175 lines
13 KiB
XML
175 lines
13 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3789792" ID-GBIF-Dataset="08b92fdc-9bba-469a-8382-0d7f0fb89bb7" ID-GBIF-Taxon="163794427" ID-ISSN="1313–2970" ID-Zenodo-Dep="3789792" ID-ZooBank="7761253A-8625-4DDA-B33B-4AD041F16667" LSID="urn:lsid:zoobank.org:act:BB668AEE-029A-4FDC-8127-E0DA27AC3C29" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1588010087092" checkinUser="plazi" docAuthor="Maruyama, Munetoshi" docDate="2010" docId="039487F4FF881D530BAAFB8AFD22F91C" docLanguage="en" docName="ZK_article_2138.pdf" docOrigin="ZooKeys 34 (34)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@3804a9a8" docStyleName="zookeys.2008.journal_article" docTitle="Pterobius itiokai Maruyama 2010, sp. n." docType="treatment" docUuid="BB668AEE-029A-4FDC-8127-E0DA27AC3C29" docUuidSource="ZooBank" docVersion="4" lastPageId="3" lastPageNumber="52" masterDocId="FFADFF8CFF8A1D500B26FFB3FFC3FF9B" masterDocTitle="A new genus and species of myrmecophilous aphodiine beetle (Coleoptera, Scarabaeidae) inhabiting the myrmecophytic epiphyte Platycerium sp. (Polypodiaceae) in the Bornean rainforest canopy" masterLastPageNumber="54" masterPageNumber="49" pageId="2" pageNumber="51" updateTime="1643562133277" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A new genus and species of myrmecophilous aphodiine beetle (Coleoptera, Scarabaeidae) inhabiting the myrmecophytic epiphyte Platycerium sp. (Polypodiaceae) in the Bornean rainforest canopy</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Maruyama, Munetoshi</mods:namePart>
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<mods:affiliation>The Kyushu University Museum, Fukuoka, Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-01-28</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>34</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>49</mods:start>
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<mods:end>54</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.34.281</mods:identifier>
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<mods:identifier type="GBIF-Dataset">08b92fdc-9bba-469a-8382-0d7f0fb89bb7</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576604</mods:identifier>
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<mods:identifier type="ZooBank">7761253A-8625-4DDA-B33B-4AD041F16667</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3789792" ID-GBIF-Taxon="163794427" ID-Zenodo-Dep="3789792" LSID="urn:lsid:zoobank.org:act:BB668AEE-029A-4FDC-8127-E0DA27AC3C29" httpUri="http://treatment.plazi.org/id/039487F4FF881D530BAAFB8AFD22F91C" lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51">
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<subSubSection pageId="2" pageNumber="51" type="nomenclature">
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<paragraph blockId="2.[140,991,1081,1143]" box="[140,556,1081,1108]" pageId="2" pageNumber="51">
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<heading box="[140,556,1081,1108]" fontSize="11" level="2" pageId="2" pageNumber="51" reason="3">
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<emphasis bold="true" box="[140,556,1081,1108]" pageId="2" pageNumber="51">
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<taxonomicName authority="Maruyama" authorityName="Maruyama" authorityYear="2010" box="[140,479,1081,1108]" class="Insecta" family="Aphodiidae" genus="Pterobius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="51" phylum="Arthropoda" rank="species" species="itiokai" status="sp. n.">
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<emphasis bold="true" box="[140,340,1081,1108]" italics="true" pageId="2" pageNumber="51">Pterobius itiokai</emphasis>
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Maruyama
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</taxonomicName>
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,
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<taxonomicNameLabel box="[491,556,1082,1108]" pageId="2" pageNumber="51" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph blockId="2.[140,991,1081,1143]" box="[140,991,1117,1143]" pageId="2" pageNumber="51">
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<uri box="[140,991,1117,1143]" pageId="2" pageNumber="51">
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urn:lsid:zoobank.org:act:
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<uuid box="[415,991,1117,1143]" pageId="2" pageNumber="51">BB668AEE-029A-4FDC-8127-E0DA27AC3C29</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,980,1187,1214]" pageId="2" pageNumber="51" type="etymology">
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<paragraph blockId="2.[140,1108,1187,1672]" box="[140,980,1187,1214]" pageId="2" pageNumber="51">
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<emphasis bold="true" box="[140,275,1187,1213]" pageId="2" pageNumber="51">
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<emphasis box="[140,270,1187,1213]" inLineHeading="true" pageId="2" pageNumber="51" reason="1">Etymology</emphasis>
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.
