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<document id="31F590843E71DA3A027CA6522C68CCE6" ID-CLB-Dataset="156941" ID-DOI="10.11646/zootaxa.5190.3.2" ID-GBIF-Dataset="d9bfc4ad-ca1f-40aa-a23a-8189507a249e" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138151" ID-ZooBank="FFD4A08E-404D-49BB-88C8-E89B497B54A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664791740547" checkinUser="plazi" docAuthor="Drew, R. A. I. &amp; Hancock, D. L." docDate="2022" docId="9F5F133EF82DB300FF3F5FA2FC3AF3C5" docLanguage="en" docName="zootaxa.5190.3.2.pdf" docOrigin="Zootaxa 5190 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bactrocera (Bactrocera) papayae Drew &amp; Hancock" docType="treatment" docVersion="7" lastPageNumber="351" masterDocId="63666B46F83FB312FFA85F35FF8DF73D" masterDocTitle="Biogeography, Speciation and Taxonomy within the genus Bactrocera Macquart with application to the Bactrocera dorsalis (Hendel) complex of fruit flies (Diptera Tephritidae: Dacinae)" masterLastPageNumber="360" masterPageNumber="333" pageNumber="351" updateTime="1699385890034" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E67EB1FFAF3D575E7C0BAB1F09C9D977">Biogeography, Speciation and Taxonomy within the genus Bactrocera Macquart with application to the Bactrocera dorsalis (Hendel) complex of fruit flies (Diptera Tephritidae: Dacinae)</mods:title>
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<mods:namePart id="0174D7F200D608914C35D8D637AF34FD">Drew, R. A. I.</mods:namePart>
<mods:affiliation id="737AC25F7A236008100D57970BF5F136">Environmental Futures Research Institute, International Centre for Management of Pest Fruit Flies, Griffith University, Nathan Campus, Brisbane, Queensland, Australia</mods:affiliation>
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<mods:namePart id="2D6EA576BD65A51E0AEEC307E5B9C527">Hancock, D. L.</mods:namePart>
<mods:affiliation id="50196E04008902C9BD2D33349D7A8D46">60 South Street, Carlisle, Cumbria CA 1 2 EP, United Kingdom.</mods:affiliation>
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<mods:date id="8E938C905FABF879A2ACA8832767727F">2022</mods:date>
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<mods:number id="EEA7F125C21BEA1ACCF0BAF8BF465FFA">5190</mods:number>
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<mods:identifier id="D5852A07CC272F5A6321BA2E42806F9C" type="DOI">10.11646/zootaxa.5190.3.2</mods:identifier>
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<subSubSection id="5FECF1A3F82DB300FF3F5FA2FC84F78C" box="[151,777,151,177]" pageId="18" pageNumber="351" type="nomenclature">
<paragraph id="1749A228F82DB300FF3F5FA2FC84F78C" blockId="18.[151,777,151,177]" box="[151,777,151,177]" pageId="18" pageNumber="351">
<heading id="4C011544F82DB300FF3F5FA2FC84F78C" bold="true" box="[151,777,151,177]" fontSize="11" level="1" pageId="18" pageNumber="351" reason="1">
<taxonomicName id="D0F6D9ABF82DB300FF3F5FA2FC84F78C" ID-CoL="87RD3" authority="Drew &amp; Hancock" authorityName="Drew &amp; Hancock" box="[151,777,151,177]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="papayae" subGenus="Bactrocera">
<emphasis id="25827E3AF82DB300FF3F5FA2FC84F78C" bold="true" box="[151,777,151,177]" pageId="18" pageNumber="351">
<emphasis id="25827E3AF82DB300FF3F5FA2FDA5F78C" bold="true" box="[151,552,151,177]" italics="true" pageId="18" pageNumber="351">Bactrocera (Bactrocera) papayae</emphasis>
Drew &amp; Hancock
