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<mods:title id="32DFFC7DEFFA28CE08EB6B7163A1F144">New records, diagnostics and preliminary checklist of the superfamily Pentatomoidea (Hemiptera: Heteroptera) from the Comoro Islands</mods:title>
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<mods:namePart id="B4CCCF16A2DB95232782C833F1F3D3C2">Nicolas, Vincent</mods:namePart>
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<mods:namePart id="F8C3848519E4C637EEF381550F608C22">Carapezza, Attilio</mods:namePart>
<mods:affiliation id="FA2C716FA4D76966894583034283A930">University of Palermo; corresponding address: via Sandro Botticelli 15, 90144 Palermo, Italy</mods:affiliation>
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<mods:namePart id="326C8DE6124796B2D3E0BED646E0B404">Rider, David A.</mods:namePart>
<mods:affiliation id="E422FE7C9A5DD008008DD088B23FBEFB">Department of Entomology, North Dakota State University, Fargo, North Dakota, USA</mods:affiliation>
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<mods:affiliation id="EEF9E0170EE9B0A32D8F6E14192B6970">Department of Entomology, National Museum of the Czech Republic, Cirkusová 1740, CZ- 193 00 Praha 9 - Horní Počernice, Czech Republic</mods:affiliation>
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<treatment id="03C2025643415C4D28FFF901D574FEC4" ID-DOI="http://doi.org/10.5281/zenodo.12742681" ID-Zenodo-Dep="12742681" LSID="urn:lsid:plazi:treatment:03C2025643415C4D28FFF901D574FEC4" httpUri="http://treatment.plazi.org/id/03C2025643415C4D28FFF901D574FEC4" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<subSubSection id="C371E0CB43415C4A28FFF901D516F8B0" box="[151,630,1783,1810]" pageId="11" pageNumber="12" type="nomenclature">
<paragraph id="8BD4B34043415C4A28FFF901D516F8B0" blockId="11.[151,630,1783,1845]" box="[151,630,1783,1810]" pageId="11" pageNumber="12">
<heading id="D09C042C43415C4A28FFF901D516F8B0" bold="true" box="[151,630,1783,1810]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
<taxonomicName id="4C6BC8C343415C4A28FFF901D516F8B0" authority="Signoret, 1851" authorityName="Signoret" authorityYear="1851" box="[151,630,1783,1810]" class="Insecta" family="Pentatomidae" genus="Eurysaspis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="transversalis">
<emphasis id="B91F6F5243415C4A28FFF901D516F8B0" bold="true" box="[151,630,1783,1810]" pageId="11" pageNumber="12">
<emphasis id="B91F6F5243415C4A28FFF901D6DCF8B3" bold="true" box="[151,444,1783,1810]" italics="true" pageId="11" pageNumber="12">Eurysaspis transversalis</emphasis>
<bibRefCitation id="EFFACEB143415C4A29ABF901D516F8B0" author="Signoret, V." box="[451,630,1783,1809]" pageId="11" pageNumber="12" pagination="329 - 348" refId="ref23109" refString="Signoret, V. (1851) Description de nouvelles especes d'Hemipteres. Annales de la Societe Entomologique de France, Serie 2, 9, 329 - 348, pl. 10." type="journal article" year="1851">Signoret, 1851</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C371E0CB43415C4A28FFF8EDD799F894" box="[151,249,1819,1845]" pageId="11" pageNumber="12" type="description">
<paragraph id="8BD4B34043415C4A28FFF8EDD799F894" blockId="11.[151,630,1783,1845]" box="[151,249,1819,1845]" pageId="11" pageNumber="12">
(
<figureCitation id="1350AFC543415C4A28F7F8EDD791F894" box="[159,241,1819,1845]" captionStart="FIGURES 1923" captionStartId="11.[151,264,1533,1557]" captionTargetBox="[151,1435,181,1509]" captionTargetId="figure-20@11.[151,1435,181,1509]" captionTargetPageId="11" captionText="FIGURES 1923. Habitus of Comoro Pentatomidae: 19—Coquerelia ventralis Horváth, 1904; 20—Eurysaspis transversalis Signoret, 1851; 21—Aesula viridissima (Signoret, 1861); 22—Neoacrosternum validum (Horváth, 1904), inset—detail of antennae; 23—Anoano pronotalis Cachan, 1952. Photos: V. Nicolas." figureDoi="http://doi.org/10.5281/zenodo.12742620" httpUri="https://zenodo.org/record/12742620/files/figure.png" pageId="11" pageNumber="12">Fig. 20</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C371E0CB43415C4A28FFF895D457F864" pageId="11" pageNumber="12" type="materials_examined">