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</emphasis>
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Dedicated to the collector of the
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<typeStatus box="[657,705,1188,1214]" pageId="2" pageNumber="51">type</typeStatus>
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series. Dr. Takao Itioka.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="2" pageNumber="51" type="materials_examined">
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<paragraph blockId="2.[140,1108,1187,1672]" pageId="2" pageNumber="51">
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<emphasis bold="true" box="[185,323,1222,1249]" pageId="2" pageNumber="51">
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<emphasis box="[185,243,1223,1249]" inLineHeading="true" pageId="2" pageNumber="51" reason="1">Type</emphasis>
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series.
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</emphasis>
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<materialsCitation ID-GBIF-Occurrence="2608997689" collectingDate="2005-09-16" country="Malaysia" location="Lambir Hills National Park" municipality="Taman Negara Bukit Lambir" pageId="2" pageNumber="51" specimenCount="2" stateProvince="Sarawak" typeStatus="holotype">
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<emphasis box="[331,426,1223,1249]" italics="true" pageId="2" pageNumber="51">
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<typeStatus box="[331,426,1223,1249]" pageId="2" pageNumber="51">Holotype</typeStatus>
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</emphasis>
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, male, “
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<collectingCountry box="[518,663,1223,1249]" name="Malaysia" pageId="2" pageNumber="51">MALAYSIA</collectingCountry>
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:
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<collectingRegion box="[674,770,1223,1249]" country="Malaysia" name="Sarawak" pageId="2" pageNumber="51">Sarawak</collectingRegion>
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,
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<collectingMunicipality box="[780,1107,1223,1249]" pageId="2" pageNumber="51">Taman Negara Bukit Lambir</collectingMunicipality>
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[
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<location LSID="urn:lsid:plazi:treatment:039487F4FF881D530BAAFB8AFD22F91C:8EE26039FF881D520BB4FB59FE0DFA9F" box="[146,462,1257,1284]" country="Malaysia" municipality="Taman Negara Bukit Lambir" name="Lambir Hills National Park" pageId="2" pageNumber="51" stateProvince="Sarawak">Lambir Hills National Park</location>
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],
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<date box="[480,614,1258,1285]" pageId="2" pageNumber="51" value="2005-09-16">
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<collectingDate box="[480,614,1258,1285]" pageId="2" pageNumber="51" value="2005-09-16">16.IX.2005</collectingDate>
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</date>
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, leg. ITIOKA-K.” Mouthparts, left legs and genitalia dissected and mounted on a small glass slide pinned under the specimen. Deposited at the Forest Department of
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<collectingRegion box="[557,653,1328,1354]" country="Malaysia" name="Sarawak" pageId="2" pageNumber="51">Sarawak</collectingRegion>
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, Malaysia. Six
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<typeStatus box="[822,928,1329,1355]" pageId="2" pageNumber="51">paratypes</typeStatus>
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with same data as holotype (deposited in the Kyushu University Museum).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="52" pageId="2" pageNumber="51" type="description">
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<paragraph blockId="2.[140,1108,1187,1672]" pageId="2" pageNumber="51">
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<emphasis bold="true" box="[185,335,1398,1424]" pageId="2" pageNumber="51">
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<emphasis box="[185,330,1398,1424]" inLineHeading="true" pageId="2" pageNumber="51" reason="1">Description</emphasis>
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.
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</emphasis>
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Length
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<quantity box="[445,626,1398,1425]" metricMagnitude="-3" metricUnit="m" metricValue="3.175" metricValueMax="3.3" metricValueMin="3.05" pageId="2" pageNumber="51" unit="mm" value="3.175" valueMax="3.3" valueMin="3.05">3.05–3.30 mm</quantity>
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.
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<emphasis box="[644,698,1399,1425]" italics="true" pageId="2" pageNumber="51">Body</emphasis>
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(Figs 1–2) elongate oval, reddish brown, glabrous shining.
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<emphasis box="[428,487,1434,1460]" italics="true" pageId="2" pageNumber="51">Head</emphasis>
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strongly gibbose medially; clypeal margin narrowly reflexed, broadly rounded on each side of shallow, median emargination; sides distinctly emarginate in front of right-angled, protruding genae; clypeal surface densely and finely punctate, with microreticulations.
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<emphasis box="[566,674,1540,1565]" italics="true" pageId="2" pageNumber="51">Pronotum</emphasis>
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rectangular, convex, side margin narrowly deplanate; posterior angles broadly rounded; surface punctures variably-sized, coarse and deep mesally, separated by 1–3 times their diameters.
