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<mods:title id="48D1E81254FEC8C783662CAF1BA22644">Revision of the Nearctic Species of the Shore-Fly Genus Scatophila Becker (Diptera: Ephydridae)</mods:title>
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<mods:affiliation id="81E296544E84F818A12752C3EDB5FFBA">Institute of Biology, Opole University, ul. Oleska 22, 45 - 052 Opole, Poland zatwar @ uni. opole. pl; https: // orcid. org / 0000 - 0003 - 2163 - 0143</mods:affiliation>
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<mods:namePart id="094B0C0C39C93AE7AFE02A67C41E19AF">Mathis, Wayne N.</mods:namePart>
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24.
<taxonomicName id="4C784D223A047B04FF1AF8DA55D0F8A3" ID-CoL="4V44K" authority="Sturtevant and Wheeler" authorityName="Sturtevant and Wheeler" box="[195,698,1854,1881]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A047B04FF1AF8DA56ECF8A3" bold="true" box="[195,390,1854,1881]" italics="true" pageId="87" pageNumber="88">Scatophila picta</emphasis>
Sturtevant and Wheeler
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<paragraph id="8BC736A13A047B04FF4EF88756B5F887" blockId="87.[151,698,1854,1917]" box="[151,479,1891,1917]" pageId="87" pageNumber="88">
<figureCitation id="13432A243A047B04FF4EF887566BF887" box="[151,257,1891,1917]" captionStart="FIGURE 167" captionStartId="73.[151,250,1107,1131]" captionTargetBox="[151,1435,181,1083]" captionTargetId="figure-27@73.[151,1435,181,1083]" captionTargetPageId="73" captionText="FIGURE 167. Distribution map of Scatophila bisignata Cresson (circles), and S. picta Sturtevant &amp; Wheeler (square)." figureDoi="http://doi.org/10.5281/zenodo.13210897" httpUri="https://zenodo.org/record/13210897/files/figure.png" pageId="87" pageNumber="88">Figs. 167</figureCitation>
,
<figureCitation id="13432A243A047B04FED4F8875650F887" box="[269,314,1891,1917]" captionStart="FIGURES 202204" captionStartId="85.[151,264,1848,1872]" captionTargetBox="[151,1435,1094,1825]" captionTargetId="figure-325@85.[151,1435,1094,1825]" captionTargetPageId="85" captionText="FIGURES 202204. Head, thorax, and wing of Scatophila. 202. S. hirsuta Sturtevant &amp; Wheeler, head and thorax, dorsal view. 203. S. mystacina sp. nov. male, wing, dorsal view. 204 S. picta Sturtevant &amp; Wheeler, wing, dorsal view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210935" httpUri="https://zenodo.org/record/13210935/files/figure.png" pageId="87" pageNumber="88">204</figureCitation>
,
<figureCitation id="13432A243A047B04FE9CF88756C0F887" box="[325,426,1891,1917]" captionStart-0="FIGURES 212214" captionStart-1="FIGURES 215219" captionStartId-0="88.[151,264,1010,1034]" captionStartId-1="89.[151,264,1540,1564]" captionTargetBox-0="[159,1427,189,977]" captionTargetBox-1="[167,1435,181,1516]" captionTargetId-0="figure-30@88.[151,1435,181,985]" captionTargetId-1="figure-27@89.[151,1435,181,1517]" captionTargetPageId-0="88" captionTargetPageId-1="89" captionText-0="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." captionText-1="FIGURES 215219. Head, thorax, and wing of Scatophila. 215216. Head of Scatophila picta Sturtevant &amp; Wheeler. 215. Latero-oblique view. 216. Lateral view. 217. Scatophila mystacina sp. nov., head and thorax, dorsal view. 218219. Wing, dorsal view. 218. S. pulchra Sturtevant &amp; Wheeler. 219. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.13210945" figureDoi-1="http://doi.org/10.5281/zenodo.13210947" httpUri-0="https://zenodo.org/record/13210945/files/figure.png" httpUri-1="https://zenodo.org/record/13210947/files/figure.png" pageId="87" pageNumber="88">212216</figureCitation>
,
<figureCitation id="13432A243A047B04FE6CF88756B5F887" box="[437,479,1891,1917]" captionStart="FIGURES 221225" captionStartId="91.[151,264,1444,1468]" captionTargetBox="[151,1435,193,1421]" captionTargetId="figure-27@91.[151,1435,181,1421]" captionTargetPageId="91" captionText="FIGURES 221225. Head and thorax of Scatophila. 221222. Head of S. pulchra Sturtevant &amp; Wheeler. 221. Latero-oblique view. 222. Lateral view. 223225. Head and thorax, dorsal view. 223. Scatophila picta Sturtevant &amp; Wheeler. 224. S. pulchra Sturtevant &amp; Wheeler. 225. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210953" httpUri="https://zenodo.org/record/13210953/files/figure.png" pageId="87" pageNumber="88">223</figureCitation>
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<treatmentCitation id="0AD910B03A047B04FF4EF84455D5F842" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[151,703,1952,1977]" page="196" pageId="87" pageNumber="88" year="1954">
<taxonomicName id="4C784D223A047B04FF4EF84455D5F842" ID-CoL="4V44K" authority="Sturtevant and Wheeler 1954: 196" authorityName="Sturtevant and Wheeler" authorityPageNumber="196" authorityYear="1954" box="[151,703,1952,1977]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="87" pageNumber="88" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A047B04FF4EF8445628F842" box="[151,322,1952,1976]" italics="true" pageId="87" pageNumber="88">Scatophila picta</emphasis>
<bibRefCitation id="EFE94B503A047B04FE97F84555D5F842" author="Sturtevant, A. H. &amp; Wheeler, M. R." box="[334,703,1952,1977]" pageId="87" pageNumber="88" pagination="151 - 257" refId="ref70834" refString="Sturtevant, A. H. &amp; Wheeler, M. R. (1954) Synopses of Nearctic Ephydridae (Diptera). Transactions of the American Entomological Society, 79, 151 - 257." type="journal article" year="1954">Sturtevant and Wheeler 1954: 196</bibRefCitation>
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.—
<treatmentCitation id="0AD910B03A047B04FD06F84454FAF842" author="Wirth, W. W." box="[735,912,1952,1976]" page="758" pageId="87" pageNumber="88" year="1965">
<bibRefCitation id="EFE94B503A047B04FD06F84454FAF842" author="Wirth, W. W." box="[735,912,1952,1976]" pageId="87" pageNumber="88" pagination="734 - 759" refId="ref70987" refString="Wirth, W. W. (1965) Ephydridae. In: Stone, A., Sabrosky, C. W., Wirth, W. W., Foote, R. W. &amp; Coulson, J. R. (Eds.), A Catalog of the Diptera of America North of Mexico. Handbook 276. U. S. Department of Agriculture, Washington, D. C., pp. 734 - 759." type="book chapter" year="1965">Wirth 1965: 758</bibRefCitation>
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[
<taxonomicName id="4C784D223A047B04FC7BF8445491F842" box="[930,1019,1952,1976]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="87" pageNumber="97" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
catalog].—
<treatmentCitation id="0AD910B03A047B04FBAAF84452F6F843" author="Zatwarnicki, T." box="[1139,1436,1952,1977]" page="293" pageId="87" pageNumber="88" year="1987">
<bibRefCitation id="EFE94B503A047B04FBAAF84452F6F843" author="Zatwarnicki, T." box="[1139,1436,1952,1977]" pageId="87" pageNumber="88" pagination="277 - 298" refId="ref71260" refString="Zatwarnicki, T. (1987) New synonyms and records of Palearctic Scatophila (Diptera, Ephydridae). Polskie Pismo Entomologiczne, 57 (2), 277 - 298." type="journal article" year="1987">Zatwarnicki 1987: 293294</bibRefCitation>
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[checklist].—
<treatmentCitation id="0AD910B03A047B04FE94F85B55D6F82D" author="Zatwarnicki, T. &amp; Mathis, W. N." box="[333,700,1983,2007]" page="366" pageId="87" pageNumber="88" year="1994">
<bibRefCitation id="EFE94B503A047B04FE94F85B55D6F82D" author="Zatwarnicki, T. &amp; Mathis, W. N." box="[333,700,1983,2007]" pageId="87" pageNumber="88" pagination="351 - 370" refId="ref71480" refString="Zatwarnicki, T. &amp; Mathis, W. N. (1994) Phylogeny and classification of the genus Scatophila Becker (Diptera: Ephydridae). Annales de la Societe Entomologique de France, 29 (4), 351 - 370. [1993] https: // doi. org / 10.1080 / 21686351.1993.12277698" type="journal article" year="1994">Zatwarnicki and Mathis 1994: 366</bibRefCitation>
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[phylogeny, not placed in a species group].—
<treatmentCitation id="0AD910B03A047B04FB7BF85B5647F80C" author="Mathis, W. N. &amp; Zatwarnicki, T." page="285" pageId="87" pageNumber="88" year="1995">
<bibRefCitation id="EFE94B503A047B04FB7BF85B5647F80C" author="Mathis, W. N. &amp; Zatwarnicki, T." pageId="87" pageNumber="88" pagination="1 - 423" refId="ref70301" refString="Mathis, W. N. &amp; Zatwarnicki, T. (1995) A world catalog of the shore flies (Diptera: Ephydridae). International Memoirs on Entomology, 4, i - vi + 1 - 423." type="journal article" year="1995">Mathis and Zatwarnicki 1995: 285</bibRefCitation>
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[world catalog].
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<emphasis id="B90CEAB33A0B7B0BFF4EFC16560CFBF0" bold="true" box="[151,358,1010,1034]" pageId="88" pageNumber="89">FIGURES 212214</emphasis>
. Male terminalia of
<taxonomicName id="4C784D223A0B7B0BFDEAFC1654AAFBF0" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[563,960,1010,1034]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="88" pageNumber="89" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A0B7B0BFDEAFC1655B2FBF0" box="[563,728,1010,1034]" italics="true" pageId="88" pageNumber="89">Scatophila picta</emphasis>
Sturtevant &amp; Wheeler.
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212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm.
</paragraph>
</caption>
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Diagnosis.—This species is distinguished from congeners by the following combination of characters: This species is mostly dark colored but with contrasted coloration; small shore flies, body length
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.
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<emphasis id="B90CEAB33A0B7B0BFF1EFB08566EFAFC" box="[199,260,1260,1286]" italics="true" pageId="88" pageNumber="89">Head</emphasis>
(
<figureCitation id="13432A243A0B7B0BFECFFB0856D6FAFD" box="[278,444,1260,1287]" captionStart="FIGURES 215219" captionStartId="89.[151,264,1540,1564]" captionTargetBox="[167,1435,181,1516]" captionTargetId="figure-27@89.[151,1435,181,1517]" captionTargetPageId="89" captionText="FIGURES 215219. Head, thorax, and wing of Scatophila. 215216. Head of Scatophila picta Sturtevant &amp; Wheeler. 215. Latero-oblique view. 216. Lateral view. 217. Scatophila mystacina sp. nov., head and thorax, dorsal view. 218219. Wing, dorsal view. 218. S. pulchra Sturtevant &amp; Wheeler. 219. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210947" httpUri="https://zenodo.org/record/13210947/files/figure.png" pageId="88" pageNumber="89">Figs. 215216</figureCitation>
,
<figureCitation id="13432A243A0B7B0BFE12FB085690FAFC" box="[459,506,1260,1286]" captionStart="FIGURES 221225" captionStartId="91.[151,264,1444,1468]" captionTargetBox="[151,1435,193,1421]" captionTargetId="figure-27@91.[151,1435,181,1421]" captionTargetPageId="91" captionText="FIGURES 221225. Head and thorax of Scatophila. 221222. Head of S. pulchra Sturtevant &amp; Wheeler. 221. Latero-oblique view. 222. Lateral view. 223225. Head and thorax, dorsal view. 223. Scatophila picta Sturtevant &amp; Wheeler. 224. S. pulchra Sturtevant &amp; Wheeler. 225. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210953" httpUri="https://zenodo.org/record/13210953/files/figure.png" pageId="88" pageNumber="89">223</figureCitation>
): Frons mostly brown with gray spots anteriorly, 1 medially, and 2 laterally, just dorsad of antennal bases; fronto-orbits blackish brown with narrow, gray stripe immediately adjacent to eye. Face of
<specimenCount id="9D7EFD283A0B7B0BFF4EFAD157C6FAB5" box="[151,172,1333,1359]" count="1" pageId="88" pageNumber="89" type="male"></specimenCount>
with medial area very shallowly depressed to flat, ventral margin slightly projected, bearing several setulae; facial coloration gray except for inverted Y-shaped marking, each arm of Y robustly developed and with patch of setulae; oral margin with 23 stout setae on each side, above them with cluster of smaller setulae arising from a brown area. Gena moderately high, height subequal to height of basal flagellomere; gena-to-eye ratio 0.27.
</paragraph>
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<emphasis id="B90CEAB33A0B7B0BFF1EFA205672FA24" box="[199,280,1476,1502]" italics="true" pageId="88" pageNumber="89">Thorax</emphasis>
(
<figureCitation id="13432A243A0B7B0BFEF5FA2056F6FA24" box="[300,412,1476,1502]" captionStart="FIGURES 202204" captionStartId="85.[151,264,1848,1872]" captionTargetBox="[151,1435,1094,1825]" captionTargetId="figure-325@85.[151,1435,1094,1825]" captionTargetPageId="85" captionText="FIGURES 202204. Head, thorax, and wing of Scatophila. 202. S. hirsuta Sturtevant &amp; Wheeler, head and thorax, dorsal view. 203. S. mystacina sp. nov. male, wing, dorsal view. 204 S. picta Sturtevant &amp; Wheeler, wing, dorsal view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210935" httpUri="https://zenodo.org/record/13210935/files/figure.png" pageId="88" pageNumber="89">Figs. 204</figureCitation>
,
<figureCitation id="13432A243A0B7B0BFE72FA2056B0FA24" box="[427,474,1476,1502]" captionStart="FIGURES 221225" captionStartId="91.[151,264,1444,1468]" captionTargetBox="[151,1435,193,1421]" captionTargetId="figure-27@91.[151,1435,181,1421]" captionTargetPageId="91" captionText="FIGURES 221225. Head and thorax of Scatophila. 221222. Head of S. pulchra Sturtevant &amp; Wheeler. 221. Latero-oblique view. 222. Lateral view. 223225. Head and thorax, dorsal view. 223. Scatophila picta Sturtevant &amp; Wheeler. 224. S. pulchra Sturtevant &amp; Wheeler. 225. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210953" httpUri="https://zenodo.org/record/13210953/files/figure.png" pageId="88" pageNumber="89">223</figureCitation>
): Mesonotum (
<figureCitation id="13432A243A0B7B0BFD52FA205585FA24" box="[651,751,1476,1502]" captionStart="FIGURES 221225" captionStartId="91.[151,264,1444,1468]" captionTargetBox="[151,1435,193,1421]" captionTargetId="figure-27@91.[151,1435,181,1421]" captionTargetPageId="91" captionText="FIGURES 221225. Head and thorax of Scatophila. 221222. Head of S. pulchra Sturtevant &amp; Wheeler. 221. Latero-oblique view. 222. Lateral view. 223225. Head and thorax, dorsal view. 223. Scatophila picta Sturtevant &amp; Wheeler. 224. S. pulchra Sturtevant &amp; Wheeler. 225. S. rubribrunnea Sturtevant &amp; Wheeler. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210953" httpUri="https://zenodo.org/record/13210953/files/figure.png" pageId="88" pageNumber="89">Fig. 223</figureCitation>
) generally mottled, mixture of brown and gray. Presutural dorsocentral setae lacking. Scutum mottled, generally mostly brown medially and mostly gray laterally, with linear gray spots between acrostichal rows subanteriorly and with gray vitta between acrostichal and dorsocentral tracks anteriorly. Scutellum mostly brown with a distinct basal and apical gray spots, lateral margins darkened; presutural dorsocentral setae lacking; pleural area extensively gray but with brown markings; anepisternum with large brown spot subdorsally; an irregular brown area at ventral portion of anepisternum and anepimeron and dorsal area of katepisternum. Wing (
<figureCitation id="13432A243A0B7B0BFE4BF9785699F94C" box="[402,499,1692,1718]" captionStart="FIGURES 202204" captionStartId="85.[151,264,1848,1872]" captionTargetBox="[151,1435,1094,1825]" captionTargetId="figure-325@85.[151,1435,1094,1825]" captionTargetPageId="85" captionText="FIGURES 202204. Head, thorax, and wing of Scatophila. 202. S. hirsuta Sturtevant &amp; Wheeler, head and thorax, dorsal view. 203. S. mystacina sp. nov. male, wing, dorsal view. 204 S. picta Sturtevant &amp; Wheeler, wing, dorsal view. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.13210935" httpUri="https://zenodo.org/record/13210935/files/figure.png" pageId="88" pageNumber="89">Fig. 204</figureCitation>
) generally infuscate, brown with distinctive white spots and markings, length 2.00 mm; Cell r
<subScript id="17FC34E43A0B7B0BFED6F936567DF91A" attach="left" box="[271,279,1746,1760]" fontSize="6" pageId="88" pageNumber="89">1</subScript>
with 4 white spots, apical spot less distinct; veins R
<subScript id="17FC34E43A0B7B0BFC8FF9365405F91A" attach="left" box="[854,879,1746,1760]" fontSize="6" pageId="88" pageNumber="89">2+3</subScript>
and R
<subScript id="17FC34E43A0B7B0BFC6FF93654BAF91A" attach="left" box="[950,976,1746,1760]" fontSize="6" pageId="88" pageNumber="89">4+5</subScript>
shallowly sinuous; costal vein ratio 0.27; M
<subScript id="17FC34E43A0B7B0BFF69F91257D2F8FE" attach="left" box="[176,184,1782,1796]" fontSize="6" pageId="88" pageNumber="89">1</subScript>
vein ratio 0.72. Knob of halter yellow. Femora and tibiae black; male midfemur lacking row of spine-like setulae along posteroventral margin; basitarsi yellow; other tarsomeres becoming darker with apical 12 brown.
</paragraph>
<paragraph id="8BC736A13A0B7B09FF1EF8C85343FE73" blockId="88.[151,1437,1188,2043]" lastBlockId="90.[151,1437,151,645]" lastPageId="90" lastPageNumber="91" pageId="88" pageNumber="89">
<emphasis id="B90CEAB33A0B7B0BFF1EF8C8565BF8BC" box="[199,305,1836,1862]" italics="true" pageId="88" pageNumber="89">Abdomen</emphasis>
(
<figureCitation id="13432A243A0B7B0BFE9FF8C85698F8BC" box="[326,498,1836,1862]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="88" pageNumber="89">Figs. 212214</figureCitation>
): Tergites dark brown from dorsal view; lateral and ventral portions with gray markings, especially toward posterior margins. Male terminalia (
<figureCitation id="13432A243A0B7B0BFCB0F8B45365F890" box="[873,1039,1872,1898]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="88" pageNumber="89">Figs. 212214</figureCitation>
): Epandrium in posterior view (
<figureCitation id="13432A243A0B7B0BFAB6F8B457A9F874" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="88" pageNumber="89">Fig. 212</figureCitation>
) generally irregularly rectangular, width almost 1/2 height, ventral margin somewhat truncate (
<figureCitation id="13432A243A0B7B0BFB23F8905231F874" box="[1274,1371,1908,1934]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="88" pageNumber="89">Fig. 212</figureCitation>
) with a very slight medial indentation, ventral margin only slightly wider than width of cercal opening; dorsal margin of epandrium in posterior view (
<figureCitation id="13432A243A0B7B0BFE3AF858552EF82C" box="[483,580,1980,2006]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="88" pageNumber="89">Fig. 212</figureCitation>
) broadly rounded, with very obtuse angles at corners, lateral margins widest at midheight, midheight indicated by lateral, shallow, bump, thereafter ventrally somewhat parallel sided to rounded posterolateral corner; epandrial setulae about equal in length, sparser medially and laterally, those along ventral and dorsal margins longer; width of cercal opening about 0.55 epandrial height, lateral margins sinuous, bulged at ventral 2/3, thereafter ventrally tapered to more narrowly rounded ventral margin; cerci in posterior view (
<figureCitation id="13432A243A0A7B0AFAB6F91B57A9F8C7" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="89" pageNumber="90">Fig. 212</figureCitation>
) robustly lunate, slightly more narrowed dorsally; aedeagus in lateral view (
<figureCitation id="13432A243A0A7B0AFBFBF8C753E9F8C7" box="[1058,1155,1827,1853]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="89" pageNumber="90">Fig. 214</figureCitation>
) robustly and irregularly rounded basally, generally forming irregular triangle, posterior portion broadly and shallowly sinuously truncate, lateral margins tapered to ventral, narrowly rounded apex, lacking a dorsal cavity, but with a long, sinuous, dorsal thumb-like projection, in ventral view (
<figureCitation id="13432A243A0A7B0AFD8FF86B55DDF853" box="[598,695,1935,1961]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="89" pageNumber="90">Fig. 213</figureCitation>
) bell-like, basal margin more narrowly rounded, thereafter flared toward ventral margin, ventrolateral corners angulate, ventral margin broadly and shallow rounded with a medial, shallow triangular notch; ventral aedeagal process sinuous; phallapodeme in lateral view (
<figureCitation id="13432A243A0A7B0AFBABF83353BAF80B" box="[1138,1232,2007,2033]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="89" pageNumber="90">Fig. 214</figureCitation>
) elongate, narrow, elbowed, dorsal extension about 1/3 ventral extension, dorsal extension with slightly expanded apical half, in ventral view (
<figureCitation id="13432A243A097B09FF06FF5F562AFF2F" box="[223,320,187,213]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="90" pageNumber="91">Fig. 213</figureCitation>
) T-shaped, width of basal crossbar about half length of medial process, crossbar tapered laterally, another crossbar below, as an inverted, robust W; postgonite in lateral view (
<figureCitation id="13432A243A097B09FBDBFF3B530EFF03" box="[1026,1124,223,249]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="90" pageNumber="91">Fig. 214</figureCitation>
) irregularly V-shaped, with dorsal arm somewhat rectangular, anteroventral corner produced, thumb-like but with acutely pointed apex, bearing setulae, ventral arm elongate, parallel-sided, nearly straight, in ventral view (
<figureCitation id="13432A243A097B09FC22FEC35331FEBB" box="[1019,1115,295,321]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="90" pageNumber="91">Fig. 213</figureCitation>
) 2 postgonites forming a Hlike structure with medially directed, narrow process at basal 2/3, apices rounded, bearing setulae along lateral and apical margins; neohypandrium in lateral view (
<figureCitation id="13432A243A097B09FD69FE8B547AFE73" box="[688,784,367,393]" captionStart="FIGURES 212214" captionStartId="88.[151,264,1010,1034]" captionTargetBox="[159,1427,189,977]" captionTargetId="figure-30@88.[151,1435,181,985]" captionTargetPageId="88" captionText="FIGURES 212214. Male terminalia of Scatophila picta Sturtevant &amp; Wheeler. 212. Epandrium and cerci, posterior view. 213. Internal male terminalia (gonite, phallapodeme, aedeagus, neohypandrium), ventral view. 214. Same, lateral view. Scale bar = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.13210945" httpUri="https://zenodo.org/record/13210945/files/figure.png" pageId="90" pageNumber="91">Fig. 214</figureCitation>
) narrow, deeply sinuous.
</paragraph>
<caption id="DF0766293A0A7B0AFF4EF9E05259F99E" ID-DOI="http://doi.org/10.5281/zenodo.13210947" ID-Zenodo-Dep="13210947" httpUri="https://zenodo.org/record/13210947/files/figure.png" pageId="89" pageNumber="90" startId="89.[151,264,1540,1564]" targetBox="[167,1435,181,1516]" targetPageId="89" targetType="figure">
<paragraph id="8BC736A13A0A7B0AFF4EF9E05259F99E" blockId="89.[151,1437,1540,1637]" pageId="89" pageNumber="90">
<emphasis id="B90CEAB33A0A7B0AFF4EF9E05605F9E7" bold="true" box="[151,367,1540,1565]" pageId="89" pageNumber="90">FIGURES 215219.</emphasis>
Head, thorax, and wing of
<emphasis id="B90CEAB33A0A7B0AFD4BF9E1546FF9E7" box="[658,773,1541,1565]" italics="true" pageId="89" pageNumber="90">
<taxonomicName id="4C784D223A0A7B0AFD4BF9E15468F9E7" box="[658,770,1541,1565]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="genus">Scatophila</taxonomicName>
.
</emphasis>
215216. Head of
<taxonomicName id="4C784D223A0A7B0AFC09F9E15202F9E7" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[976,1384,1541,1565]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A0A7B0AFC09F9E15312F9E6" box="[976,1144,1541,1565]" italics="true" pageId="89" pageNumber="90">Scatophila picta</emphasis>
Sturtevant &amp; Wheeler.
</taxonomicName>
215. Latero-oblique view. 216. Lateral view. 217.
<taxonomicName id="4C784D223A0A7B0AFDA8F9CD5424F9BA" authorityName="Zatwarnicki &amp; Mathis USNM" authorityYear="2024" box="[625,846,1577,1601]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="mystacina" status="sp. nov.">
<emphasis id="B90CEAB33A0A7B0AFDA8F9CD5424F9BA" box="[625,846,1577,1601]" italics="true" pageId="89" pageNumber="90">Scatophila mystacina</emphasis>
</taxonomicName>
<emphasis id="B90CEAB33A0A7B0AFC8FF9CD54C0F9BB" bold="true" box="[854,938,1577,1601]" pageId="89" pageNumber="90">
<taxonomicNameLabel id="A23F57C83A0A7B0AFC8FF9CD54C0F9BB" box="[854,938,1577,1601]" pageId="89" pageNumber="90" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, head and thorax, dorsal view. 218
<emphasis id="B90CEAB33A0A7B0AFAC4F9CD5240F9BB" bold="true" box="[1309,1322,1577,1601]" pageId="89" pageNumber="90"></emphasis>
219. Wing, dorsal view. 218.
<taxonomicName id="4C784D223A0A7B0AFE93F9A955F7F99F" authority="Sturtevant &amp; Wheeler." authorityName="Sturtevant &amp; Wheeler." box="[330,669,1613,1637]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="pulchra">
<emphasis id="B90CEAB33A0A7B0AFE93F9A956D9F99F" box="[330,435,1613,1637]" italics="true" pageId="89" pageNumber="90">S. pulchra</emphasis>
Sturtevant &amp; Wheeler.
</taxonomicName>
219.
<taxonomicName id="4C784D223A0A7B0AFD0CF9A953F7F99F" authority="Sturtevant &amp; Wheeler. Scale" authorityName="Sturtevant &amp; Wheeler. Scale" box="[725,1181,1613,1637]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="89" pageNumber="90" phylum="Arthropoda" rank="species" species="rubribrunnea">
<emphasis id="B90CEAB33A0A7B0AFD0CF9A9541DF99F" box="[725,887,1613,1637]" italics="true" pageId="89" pageNumber="90">S. rubribrunnea</emphasis>
Sturtevant &amp; Wheeler. Scale
</taxonomicName>
bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8BC736A13A097B09FF1EFE775393FE0F" blockId="90.[151,1437,151,645]" pageId="90" pageNumber="91">
Type Material.—The
<typeStatus id="54C388033A097B09FE6DFE77557CFE57" box="[436,534,403,429]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
male of
<taxonomicName id="4C784D223A097B09FDB6FE775449FE56" box="[623,803,403,429]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="picta">
<emphasis id="B90CEAB33A097B09FDB6FE775449FE56" box="[623,803,403,429]" italics="true" pageId="90" pageNumber="91">Scatophila picta</emphasis>
</taxonomicName>
is labeled “MatanuskaAlaska
<date id="FFC610613A097B09FBB2FE775382FE57" box="[1131,1256,402,429]" pageId="90" pageNumber="91" value="1945-10-02">Oct.2 1945</date>
RotaryTrapColl J.C.Chamberlin [“2” and “5” handwritten]/
<typeStatus id="54C388033A097B09FD51FE535476FE2B" box="[648,796,439,465]" pageId="90" pageNumber="91" type="holotype">HOLOTYPE</typeStatus>
<taxonomicName id="4C784D223A097B09FCFEFE535310FE2B" authority="Stvt &amp; Whlr" authorityName="Stvt &amp; Whlr" box="[807,1146,439,465]" class="Insecta" family="Ephydridae" genus="Scatophila" kingdom="Animalia" order="Diptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="species" species="picta">Scatophila picta Stvt &amp; Whlr</taxonomicName>
[pink].” The
<typeStatus id="54C388033A097B09FAC4FE535215FE2B" box="[1309,1407,439,465]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
is double mounted (glued to a paper triangle), is in excellent condition, and is deposited in the USNM.
</paragraph>
<paragraph id="8BC736A13A097B09FF1EFE1B5263FDE3" blockId="90.[151,1437,151,645]" box="[199,1289,510,537]" pageId="90" pageNumber="91">
<materialsCitation id="3B103CFC3A097B09FF1EFE1B5269FDE3" box="[199,1283,510,537]" country="United States of America" latitude="61.541668" location="Matanuska" longLatPrecision="102" longitude="-149.23" municipality="Matanuska-Susitna" pageId="90" pageNumber="91" specimenCount="1" stateProvince="Alaska" typeStatus="holotype">
<typeStatus id="54C388033A097B09FF1EFE1B5795FDE3" box="[199,255,511,537]" pageId="90" pageNumber="91">Type</typeStatus>
Locality.—
<collectingCountry id="F36F76313A097B09FE5DFE1B5577FDE3" box="[388,541,511,537]" name="United States of America" pageId="90" pageNumber="91">United States</collectingCountry>
.
<collectingRegion id="49BCF8433A097B09FDFEFE1B5512FDE3" box="[551,632,511,537]" country="United States of America" name="Alaska" pageId="90" pageNumber="91">Alaska</collectingRegion>
.
<collectingMunicipality id="6BA3ACDB3A097B09FD5AFE1B5434FDE3" box="[643,862,511,537]" pageId="90" pageNumber="91">Matanuska-Susitna</collectingMunicipality>
:
<location id="8EA7607A3A097B09FCB1FE1B5489FDE3" LSID="urn:lsid:plazi:treatment:03D187B73A047B09FF4EF8DA57BFFD7F:8EA7607A3A097B09FCB1FE1B5489FDE3" box="[872,995,511,537]" country="United States of America" latitude="61.541668" longLatPrecision="102" longitude="-149.23" municipality="Matanuska-Susitna" name="Matanuska" pageId="90" pageNumber="91" stateProvince="Alaska">Matanuska</location>
(
<geoCoordinate id="EE4C50663A097B09FC2BFE1A530DFDE3" box="[1010,1127,510,537]" degrees="61" direction="north" minutes="32.5" orientation="latitude" pageId="90" pageNumber="91" precision="92" value="61.541668">61°32.5'N</geoCoordinate>
,
<geoCoordinate id="EE4C50663A097B09FBAAFE1B5392FDE3" box="[1139,1272,511,537]" degrees="149" direction="west" minutes="13.8" orientation="longitude" pageId="90" pageNumber="91" precision="92" value="-149.23">149°13.8'W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BC736A13A097B09FF1EFDC75437FDC7" blockId="90.[151,1437,151,645]" box="[199,861,547,573]" pageId="90" pageNumber="91">
Distribution (
<figureCitation id="13432A243A097B09FE87FDC756D7FDC7" box="[350,445,547,573]" captionStart="FIGURE 167" captionStartId="73.[151,250,1107,1131]" captionTargetBox="[151,1435,181,1083]" captionTargetId="figure-27@73.[151,1435,181,1083]" captionTargetPageId="73" captionText="FIGURE 167. Distribution map of Scatophila bisignata Cresson (circles), and S. picta Sturtevant &amp; Wheeler (square)." figureDoi="http://doi.org/10.5281/zenodo.13210897" httpUri="https://zenodo.org/record/13210897/files/figure.png" pageId="90" pageNumber="91">Fig. 167</figureCitation>
).—
<emphasis id="B90CEAB33A097B09FE30FDC7553BFDC6" box="[489,593,547,572]" italics="true" pageId="90" pageNumber="91">
<taxonomicName id="4C784D223A097B09FE30FDC75521FDC6" box="[489,587,547,572]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="90" pageNumber="91" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
:
</emphasis>
<collectingCountry id="F36F76313A097B09FD81FDC75584FDC7" box="[600,750,547,573]" name="United States of America" pageId="90" pageNumber="91">United States</collectingCountry>
(
<collectingRegion id="49BCF8433A097B09FD24FDC7543BFDC7" box="[765,849,547,573]" country="United States of America" name="Alaska" pageId="90" pageNumber="91">Alaska</collectingRegion>
).
</paragraph>
<paragraph id="8BC736A13A097B09FF1EFDA357BFFD7F" blockId="90.[151,1437,151,645]" pageId="90" pageNumber="91">
Remarks.—This species is placed in the
<emphasis id="B90CEAB33A097B09FD4EFDA3545BFD9B" box="[663,817,583,609]" italics="true" pageId="90" pageNumber="91">quadriguttata</emphasis>
group, and is only known, thus far, from the
<typeStatus id="54C388033A097B09FAE3FDA352F6FD9B" box="[1338,1436,583,609]" pageId="90" pageNumber="91" type="holotype">holotype</typeStatus>
male.
</paragraph>
</subSubSection>
</treatment>
</document>