<paragraph id="8BD4B34043415C4A28FFF895D457F864" blockId="11.[151,1436,1891,2026]" pageId="11" pageNumber="12">
<emphasis id="B91F6F5243415C4A28FFF895D541F8DC" bold="true" box="[151,545,1891,1917]" pageId="11" pageNumber="12">Material examined. ANJOUAN:</emphasis>
<materialsCitation id="3B03B91D43415C4A2A44F895D274F8DC" box="[556,1300,1891,1918]" collectingDate="1974-01-15" collectionCode="MNHN" collectorName="L. Matile" determinerName="P. Kment" location="Bambao" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1">
<location id="8EB4E59B43415C4A2A44F895D5ECF8DC" LSID="urn:lsid:plazi:treatment:03C2025643415C4D28FFF901D574FEC4:8EB4E59B43415C4A2A44F895D5ECF8DC" box="[556,652,1891,1917]" name="Bambao" pageId="11" pageNumber="12">Bambao</location>
,
<date id="FFD5958043415C4A2AF4F895D468F8DF" box="[668,776,1891,1918]" pageId="11" pageNumber="12" value="1974-01-15">
<collectingDate id="EF916C6843415C4A2AF4F895D468F8DF" box="[668,776,1891,1918]" pageId="11" pageNumber="12" value="1974-01-15">15.i.1974</collectingDate>
</date>
,
<specimenCount id="9D6D78C943415C4A2B70F895D45DF8DC" box="[792,829,1891,1917]" count="1" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
,
<collectorName id="269ED69643415C4A2B26F892D4DAF8DC" box="[846,954,1891,1918]" pageId="11" pageNumber="12">L. Matile</collectorName>
lgt.,
<determinerName id="7E085F8443415C4A2B95F892D308F8DF" box="[1021,1128,1892,1918]" pageId="11" pageNumber="12">P. Kment</determinerName>
det. (
<collectionCode id="ED7A2B8543415C4A2CD8F895D26CF8DC" box="[1200,1292,1891,1917]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="11" pageNumber="12">MNHN</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B03B91D43415C4A2D4BF895D39AF800" collectingDate="1991-05-05" collectionCode="BMNH" collectorName="Le Galawa H. &amp; K. M. Guichard" determinerName="P. Kment" pageId="11" pageNumber="12" specimenCount="3" specimenCount-female="2" specimenCount-male="1">
<emphasis id="B91F6F5243415C4A2D4BF895D642F800" bold="true" pageId="11" pageNumber="12">GRANDE COMORE:</emphasis>
<collectorName id="269ED69643415C4A2941F87ED6A7F803" box="[297,455,1927,1954]" pageId="11" pageNumber="12">Le Galawa H.</collectorName>
, 22.iv. &amp;
<date id="FFD5958043415C4A2A5DF871D5F8F803" box="[565,664,1927,1954]" pageId="11" pageNumber="12" value="1991-05-05">
<collectingDate id="EF916C6843415C4A2A5DF871D5F8F803" box="[565,664,1927,1954]" pageId="11" pageNumber="12" value="1991-05-05">5.v.1991</collectingDate>
</date>
,
<specimenCount id="9D6D78C943415C4A2ACCF871D5A8F800" box="[676,712,1927,1953]" count="1" pageId="11" pageNumber="12" type="male">1♂</specimenCount>
,
<specimenCount id="9D6D78C943415C4A2ABDF871D599F800" box="[725,761,1927,1953]" count="2" pageId="11" pageNumber="12" type="female">2♀</specimenCount>
,
<collectorName id="269ED69643415C4A2B6EF87ED4D8F800" box="[774,952,1927,1954]" pageId="11" pageNumber="12">K. M. Guichard</collectorName>
lgt.,
<determinerName id="7E085F8443415C4A2B9AF87ED338F803" box="[1010,1112,1928,1954]" pageId="11" pageNumber="12">P. Kment</determinerName>
det. (
<collectionCode id="ED7A2B8543415C4A2CF0F871D392F800" box="[1176,1266,1927,1953]" pageId="11" pageNumber="12">BMNH</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B03B91D43415C4A2D6DF871D453F864" collectingDate="2019-10-02" collectionCode="VNBF" collectorName="V. Nicolas" country="Mayotte" location="Pengoualajou" pageId="11" pageNumber="12" specimenCount="1" specimenCount-female="1">
<emphasis id="B91F6F5243415C4A2D6DF871D2FCF800" bold="true" box="[1285,1436,1927,1953]" pageId="11" pageNumber="12">
<collectingCountry id="F37CF3D043415C4A2D6DF871D2F8F800" box="[1285,1432,1927,1953]" name="Mayotte" pageId="11" pageNumber="12">MAYOTTE</collectingCountry>
:
</emphasis>
<location id="8EB4E59B43415C4A28FFF85DD652F864" LSID="urn:lsid:plazi:treatment:03C2025643415C4D28FFF901D574FEC4:8EB4E59B43415C4A28FFF85DD652F864" box="[151,306,1963,1989]" country="Mayotte" name="Pengoualajou" pageId="11" pageNumber="12">Pengoualajou</location>
,
<date id="FFD5958043415C4A2956F85DD6C2F864" box="[318,418,1963,1989]" pageId="11" pageNumber="12" value="2019-10-02">
<collectingDate id="EF916C6843415C4A2956F85DD6C2F864" box="[318,418,1963,1989]" pageId="11" pageNumber="12" value="2019-10-02">2.x.2019</collectingDate>
</date>
,
<specimenCount id="9D6D78C943415C4A29C6F85DD6B1F864" box="[430,465,1963,1989]" count="1" pageId="11" pageNumber="12" type="female">1♀</specimenCount>
,
<collectorName id="269ED69643415C4A29B6F85AD533F864" box="[478,595,1963,1990]" pageId="11" pageNumber="12">V. Nicolas</collectorName>
lgt. &amp; det. (
<collectionCode id="ED7A2B8543415C4A2AB2F85DD44BF864" box="[730,811,1963,1989]" pageId="11" pageNumber="12">VNBF</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C371E0CB43415C4A28AFF839D2E3F848" box="[199,1411,1999,2026]" pageId="11" pageNumber="12" type="biology_ecology">
<paragraph id="8BD4B34043415C4A28AFF839D2E3F848" blockId="11.[151,1436,1891,2026]" box="[199,1411,1999,2026]" pageId="11" pageNumber="12">
<emphasis id="B91F6F5243415C4A28AFF839D647F848" bold="true" box="[199,295,1999,2025]" pageId="11" pageNumber="12">Biology.</emphasis>
This species has been found on
<taxonomicName id="4C6BC8C343415C4A2AFAF826D430F848" box="[658,848,2000,2026]" class="Magnoliopsida" family="Fabaceae" genus="Senna" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="species" species="singueana">
<emphasis id="B91F6F5243415C4A2AFAF826D430F848" box="[658,848,2000,2026]" italics="true" pageId="11" pageNumber="12">Senna singueana</emphasis>
</taxonomicName>
(
<taxonomicName id="4C6BC8C343415C4A2B37F839D4ABF848" box="[863,971,1999,2025]" class="Magnoliopsida" family="Fabaceae" kingdom="Plantae" order="Fabales" pageId="11" pageNumber="12" phylum="Tracheophyta" rank="family">Fabaceae</taxonomicName>
) in
<collectingCountry id="F37CF3D043415C4A2B9EF826D333F84B" box="[1014,1107,2000,2026]" name="Mayotte" pageId="11" pageNumber="12">Mayotte</collectingCountry>
(V. Nicolas, pers. observ.).
</paragraph>
</subSubSection>
<subSubSection id="C371E0CB43465C4D28AFFF60D6B7FF74" pageId="12" pageNumber="13" type="distribution">
<paragraph id="8BD4B34043465C4D28AFFF60D6B7FF74" blockId="12.[151,1437,150,357]" pageId="12" pageNumber="13">
<emphasis id="B91F6F5243465C4D28AFFF60D601FF11" bold="true" box="[199,353,150,176]" pageId="12" pageNumber="13">Distribution.</emphasis>
<collectingCountry id="F37CF3D043465C4D290FFF61D6B2FF10" box="[359,466,151,177]" name="Comoros" pageId="12" pageNumber="13">Comoros</collectingCountry>
:
<collectingRegion id="49AF7DA243465C4D29B3FF61D55EFF10" box="[475,574,151,177]" country="Comoros" name="Anjouan" pageId="12" pageNumber="13">Anjouan</collectingRegion>
,
<collectingRegion id="49AF7DA243465C4D2A22FF61D460FF10" box="[586,768,151,177]" country="Comoros" name="Grande Comore" pageId="12" pageNumber="13">Grande Comore</collectingRegion>
,
<collectingCountry id="F37CF3D043465C4D2B63FF61D408FF10" box="[779,872,151,177]" name="Mayotte" pageId="12" pageNumber="13">Mayotte</collectingCountry>
(new records);
<collectingCountry id="F37CF3D043465C4D2C71FF61D3C0FF10" box="[1049,1184,151,177]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
(
<bibRefCitation id="EFFACEB143465C4D2CC6FF61D227FF10" author="Cachan, P." box="[1198,1351,150,177]" pageId="12" pageNumber="13" pagination="231 - 462" refId="ref14750" refString="Cachan, P. (1952) Les Pentatomidae de Madagascar (Hemipteres Heteropteres). Memoires de l'Institut Scientifique de Madagascar (E), 1 (2), 231 - 462, pls. 6 - 14." type="journal article" year="1952">Cachan 1952</bibRefCitation>
);
<collectingCountry id="F37CF3D043465C4D2D33FF61D2F9FF10" box="[1371,1433,151,177]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
:
<collectingCountry id="F37CF3D043465C4D28FFFF4DD678FF74" box="[151,280,187,213]" name="India" pageId="12" pageNumber="13">Puducherry</collectingCountry>
(
<bibRefCitation id="EFFACEB143465C4D294FFF4DD6AAFF74" author="Signoret, V." box="[295,458,186,213]" pageId="12" pageNumber="13" pagination="329 - 348" refId="ref23109" refString="Signoret, V. (1851) Description de nouvelles especes d'Hemipteres. Annales de la Societe Entomologique de France, Serie 2, 9, 329 - 348, pl. 10." type="journal article" year="1851">Signoret 1851</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C371E0CB43465C4D28AFFF28D574FEC4" pageId="12" pageNumber="13" type="discussion">
<paragraph id="8BD4B34043465C4D28AFFF28D574FEC4" blockId="12.[151,1437,150,357]" pageId="12" pageNumber="13">
<emphasis id="B91F6F5243465C4D28AFFF28D62FFF59" bold="true" box="[199,335,222,248]" pageId="12" pageNumber="13">Comments.</emphasis>
This species was described from
<collectingCountry id="F37CF3D043465C4D2AB0FF29D438FF58" box="[728,856,223,249]" name="India" pageId="12" pageNumber="13">Pondichery</collectingCountry>
[=
<collectingCountry id="F37CF3D043465C4D2BECFF29D368FF58" box="[900,1032,223,249]" name="India" pageId="12" pageNumber="13">Puducherry</collectingCountry>
] by
<bibRefCitation id="EFFACEB143465C4D2C56FF29D394FF58" author="Signoret, V." box="[1086,1268,222,249]" pageId="12" pageNumber="13" pagination="329 - 348" refId="ref23109" refString="Signoret, V. (1851) Description de nouvelles especes d'Hemipteres. Annales de la Societe Entomologique de France, Serie 2, 9, 329 - 348, pl. 10." type="journal article" year="1851">Signoret (1851)</bibRefCitation>
, but has never been reported from
<collectingCountry id="F37CF3D043465C4D291CFEF5D6CEFEBC" box="[372,430,259,285]" name="India" pageId="12" pageNumber="13">India</collectingCountry>
(cf.
<bibRefCitation id="EFFACEB143465C4D2988FEF5D5E9FEBC" author="Atkinson, E. T." box="[480,649,259,285]" pageId="12" pageNumber="13" pagination="118 - 184" refId="ref13914" refString="Atkinson, E. T. (1888) Notes on Indian Rhynchota, Heteroptera No. 4. Journal of the Asiatic Society of Bengal, 57, 118 - 184." type="journal article" year="1888">Atkinson 1888</bibRefCitation>
,
<bibRefCitation id="EFFACEB143465C4D2AFCFEF5D446FEBC" author="Distant, W. L." box="[660,806,259,285]" pageId="12" pageNumber="13" pagination="1 - 438" refId="ref15518" refString="Distant, W. L. (1902) Rhynchota. Vol. I. Heteroptera. In: Blanford, W. T. (Ed.), The Fauna of British India Including Ceylon and Burma. Taylor and Francis, London, pp. i - xxxvii + 1 - 438. https: // doi. org / 10.5962 / bhl. title. 9193" type="book chapter" year="1902">Distant 1902</bibRefCitation>
, Salini &amp; Vitaktamath 2015). This suggests that the
<typeStatus id="54D00DE243465C4D2D04FEF5D2FCFEBC" box="[1388,1436,259,285]" pageId="12" pageNumber="13">type</typeStatus>
specimen(s) were mislabelled. Accordingly, we treat the species here as subendemic to
<collectingCountry id="F37CF3D043465C4D2C19FED1D3B8FEE0" box="[1137,1240,295,321]" name="Comoros" pageId="12" pageNumber="13">Comoros</collectingCountry>
and
<collectingCountry id="F37CF3D043465C4D2D67FED1D2F8FEE0" box="[1295,1432,295,321]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
, and it is not a widespread species.
</paragraph>
</subSubSection>
</treatment>
</document>