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<taxonomicName box="[891,998,1610,1636]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="51" phylum="Arthropoda" rank="genus">
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<emphasis box="[891,998,1610,1636]" italics="true" pageId="2" pageNumber="51">Scutellum</emphasis>
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</taxonomicName>
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narrowly subtriangular.
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<emphasis box="[302,367,1645,1671]" italics="true" pageId="2" pageNumber="51">Elytra</emphasis>
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slightly wider than base of pronotum and about 2.3 times as long
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</paragraph>
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<paragraph blockId="3.[140,1107,1063,1149]" pageId="3" pageNumber="52">
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<emphasis bold="true" box="[140,275,1063,1085]" pageId="3" pageNumber="52">Figures 3–9.</emphasis>
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<taxonomicName authority="Maruyama, 2010" authorityName="Maruyama" authorityYear="2010" box="[282,429,1063,1084]" class="Insecta" family="Aphodiidae" genus="Pterobius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="species" species="itiokai" status="new genus and species">
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<emphasis box="[282,429,1063,1084]" italics="true" pageId="3" pageNumber="52">Pterobius itiokai</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel box="[442,651,1063,1085]" pageId="3" pageNumber="52" rank="species">new genus and species</taxonomicNameLabel>
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(
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<typeStatus box="[665,752,1063,1085]" pageId="3" pageNumber="52">holotype</typeStatus>
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).
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<emphasis bold="true" box="[768,781,1063,1085]" pageId="3" pageNumber="52">3</emphasis>
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epipharynx
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<emphasis bold="true" box="[900,913,1063,1085]" pageId="3" pageNumber="52">4</emphasis>
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left antenna, ventral view
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<emphasis bold="true" box="[188,201,1095,1117]" pageId="3" pageNumber="52">5</emphasis>
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foreleg, dorsal view
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<emphasis bold="true" box="[391,404,1095,1117]" pageId="3" pageNumber="52">6</emphasis>
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middle leg, dorsal view
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<emphasis bold="true" box="[629,642,1095,1117]" pageId="3" pageNumber="52">7</emphasis>
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hindleg, dorsal view
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<emphasis bold="true" box="[839,852,1095,1117]" pageId="3" pageNumber="52">8</emphasis>
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aedeagus, dorsal view
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<emphasis bold="true" box="[1061,1074,1095,1117]" pageId="3" pageNumber="52">9</emphasis>
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aedeagus, lateral view.
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</paragraph>
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<paragraph blockId="3.[140,1108,1187,1672]" pageId="3" pageNumber="52">
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as pronotum, epipleural margin slightly reflexed, forming obtuse tooth at shoulder; striae finely impressed, strial punctures coarse, deep; intervals slightly convex, smooth, impunctate. Ventral sclerites shining; mesosternum gibbose and smooth in anterior half, posterior half concave, finely and sparsely punctate, at middle with triangular, smooth convexity prolonged to metasternum; metasternum convex, midline fine; surface finely and sparsely punctate; abdominal sternites with fine fluting along sutures and smooth, impunctate surface including that of pygidium.
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<emphasis box="[833,878,1399,1424]" italics="true" pageId="3" pageNumber="52">Legs</emphasis>
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(Figs 5–7) short; all femora sparsely and finely punctate; profemur subquadrate; meso- and metafemora with posterior lines; metatarsus stout, 2/3 times as long as tibia.
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</paragraph>
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<paragraph blockId="3.[140,1108,1187,1672]" pageId="3" pageNumber="52">
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<emphasis box="[185,249,1504,1530]" italics="true" pageId="3" pageNumber="52">Male.</emphasis>
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Penultimate abdominal sternite and disc of pygidium shorter than female; aedeagus as in Figs 8–9.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="3" pageNumber="52" type="discussion">
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<paragraph blockId="3.[140,1108,1187,1672]" pageId="3" pageNumber="52">
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<emphasis bold="true" box="[185,298,1575,1601]" pageId="3" pageNumber="52">
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<emphasis box="[185,293,1575,1601]" inLineHeading="true" pageId="3" pageNumber="52" reason="1">Remarks</emphasis>
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.
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</emphasis>
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This species is similar to species of the genus
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<taxonomicName authorityName="Schmidt" authorityYear="1913" box="[828,987,1575,1601]" class="Insecta" family="Aphodiidae" genus="Cnematoplatys" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="52" phylum="Arthropoda" rank="genus">
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<emphasis box="[828,987,1575,1601]" italics="true" pageId="3" pageNumber="52">Cnematoplatys</emphasis>
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</taxonomicName>
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in general appearance but are easily distinguished from them by the margined elytra, the thick meso- and metatarsi. and the 4-segmented metatarsi.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |