A1A7B1AFF4EFB055799FB01" box="[151,243,1249,1275]" captionStart="FIGURE 30" captionStartId="22.[151,250,1023,1047]" captionTargetBox="[151,1435,181,999]" captionTargetId="figure-30@22.[151,1435,181,999]" captionTargetPageId="22" captionText="FIGURE 30. Distribution map of Scatophila angusta sp. nov. (circles), S. conifera Sturtevant &amp; Wheeler (triangle), and S. contaminata (Stenhammar) (squares) in Nearctic Region." figureDoi="http://doi.org/10.5281/zenodo.13210791" httpUri="https://zenodo.org/record/13210791/files/figure.png" pageId="73" pageNumber="74">Figs. 30</figureCitation>
,
<figureCitation id="13432A243A1A7B1AFF26FB055641FB06" box="[255,299,1249,1276]" captionStart="FIGURES 156158" captionStartId="68.[151,264,1913,1937]" captionTargetBox="[151,1435,1130,1889]" captionTargetId="figure-70@68.[151,1435,1130,1890]" captionTargetPageId="68" captionText="FIGURES 156158. Head, thorax and wing of Scatophila. 156 Scatophila bipiliaris Sturtevant &amp; Wheeler, head and thorax, dorsal view. 157158. Wing, dorsal view. 157. S. conifera Sturtevant &amp; Wheeler. 158. S. cribrata (Stenhammar). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210887" httpUri="https://zenodo.org/record/13210887/files/figure.png" pageId="73" pageNumber="74">157</figureCitation>
,
<figureCitation id="13432A243A1A7B1AFEEEFB0556F6FB01" box="[311,412,1249,1275]" captionStart-0="FIGURES 168171" captionStart-1="FIGURES 172174" captionStartId-0="74.[151,264,1328,1352]" captionStartId-1="75.[151,264,1078,1102]" captionTargetBox-0="[170,1435,206,1304]" captionTargetBox-1="[159,1427,189,1047]" captionTargetId-0="figure-30@74.[151,1435,181,1304]" captionTargetId-1="figure-27@75.[151,1435,181,1055]" captionTargetPageId-0="74" captionTargetPageId-1="75" captionText-0="FIGURES 168171. Head of Scatophila conifera Sturtevant &amp; Wheeler. 168. Male, latero-oblique view. 169. Male, lateral view. 170. Female, latero-oblique view. 171. Female, lateral view. Scale bar = 0.2 mm." captionText-1="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13210901" figureDoi-1="http://doi.org/10.5281/zenodo.13210903" httpUri-0="https://zenodo.org/record/13210901/files/figure.png" httpUri-1="https://zenodo.org/record/13210903/files/figure.png" pageId="73" pageNumber="74">168174</figureCitation>
,
<figureCitation id="13432A243A1A7B1AFE7EFB0556BBFB01" box="[423,465,1249,1275]" captionStart="FIGURES 175179" captionStartId="76.[151,264,1403,1427]" captionTargetBox="[164,1423,181,1379]" captionTargetId="figure-30@76.[164,1423,181,1379]" captionTargetPageId="76" captionText="FIGURES 175179. Head and thorax of Scatophila. 175177. Head and thorax, dorsal view. 175. Scatophila bisignata Cresson. 176. S. conifera Sturtevant &amp; Wheeler. 177. S. cribrata (Stenhammar). 178179. Head of Scatophila cribrata (Stenhammar). 178. Latero-oblique view. 179. Lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210907" httpUri="https://zenodo.org/record/13210907/files/figure.png" pageId="73" pageNumber="74">176</figureCitation>
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<treatmentCitation id="0AD910B03A1A7B1AFF4EFAFB5583FACD" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[151,745,1311,1335]" page="189" pageId="73" pageNumber="74" year="1954">
<taxonomicName id="4C784D223A1A7B1AFF4EFAFB5583FACD" ID-CoL="4V43Q" authority="Sturtevant and Wheeler 1954: 189" authorityName="Sturtevant and Wheeler" authorityPageNumber="189" authorityYear="1954" box="[151,745,1311,1335]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="73" pageNumber="74" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A1A7B1AFF4EFAFB560FFACD" box="[151,357,1311,1335]" italics="true" pageId="73" pageNumber="74">Scatophila conifera</emphasis>
<bibRefCitation id="EFE94B503A1A7B1AFEAAFAFB5583FACD" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[371,745,1311,1335]" pageId="73" pageNumber="74" pagination="151 - 257" refId="ref70834" refString="Sturtevant, A. H. &amp; Wheeler, M. R. (1954) Synopses of Nearctic Ephydridae (Diptera). Transactions of the American Entomological Society, 79, 151 - 257." type="journal article" year="1954">Sturtevant and Wheeler 1954: 189</bibRefCitation>
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.—
<treatmentCitation id="0AD910B03A1A7B1AFCD0FAFB54D7FACD" author="Wirth, W. W." box="[777,957,1311,1335]" page="758" pageId="73" pageNumber="74" year="1965">
<bibRefCitation id="EFE94B503A1A7B1AFCD0FAFB54D7FACD" author="Wirth, W. W." box="[777,957,1311,1335]" pageId="73" pageNumber="74" pagination="734 - 759" refId="ref70987" refString="Wirth, W. W. (1965) Ephydridae. In: Stone, A., Sabrosky, C. W., Wirth, W. W., Foote, R. W. &amp; Coulson, J. R. (Eds.), A Catalog of the Diptera of America North of Mexico. Handbook 276. U. S. Department of Agriculture, Washington, D. C., pp. 734 - 759." type="book chapter" year="1965">Wirth 1965: 758</bibRefCitation>
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[
<taxonomicName id="4C784D223A1A7B1AFC09FAFB5343FACD" box="[976,1065,1311,1335]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="73" pageNumber="97" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
catalog].—
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<bibRefCitation id="EFE94B503A1A7B1AFB7AFAFB52F6FACD" author="Zatwarnicki, T." box="[1187,1436,1311,1335]" pageId="73" pageNumber="74" pagination="277 - 298" refId="ref71260" refString="Zatwarnicki, T. (1987) New synonyms and records of Palearctic Scatophila (Diptera, Ephydridae). Polskie Pismo Entomologiczne, 57 (2), 277 - 298." type="journal article" year="1987">Zatwarnicki 1987: 291</bibRefCitation>
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[checklist].—
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<bibRefCitation id="EFE94B503A1A7B1AFE94FADA55DDFAAC" author="Zatwarnicki, T. &amp; Mathis, W. N." box="[333,695,1342,1366]" pageId="73" pageNumber="74" pagination="351 - 370" refId="ref71480" refString="Zatwarnicki, T. &amp; Mathis, W. N. (1994) Phylogeny and classification of the genus Scatophila Becker (Diptera: Ephydridae). Annales de la Societe Entomologique de France, 29 (4), 351 - 370. [1993] https: // doi. org / 10.1080 / 21686351.1993.12277698" type="journal article" year="1994">Zatwarnicki and Mathis 1994: 365</bibRefCitation>
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[phylogeny,
<emphasis id="B90CEAB33A1A7B1AFC99FADA54A1FAAC" box="[832,971,1342,1366]" italics="true" pageId="73" pageNumber="74">quadriguttata</emphasis>
group].—
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<bibRefCitation id="EFE94B503A1A7B1AFBEDFADA52F6FAAC" author="Mathis, W. N. &amp; Zatwarnicki, T." box="[1076,1436,1342,1366]" pageId="73" pageNumber="74" pagination="1 - 423" refId="ref70301" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). International Memoirs on Entomology, 4, i - vi + 1 - 423." type="journal article" year="1995">Mathis and Zatwarnicki 1995: 282</bibRefCitation>
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[world catalog].
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</paragraph>
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<subSubSection id="C362652A3A1A7B18FF4EFA7855BEF9C3" lastPageId="75" lastPageNumber="76" pageId="73" pageNumber="74" type="diagnosis">
<paragraph id="8BC736A13A1A7B1AFF4EFA785252FA21" blockId="73.[151,1437,1436,2039]" pageId="73" pageNumber="74">
Diagnosis.—This species is similar to congeners in the
<emphasis id="B90CEAB33A1A7B1AFCDBFA7854F6FA4C" box="[770,924,1436,1462]" italics="true" pageId="73" pageNumber="74">quadriguttata</emphasis>
group but is distinguished from congeners by the following combination of characters: Generally grayish; small shore flies, body length
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.
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<emphasis id="B90CEAB33A1A7B1AFF1EFA00566EFA04" box="[199,260,1508,1534]" italics="true" pageId="73" pageNumber="74">Head</emphasis>
(
<figureCitation id="13432A243A1A7B1AFECBFA0056DDFA04" box="[274,439,1508,1534]" captionStart="FIGURES 168171" captionStartId="74.[151,264,1328,1352]" captionTargetBox="[170,1435,206,1304]" captionTargetId="figure-30@74.[151,1435,181,1304]" captionTargetPageId="74" captionText="FIGURES 168171. Head of Scatophila conifera Sturtevant &amp; Wheeler. 168. Male, latero-oblique view. 169. Male, lateral view. 170. Female, latero-oblique view. 171. Female, lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210901" httpUri="https://zenodo.org/record/13210901/files/figure.png" pageId="73" pageNumber="74">Figs. 168171</figureCitation>
): Mesofrons gray; distance between insertion of lateroclinate seta and medial vertical seta only slightly greater than distance between vertical setae. Face of
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convex, ventral portion projected, somewhat conical, setulae on either side somewhat clustered but not strongly expressed. Gena relatively high, gena-to-eye ratio 0.320.37.
</paragraph>
<paragraph id="8BC736A13A1A7B1AFF1EF99053CFF900" blockId="73.[151,1437,1436,2039]" pageId="73" pageNumber="74">
<emphasis id="B90CEAB33A1A7B1AFF1EF9905672F974" box="[199,280,1652,1678]" italics="true" pageId="73" pageNumber="74">Thorax</emphasis>
(
<figureCitation id="13432A243A1A7B1AFEF1F99056F9F975" box="[296,403,1652,1679]" captionStart="FIGURES 156158" captionStartId="68.[151,264,1913,1937]" captionTargetBox="[151,1435,1130,1889]" captionTargetId="figure-70@68.[151,1435,1130,1890]" captionTargetPageId="68" captionText="FIGURES 156158. Head, thorax and wing of Scatophila. 156 Scatophila bipiliaris Sturtevant &amp; Wheeler, head and thorax, dorsal view. 157158. Wing, dorsal view. 157. S. conifera Sturtevant &amp; Wheeler. 158. S. cribrata (Stenhammar). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210887" httpUri="https://zenodo.org/record/13210887/files/figure.png" pageId="73" pageNumber="74">Figs. 157</figureCitation>
,
<figureCitation id="13432A243A1A7B1AFE46F99056A4F974" box="[415,462,1652,1678]" captionStart="FIGURES 175179" captionStartId="76.[151,264,1403,1427]" captionTargetBox="[164,1423,181,1379]" captionTargetId="figure-30@76.[164,1423,181,1379]" captionTargetPageId="76" captionText="FIGURES 175179. Head and thorax of Scatophila. 175177. Head and thorax, dorsal view. 175. Scatophila bisignata Cresson. 176. S. conifera Sturtevant &amp; Wheeler. 177. S. cribrata (Stenhammar). 178179. Head of Scatophila cribrata (Stenhammar). 178. Latero-oblique view. 179. Lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210907" httpUri="https://zenodo.org/record/13210907/files/figure.png" pageId="73" pageNumber="74">176</figureCitation>
): Mesonotum (
<figureCitation id="13432A243A1A7B1AFDAEF99055BCF974" box="[631,726,1652,1678]" captionStart="FIGURES 175179" captionStartId="76.[151,264,1403,1427]" captionTargetBox="[164,1423,181,1379]" captionTargetId="figure-30@76.[164,1423,181,1379]" captionTargetPageId="76" captionText="FIGURES 175179. Head and thorax of Scatophila. 175177. Head and thorax, dorsal view. 175. Scatophila bisignata Cresson. 176. S. conifera Sturtevant &amp; Wheeler. 177. S. cribrata (Stenhammar). 178179. Head of Scatophila cribrata (Stenhammar). 178. Latero-oblique view. 179. Lateral view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210907" httpUri="https://zenodo.org/record/13210907/files/figure.png" pageId="73" pageNumber="74">Fig. 176</figureCitation>
) generally gray with some irregular brownish pattern medially, variable; scutellum gray. Wing (
<figureCitation id="13432A243A1A7B1AFE2FF97C553EF949" box="[502,596,1688,1715]" captionStart="FIGURES 156158" captionStartId="68.[151,264,1913,1937]" captionTargetBox="[151,1435,1130,1889]" captionTargetId="figure-70@68.[151,1435,1130,1890]" captionTargetPageId="68" captionText="FIGURES 156158. Head, thorax and wing of Scatophila. 156 Scatophila bipiliaris Sturtevant &amp; Wheeler, head and thorax, dorsal view. 157158. Wing, dorsal view. 157. S. conifera Sturtevant &amp; Wheeler. 158. S. cribrata (Stenhammar). Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210887" httpUri="https://zenodo.org/record/13210887/files/figure.png" pageId="73" pageNumber="74">Fig. 157</figureCitation>
) generally pale colored, spots pale; costal vein ratio 0.210.25; M
<subScript id="17FC34E43A1A7B1AFAFFF94E5244F942" attach="left" box="[1318,1326,1706,1720]" fontSize="6" pageId="73" pageNumber="74">1</subScript>
vein ratio 0.600.68; knob of halter yellow. Legs generally dark colored; femora and tibiae dark with gray microtomentum; male midfemur lacking row of spine-like setulae along posteroventral margin; tarsi blackish.
</paragraph>
<paragraph id="8BC736A13A1A7B19FF1EF8E05511F82A" blockId="73.[151,1437,1436,2039]" lastBlockId="74.[151,1437,1470,2001]" lastPageId="74" lastPageNumber="75" pageId="73" pageNumber="74">
<emphasis id="B90CEAB33A1A7B1AFF1EF8E0565BF8E4" box="[199,305,1796,1822]" italics="true" pageId="73" pageNumber="74">Abdomen</emphasis>
(
<figureCitation id="13432A243A1A7B1AFE9AF8E05687F8E4" box="[323,493,1796,1822]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="73" pageNumber="74">Figs. 172174</figureCitation>
): Abdomen mostly black, subshiny to shiny, with small gray to whitish gray area laterally toward posterior margin on basal 13 tergites. Male terminalia (
<figureCitation id="13432A243A1A7B1AFC74F8CC5338F8B8" box="[941,1106,1832,1858]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="73" pageNumber="74">Figs. 172174</figureCitation>
): Epandrium in posterior view (
<figureCitation id="13432A243A1A7B1AFF46F8A8566AF89C" box="[159,256,1868,1894]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="73" pageNumber="74">Fig. 172</figureCitation>
) generally irregularly and broadly rectangular to oval, width about 3/4 height, ventral margin somewhat truncate (
<figureCitation id="13432A243A1A7B1AFED8F894560BF870" box="[257,353,1904,1930]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="73" pageNumber="74">Fig. 172</figureCitation>
) with medial, shallowly rounded projection, ventral margin distinctly wider than width of cercal opening; dorsal margin of epandrium in posterior view (
<figureCitation id="13432A243A1A7B1AFCF9F87054E8F854" box="[800,898,1940,1966]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="73" pageNumber="74">Fig. 172</figureCitation>
) rounded, lateral margins widest at midheight, thereafter ventrally tapered to ventral margin, ventrolateral corners rounded; epandrial setulae about equal in length, sparser medially and laterally, those along ventral margin more abundant but shorter; height of cercal opening more than 1/3 epandrial height; cerci in posterior view (
<figureCitation id="13432A243A197B19FD0CFA5A545CFA23" box="[725,822,1470,1497]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 172</figureCitation>
) semihemispherical, slightly more narrowed dorsally; aedeagus in lateral view (
<figureCitation id="13432A243A197B19FE6EFA06557DFA07" box="[439,535,1506,1533]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 174</figureCitation>
) obtusely pointed basally, forming irregular rectangle, ventral margin shallowly rounded, posterior margin broadly rounded, right angle at posterodorsal corner, dorsal margin deeply and widely excavated basally, cavity deeper than width of opening, in ventral view (
<figureCitation id="13432A243A197B19FC2BF9CE533CF9BF" box="[1010,1110,1578,1605]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 173</figureCitation>
) ovately rectangular, lateral margins shallowly curved, wider basally, tapered gradually toward apex, apical margin truncated and with a long, acutely pointed, medial point; ventral aedeagal process shallowly C-shaped; phallapodeme in lateral view (
<figureCitation id="13432A243A197B19FAB6F99657A8F94B" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 174</figureCitation>
) L-shaped, elongate, narrow, parallel sided, dorsal half gently and shallowly curved, ventral extension nearly straight, elongate, narrow, tapered, in ventral view (
<figureCitation id="13432A243A197B19FD02F95E5451F92F" box="[731,827,1722,1749]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 173</figureCitation>
) T-shaped, width of basal crossbar shorter than length of medial process, crossbar nearly straight, nearly as thick as medial process, medial process with lateral margins shallowly sinuous; postgonite in lateral view (
<figureCitation id="13432A243A197B19FD49F8E65584F8E7" box="[656,750,1794,1821]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 174</figureCitation>
) irregularly V-shaped, with dorsal arm irregularly rectangular, anterodorsal corner produced to narrow, digitiform extension, anterodorsal corner moderately produced with posterodorsal angle produced to short, pointed extension, ventral arm elongate, thin, parallel-sided, in ventral view (
<figureCitation id="13432A243A197B19FF46F88A566BF873" box="[159,257,1902,1929]" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 173</figureCitation>
) with base slopping laterally, nearly straight, not excavated, posterior, thumb-like extensions narrowly developed, apices produced laterally, bearing setulae along medial margin; neohypandrium in lateral view (
<figureCitation id="13432A243A197B19FAB6F87657A8F82B" captionStart="FIGURES 172174" captionStartId="75.[151,264,1078,1102]" captionTargetBox="[159,1427,189,1047]" captionTargetId="figure-27@75.[151,1435,181,1055]" captionTargetPageId="75" captionText="FIGURES 172174. Male terminalia of Scatophila conifera Sturtevant &amp; Wheeler. 172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="74" pageNumber="75">Fig. 174</figureCitation>
) narrow, elongate, distinctly C-shaped.
</paragraph>
<caption id="DF0766293A197B19FF4EFAD4536DFA96" ID-DOI="http://doi.org/10.5281/zenodo.13210901" ID-Zenodo-Dep="13210901" httpUri="https://zenodo.org/record/13210901/files/figure.png" pageId="74" pageNumber="75" startId="74.[151,264,1328,1352]" targetBox="[170,1435,206,1304]" targetPageId="74" targetType="figure">
<paragraph id="8BC736A13A197B19FF4EFAD4536DFA96" blockId="74.[151,1436,1328,1388]" pageId="74" pageNumber="75">
<emphasis id="B90CEAB33A197B19FF4EFAD45603FAB2" bold="true" box="[151,361,1328,1352]" pageId="74" pageNumber="75">FIGURES 168171</emphasis>
. Head of
<taxonomicName id="4C784D223A197B19FE0DFAD454E5FAB2" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[468,911,1328,1352]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="74" pageNumber="75" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A197B19FE0DFAD455F7FAB2" box="[468,669,1328,1352]" italics="true" pageId="74" pageNumber="75">Scatophila conifera</emphasis>
Sturtevant &amp; Wheeler.
</taxonomicName>
168. Male, latero-oblique view. 169. Male, lateral view. 170. Female, latero-oblique view. 171. Female, lateral view. Scale bar = 0.2 mm.
</paragraph>
</caption>
<caption id="DF0766293A187B18FF4EFBD2564DFB6C" ID-DOI="http://doi.org/10.5281/zenodo.13210903" ID-Zenodo-Dep="13210903" httpUri="https://zenodo.org/record/13210903/files/figure.png" pageId="75" pageNumber="76" startId="75.[151,264,1078,1102]" targetBox="[159,1427,189,1047]" targetPageId="75" targetType="figure">
<paragraph id="8BC736A13A187B18FF4EFBD2564DFB6C" blockId="75.[151,1437,1078,1175]" pageId="75" pageNumber="76">
<emphasis id="B90CEAB33A187B18FF4EFBD25602FBB4" bold="true" box="[151,360,1078,1102]" pageId="75" pageNumber="76">FIGURES 172174</emphasis>
. Male terminalia of
<taxonomicName id="4C784D223A187B18FDE0FBD35486FBB5" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[569,1004,1079,1103]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A187B18FDE0FBD3546AFBB5" box="[569,768,1079,1103]" italics="true" pageId="75" pageNumber="76">Scatophila conifera</emphasis>
Sturtevant &amp; Wheeler.
</taxonomicName>
172. Epandrium and cerci, posterior view. 173. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 174. Same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A187B18FF1EFB385460FA9B" blockId="75.[151,1437,1243,1594]" pageId="75" pageNumber="76">
Type Material.—
<materialsCitation id="3B103CFC3A187B18FE57FB3F55D8FA98" collectingDate="1951-07-24" collectionCode="R, TYPE, ANSP" pageId="75" pageNumber="76" specimenCount="3" typeStatus="holotype">
The
<typeStatus id="54C388033A187B18FE11FB3F5540FB0F" box="[456,554,1243,1269]" pageId="75" pageNumber="76" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C784D223A187B18FD78FB3F53CDFB0F" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" box="[673,1191,1243,1270]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="conifera">
<emphasis id="B90CEAB33A187B18FD78FB3F54E9FB0C" box="[673,899,1243,1270]" italics="true" pageId="75" pageNumber="76">Scatophila conifera</emphasis>
Sturtevant and Wheeler
</taxonomicName>
is labeled “Weott[,] Calif[ornia][,] 7-24-51 [
<date id="FFC610613A187B18FE70FAE45559FAE0" box="[425,563,1279,1306]" pageId="75" pageNumber="76" value="1951-07-24">
<collectingDate id="EF82E9893A187B18FE70FAE45559FAE0" box="[425,563,1279,1306]" pageId="75" pageNumber="76" value="1951-07-24">24 Jul 1951</collectingDate>
</date>
][handwritten]/M.
<collectionCode id="ED69AE643A187B18FD27FAE4547BFAE0" box="[766,785,1280,1306]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="75" pageNumber="76">R</collectionCode>
. Wheeler/
<typeStatus id="54C388033A187B18FC51FAE45376FAE0" box="[904,1052,1280,1306]" pageId="75" pageNumber="76" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C784D223A187B18FBFFFB1B52F7FAE3" authority="Stvt &amp; Whlr" authorityName="Stvt &amp; Whlr" box="[1062,1437,1279,1306]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="conifera">Scatophila conifera Stvt &amp; Whlr</taxonomicName>
[pink]/6683
<collectionCode id="ED69AE643A187B18FEFAFAC05603FAC4" box="[291,361,1316,1342]" pageId="75" pageNumber="76">TYPE</collectionCode>
[dark pink; number handwritten].” The
<typeStatus id="54C388033A187B18FCF6FAC754FBFAC7" box="[815,913,1315,1341]" pageId="75" pageNumber="76" type="holotype">holotype</typeStatus>
is double mounted (glued to a paper point), is in good condition, and is deposited in the
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</materialsCitation>
(6683).
</paragraph>
<paragraph id="8BC736A13A187B18FF1EFA8F55C9FA50" blockId="75.[151,1437,1243,1594]" pageId="75" pageNumber="76">
Other Specimens Examined.—
<materialsCitation id="3B103CFC3A187B18FDFDFA885676FA50" collectingDate="1951-07-24" country="United States of America" latitude="40.321667" location="Weott" longLatPrecision="116" longitude="-123.92167" municipality="Humboldt" pageId="75" pageNumber="76" specimenCount="1" stateProvince="California">
<collectingCountry id="F36F76313A187B18FDFDFA885593FA7C" box="[548,761,1388,1414]" name="United States of America" pageId="75" pageNumber="76">UNITED STATES</collectingCountry>
.
<emphasis id="B90CEAB33A187B18FCDDFA885346FA7F" box="[772,1068,1387,1414]" italics="true" pageId="75" pageNumber="76">
<collectingRegion id="49BCF8433A187B18FCDDFA8854C2FA7C" box="[772,936,1388,1414]" country="United States of America" name="California" pageId="75" pageNumber="76">CALIFORNIA</collectingRegion>
.
<collectingMunicipality id="6BA3ACDB3A187B18FC6AFA8F534CFA7F" box="[947,1062,1387,1413]" pageId="75" pageNumber="76">Humboldt</collectingMunicipality>
:
</emphasis>
<location id="8EA7607A3A187B18FBEAFA88531FFA7C" LSID="urn:lsid:plazi:treatment:03D187B73A1A7B18FF4EFB5955BEF9C3:8EA7607A3A187B18FBEAFA88531FFA7C" box="[1075,1141,1388,1414]" country="United States of America" latitude="40.321667" longLatPrecision="116" longitude="-123.92167" municipality="Humboldt" name="Weott" pageId="75" pageNumber="76" stateProvince="California">Weott</location>
(
<geoCoordinate id="EE4C50663A187B18FB5CFA885391FA7C" box="[1157,1275,1388,1414]" degrees="40" direction="north" minutes="19.3" orientation="latitude" pageId="75" pageNumber="76" precision="92" value="40.321667">40°19.3'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A187B18FADFFA8F52E1FA7C" box="[1286,1419,1387,1414]" degrees="123" direction="west" minutes="55.3" orientation="longitude" pageId="75" pageNumber="76" precision="92" value="-123.92167">123°55.3'W</geoCoordinate>
),
<date id="FFC610613A187B18FF4EFA745676FA50" box="[151,284,1423,1450]" pageId="75" pageNumber="76" value="1951-07-24">
<collectingDate id="EF82E9893A187B18FF4EFA745676FA50" box="[151,284,1423,1450]" pageId="75" pageNumber="76" value="1951-07-24">24 Jul 1951</collectingDate>
</date>
</materialsCitation>
,
<materialsCitation id="3B103CFC3A187B18FEF1FA7455F5FA50" box="[296,671,1423,1450]" collectionCode="USNM" country="United States of America" location="Wheeler" pageId="75" pageNumber="76" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="M">
<collectingRegion id="49BCF8433A187B18FEF1FA74562EFA50" box="[296,324,1424,1450]" country="United States of America" name="M" pageId="75" pageNumber="76">M</collectingRegion>
.
<collectingRegion id="49BCF8433A187B18FE96FA74560FFA50" box="[335,357,1424,1450]" country="United States of America" name="R" pageId="75" pageNumber="76">R</collectingRegion>
.
<location id="8EA7607A3A187B18FEB6FA6B56A4FA53" LSID="urn:lsid:plazi:treatment:03D187B73A1A7B18FF4EFB5955BEF9C3:8EA7607A3A187B18FEB6FA6B56A4FA53" box="[367,462,1423,1449]" country="United States of America" name="Wheeler" pageId="75" pageNumber="76" stateProvince="M">Wheeler</location>
(
<specimenCount id="9D7EFD283A187B18FE07FA6B5569FA50" box="[478,515,1423,1450]" count="5" pageId="75" pageNumber="76" type="male">5♂</specimenCount>
,
<specimenCount id="9D7EFD283A187B18FDD6FA74555EFA50" box="[527,564,1424,1450]" count="2" pageId="75" pageNumber="76" type="female">2♀</specimenCount>
;
<collectionCode id="ED69AE643A187B18FD98FA7455FDFA50" box="[577,663,1424,1450]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="75" pageNumber="76" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A187B18FF1EFA5053FAFA34" blockId="75.[151,1437,1243,1594]" box="[199,1168,1459,1486]" pageId="75" pageNumber="76">
<materialsCitation id="3B103CFC3A187B18FF1EFA5053E0FA34" box="[199,1162,1459,1486]" country="United States of America" latitude="40.321667" location="Weott" longLatPrecision="116" longitude="-123.92167" municipality="Humboldt" pageId="75" pageNumber="76" specimenCount="1" stateProvince="California" typeStatus="holotype">
<typeStatus id="54C388033A187B18FF1EFA505795FA34" box="[199,255,1460,1486]" pageId="75" pageNumber="76">Type</typeStatus>
Locality.—
<collectingCountry id="F36F76313A187B18FE5DFA575577FA34" box="[388,541,1459,1486]" name="United States of America" pageId="75" pageNumber="76">United States</collectingCountry>
.
<collectingRegion id="49BCF8433A187B18FDF1FA5755F7FA37" box="[552,669,1459,1485]" country="United States of America" name="California" pageId="75" pageNumber="76">California</collectingRegion>
.
<collectingMunicipality id="6BA3ACDB3A187B18FD71FA575474FA37" box="[680,798,1459,1485]" pageId="75" pageNumber="76">Humboldt</collectingMunicipality>
:
<location id="8EA7607A3A187B18FCF1FA505400FA34" LSID="urn:lsid:plazi:treatment:03D187B73A1A7B18FF4EFB5955BEF9C3:8EA7607A3A187B18FCF1FA505400FA34" box="[808,874,1460,1486]" country="United States of America" latitude="40.321667" longLatPrecision="116" longitude="-123.92167" municipality="Humboldt" name="Weott" pageId="75" pageNumber="76" stateProvince="California">Weott</location>
(
<geoCoordinate id="EE4C50663A187B18FCA0FA505485FA34" box="[889,1007,1460,1486]" degrees="40" direction="north" minutes="19.3" orientation="latitude" pageId="75" pageNumber="76" precision="92" value="40.321667">40°19.3'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A187B18FC23FA575315FA34" box="[1018,1151,1459,1486]" degrees="123" direction="west" minutes="55.3" orientation="longitude" pageId="75" pageNumber="76" precision="92" value="-123.92167">123°55.3'W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A187B18FF1EFA335419FA0B" blockId="75.[151,1437,1243,1594]" box="[199,883,1495,1522]" pageId="75" pageNumber="76">
Distribution (
<figureCitation id="13432A243A187B18FE87FA3356C5FA08" box="[350,431,1495,1522]" captionStart="FIGURE 30" captionStartId="22.[151,250,1023,1047]" captionTargetBox="[151,1435,181,999]" captionTargetId="figure-30@22.[151,1435,181,999]" captionTargetPageId="22" captionText="FIGURE 30. Distribution map of Scatophila angusta sp. nov. (circles), S. conifera Sturtevant &amp; Wheeler (triangle), and S. contaminata (Stenhammar) (squares) in Nearctic Region." figureDoi="http://doi.org/10.5281/zenodo.13210791" httpUri="https://zenodo.org/record/13210791/files/figure.png" pageId="75" pageNumber="76">Fig. 30</figureCitation>
).—
<emphasis id="B90CEAB33A187B18FE02FA3C5529FA0B" box="[475,579,1496,1521]" italics="true" pageId="75" pageNumber="76">
<taxonomicName id="4C784D223A187B18FE02FA3C5557FA0B" box="[475,573,1496,1521]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
:
</emphasis>
<collectingCountry id="F36F76313A187B18FD93FA33558AFA08" box="[586,736,1495,1522]" name="United States of America" pageId="75" pageNumber="76">United States</collectingCountry>
(
<collectingRegion id="49BCF8433A187B18FD36FA335402FA0B" box="[751,872,1495,1521]" country="United States of America" name="California" pageId="75" pageNumber="76">California</collectingRegion>
).
</paragraph>
<paragraph id="8BC736A13A187B18FF1EFA1F55BEF9C3" blockId="75.[151,1437,1243,1594]" pageId="75" pageNumber="76">
Remarks.—This species is placed in the
<emphasis id="B90CEAB33A187B18FD4FFA1F545AF9EF" box="[662,816,1531,1557]" italics="true" pageId="75" pageNumber="76">quadriguttata</emphasis>
group and is known thus far only from the
<typeStatus id="54C388033A187B18FAFDFA18523EF9EC" box="[1316,1364,1532,1558]" pageId="75" pageNumber="76">type</typeStatus>
series that was collected in Humboldt County,
<collectingRegion id="49BCF8433A187B18FD82F9FB55BAF9C3" box="[603,720,1567,1593]" country="United States of America" name="California" pageId="75" pageNumber="76">California</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>