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="1749A228F82DB300FF3F5FE9FA11F7CE" blockId="18.[151,1436,219,341]" box="[151,1436,219,244]" pageId="18" pageNumber="351">
<treatmentCitationGroup id="37E68506F82DB300FF3F5FE9FA11F7CE" box="[151,1436,219,244]" pageId="18" pageNumber="351">
<treatmentCitation id="96578439F82DB300FF3F5FE9FC9FF7C9" author="Drew, R. A. I. &amp; Hancock, D. L." box="[151,786,220,244]" page="48" pageId="18" pageNumber="351" year="1994">
<taxonomicName id="D0F6D9ABF82DB300FF3F5FE9FC9FF7C9" ID-CoL="87RD3" authority="Drew &amp; Hancock, 1994: 48" authorityName="Drew &amp; Hancock" authorityPageNumber="48" authorityYear="1994" box="[151,786,220,244]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="papayae" subGenus="Bactrocera">
<emphasis id="25827E3AF82DB300FF3F5FE9FE63F7CE" box="[151,494,220,243]" italics="true" pageId="18" pageNumber="351">Bactrocera (Bactrocera) papayae</emphasis>
<bibRefCitation id="7367DFD9F82DB300FE5C5FE9FC9FF7C9" author="Drew, R. A. I. &amp; Hancock, D. L." box="[500,786,220,244]" pageId="18" pageNumber="351" pagination="1 - 68" refId="ref21807" refString="Drew, R. A. I. &amp; Hancock, D. L. (1994) The Bactrocera dorsalis complex of fruit flies (Diptera: Tephritidae: Dacinae) in Asia. Bulletin of Entomological Research, Supplement 2, i - iii + 1 - 68. https: // doi. org / 10.1017 / S 1367426900000278" type="journal article" year="1994">Drew &amp; Hancock, 1994: 48</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="96578439F82DB300FCB55FE9FB93F7C9" author="Norrbom, A. L. &amp; Carroll, L. E. &amp; Thompson, F. C. &amp; White, I. M. &amp; Freidberg, A." box="[797,1054,220,244]" page="93" pageId="18" pageNumber="351" year="1998">
<bibRefCitation id="7367DFD9F82DB300FCB55FE9FB93F7C9" author="Norrbom, A. L. &amp; Carroll, L. E. &amp; Thompson, F. C. &amp; White, I. M. &amp; Freidberg, A." box="[797,1054,220,244]" pageId="18" pageNumber="351" pagination="65 - 251" refId="ref24108" refString="Norrbom, A. L., Carroll, L. E., Thompson, F. C., White, I. M. &amp; Freidberg, A. (1998) Systematic database of names. In: Thompson, F. C., (Ed.), Fruit Fly Expert Identification System and Systematic Information Database. Myia, 9, 65 - 251." type="journal article" year="1998">
Norrbom
<emphasis id="25827E3AF82DB300FC295FE9FC23F7C9" box="[897,942,220,244]" italics="true" pageId="18" pageNumber="351">et al</emphasis>
., 1998: 93
</bibRefCitation>
</treatmentCitation>
;
<treatmentCitation id="96578439F82DB300FB815FE9FAB0F7C9" author="Drew, R. A. I. &amp; Romig, M. C." box="[1065,1341,220,244]" page="142" pageId="18" pageNumber="351" year="2013">
<bibRefCitation id="7367DFD9F82DB300FB815FE9FAB0F7C9" author="Drew, R. A. I. &amp; Romig, M. C." box="[1065,1341,220,244]" pageId="18" pageNumber="351" refId="ref22177" refString="Drew, R. A. I. &amp; Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia, Indomalaya to North-West Australasia. CABI, Wallingford, 653 pp. https: // doi. org / 10.1079 / 9781780640358.0000" type="book" year="2013">Drew &amp; Romig, 2013: 142</bibRefCitation>
</treatmentCitation>
,
<treatmentCitation id="96578439F82DB300FAE15FEEFA15F7CE" author="Drew, R. A. I. &amp; Romig, M. C." box="[1353,1432,219,243]" page="7" pageId="18" pageNumber="351" year="2016">
<bibRefCitation id="7367DFD9F82DB300FAE15FEEFA15F7CE" author="Drew, R. A. I. &amp; Romig, M. C." box="[1353,1432,219,243]" pageId="18" pageNumber="351" refId="ref22229" refString="Drew, R. A. I. &amp; Romig, M. C. (2016) Keys to the tropical fruit flies (Tephritidae: Dacinae) of South-East Asia, Indomalaya to North-West Australasia. CABI, Wallingford, 485 pp. https: // doi. org / 10.1079 / 9781780644196.0000" type="book" year="2016">2016: 7</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
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<paragraph id="1749A228F82DB300FF6F5E35FE14F62A" blockId="18.[151,1436,219,341]" box="[199,409,256,280]" pageId="18" pageNumber="351">
<materialsCitation id="A79EA875F82DB300FF6F5E35FE14F62A" ID-GBIF-Occurrence="3920574304" box="[199,409,256,280]" collectionCode="BMNH" pageId="18" pageNumber="351" specimenCount="1" typeStatus="holotype">
<typeStatus id="C84D1C8AF82DB300FF6F5E35FEA8F625" box="[199,293,256,280]" pageId="18" pageNumber="351" type="holotype">Holotype</typeStatus>
in
<collectionCode id="71E73AEDF82DB300FEEE5E35FE18F62A" box="[326,405,256,279]" pageId="18" pageNumber="351">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="5FECF1A3F82DB300FF3F5E2AFDD2F6A5" pageId="18" pageNumber="351" type="reference_group">
<paragraph id="1749A228F82DB300FF3F5E2AFE2DF668" blockId="18.[151,1436,219,341]" pageId="18" pageNumber="351">
<treatmentCitationGroup id="37E68506F82DB300FF3F5E2AFE2DF668" pageId="18" pageNumber="351">
<treatmentCitation id="96578439F82DB300FF3F5E2AFCC5F60B" author="Drew, R. A. I. &amp; Hancock, D. L." box="[151,840,286,310]" page="52" pageId="18" pageNumber="351" year="1994">
<taxonomicName id="D0F6D9ABF82DB300FF3F5E2AFCC5F60B" ID-CoL="87RDL" authority="Drew &amp; Hancock, 1994: 52" authorityName="Drew &amp; Hancock" authorityPageNumber="52" authorityYear="1994" box="[151,840,286,310]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="philippinensis" subGenus="Bactrocera">
<emphasis id="25827E3AF82DB300FF3F5E2AFDA8F60B" box="[151,549,286,310]" italics="true" pageId="18" pageNumber="351">Bactrocera (Bactrocera) philippinensis</emphasis>
<bibRefCitation id="7367DFD9F82DB300FD845E2AFCC5F60B" author="Drew, R. A. I. &amp; Hancock, D. L." box="[556,840,286,310]" pageId="18" pageNumber="351" pagination="1 - 68" refId="ref21807" refString="Drew, R. A. I. &amp; Hancock, D. L. (1994) The Bactrocera dorsalis complex of fruit flies (Diptera: Tephritidae: Dacinae) in Asia. Bulletin of Entomological Research, Supplement 2, i - iii + 1 - 68. https: // doi. org / 10.1017 / S 1367426900000278" type="journal article" year="1994">Drew &amp; Hancock, 1994: 52</bibRefCitation>
</taxonomicName>
</treatmentCitation>
;
<treatmentCitation id="96578439F82DB300FCFB5E2BFBDEF60B" author="Norrbom, A. L. &amp; Carroll, L. E. &amp; Thompson, F. C. &amp; White, I. M. &amp; Freidberg, A." box="[851,1107,286,310]" page="94" pageId="18" pageNumber="351" year="1998">
<bibRefCitation id="7367DFD9F82DB300FCFB5E2BFBDEF60B" author="Norrbom, A. L. &amp; Carroll, L. E. &amp; Thompson, F. C. &amp; White, I. M. &amp; Freidberg, A." box="[851,1107,286,310]" pageId="18" pageNumber="351" pagination="65 - 251" refId="ref24108" refString="Norrbom, A. L., Carroll, L. E., Thompson, F. C., White, I. M. &amp; Freidberg, A. (1998) Systematic database of names. In: Thompson, F. C., (Ed.), Fruit Fly Expert Identification System and Systematic Information Database. Myia, 9, 65 - 251." type="journal article" year="1998">
Norrbom
<emphasis id="25827E3AF82DB300FC1F5E2AFC69F60B" box="[951,996,286,310]" italics="true" pageId="18" pageNumber="351">et al</emphasis>
., 1998: 94
</bibRefCitation>
</treatmentCitation>
.
<typeStatus id="C84D1C8AF82DB300FBF55E2BFB36F60B" box="[1117,1211,286,310]" pageId="18" pageNumber="351" type="holotype">Holotype</typeStatus>
in BPBM. Syn.
<treatmentCitation id="96578439F82DB300FACC5E2AFE11F668" author="Drew, R. A. I. &amp; Romig, M. C." page="142" pageId="18" pageNumber="351" year="2013">
<bibRefCitation id="7367DFD9F82DB300FACC5E2AFE11F668" author="Drew, R. A. I. &amp; Romig, M. C." pageId="18" pageNumber="351" refId="ref22177" refString="Drew, R. A. I. &amp; Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia, Indomalaya to North-West Australasia. CABI, Wallingford, 653 pp. https: // doi. org / 10.1079 / 9781780640358.0000" type="book" year="2013">Drew &amp; Romig, 2013: 142</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="1749A228F82DB300FF3F5E4BFDD2F6A5" blockId="18.[151,1437,382,1272]" box="[151,607,382,408]" pageId="18" pageNumber="351">Common Name: Asian Papaya Fruit Fly.</paragraph>
</subSubSection>
<subSubSection id="5FECF1A3F82DB300FF6F5E97FC3AF3C5" pageId="18" pageNumber="351" type="description">
<paragraph id="1749A228F82DB300FF6F5E97FD79F475" blockId="18.[151,1437,382,1272]" pageId="18" pageNumber="351">
Definition: Face fulvous with a pair of large oval black spots; postpronotal lobes and notopleura yellow; scutum black with dark brown laterally and around notopleural suture; broad parallel-sided lateral postsutural yellow vittae ending at or behind
<emphasis id="25827E3AF82DB300FED55EDEFE17F539" box="[381,410,491,516]" italics="true" pageId="18" pageNumber="351">ia.</emphasis>
seta; medial postsutural yellow vitta absent; anepisternal stripe reaching midway between anterior margin of notopleuron and anterior
<emphasis id="25827E3AF82DB300FD385D3BFD31F515" box="[656,700,526,552]" italics="true" pageId="18" pageNumber="351">npl.</emphasis>
seta dorsally; scutellum yellow with a narrow black basal band; legs with femora entirely fulvous, fore and hind tibiae dark fuscous, mid tibiae fuscous basally (darker stripe on ventral surface of fore tibiae more distinct on fresh specimens); wing with cells bc and c colourless, microtrichia in outer corner of cell c only, a narrow fuscous costal band confluent with R
<subScript id="8B72A06DF82DB300FC765DB9FC7AF5A7" attach="left" box="[990,1015,652,666]" fontSize="6" pageId="18" pageNumber="351">2+3</subScript>
and may widen slightly around apex of wing, a narrow fuscous anal streak, supernumerary lobe of medium development; abdominal terga III-V orangebrown with a narrow transverse black band across anterior margin of tergum III which expands laterally into narrow margins (in some specimens this band is broken in the midline), a narrow to medium width medial longitudinal black band over all three terga, anterolateral corners of terga IV and V dark fuscous to black, ceromata on tergum V orange-brown, abdominal terga dark fuscous to black.
</paragraph>
<paragraph id="1749A228F82DB300FF6F5C67FDD9F489" blockId="18.[151,1437,382,1272]" pageId="18" pageNumber="351">
Distribution: Peninsular
<collectingCountry id="6FE1E2B8F82DB300FE7E5C67FDB2F451" box="[470,575,850,876]" name="Malaysia" pageId="18" pageNumber="351">Malaysia</collectingCountry>
, East
<collectingCountry id="6FE1E2B8F82DB300FDD55C67FD6BF451" box="[637,742,850,876]" name="Malaysia" pageId="18" pageNumber="351">Malaysia</collectingCountry>
, Central to Southern
<collectingCountry id="6FE1E2B8F82DB300FC615C67FBA0F451" box="[969,1069,850,876]" name="Thailand" pageId="18" pageNumber="351">Thailand</collectingCountry>
(at least as far north as
<collectingRegion id="D5326CCAF82DB300FA8F5C67FA1CF451" box="[1319,1425,850,876]" country="Thailand" name="Krung Thep Mahanakhon" pageId="18" pageNumber="351">Bangkok</collectingRegion>
),
<collectingCountry id="6FE1E2B8F82DB300FF3F5C43FE95F4AD" box="[151,280,886,912]" name="Philippines" pageId="18" pageNumber="351">Philippines</collectingCountry>
,
<collectingCountry id="6FE1E2B8F82DB300FE8C5C43FEE8F4AD" box="[292,357,886,912]" name="Palau" pageId="18" pageNumber="351">Palau</collectingCountry>
, Borneo,
<collectingCountry id="6FE1E2B8F82DB300FE7B5C43FDCEF4AD" box="[467,579,886,912]" name="Indonesia" pageId="18" pageNumber="351">Indonesia</collectingCountry>
,
<collectingCountry id="6FE1E2B8F82DB300FDE75C43FD4EF4AD" box="[591,707,886,912]" name="Singapore" pageId="18" pageNumber="351">Singapore</collectingCountry>
,
<collectingCountry id="6FE1E2B8F82DB300FD675C43FCB2F4AD" box="[719,831,886,912]" name="Indonesia" pageId="18" pageNumber="351">Moluccas</collectingCountry>
, Indonesian
<collectingRegion id="D5326CCAF82DB300FC665C43FB98F4AD" box="[974,1045,886,912]" country="Indonesia" name="Papua Barat" pageId="18" pageNumber="351">Papua</collectingRegion>
,
<collectingCountry id="6FE1E2B8F82DB300FB895C43FB72F4AD" box="[1057,1279,886,912]" name="Papua New Guinea" pageId="18" pageNumber="351">Papua New Guinea</collectingCountry>
,
<collectingRegion id="D5326CCAF82DB300FAA35C43FA14F4AD" box="[1291,1433,886,912]" country="Papua New Guinea" name="West New Britain" pageId="18" pageNumber="351">New Britain</collectingRegion>
,
<collectingCountry id="6FE1E2B8F82DB300FF3F5CAFFED9F489" box="[151,340,922,948]" name="Christmas Island" pageId="18" pageNumber="351">Christmas Island</collectingCountry>
(Australian Territory).
</paragraph>
<paragraph id="1749A228F82DB300FF6F5C8BFB3DF4E5" blockId="18.[151,1437,382,1272]" box="[199,1200,958,984]" pageId="18" pageNumber="351">
Hosts: A wide range of commercial/edible and wild host fruits (see
<bibRefCitation id="7367DFD9F82DB300FC115C8BFB29F4E5" author="Allwood, A. J. &amp; Chinajariyawong A. &amp; Drew R. A. I. &amp; Hamacek, E. L. &amp; Hancock, D. L. &amp; Hengsawad, C. &amp; Jipanin, J. C. &amp; Jirasurat, M. &amp; Kong Krong, C. &amp; Kritsaneepaiboon, S. &amp; Leong, C. T. S. &amp; Vijaysegaran, S." box="[953,1188,958,984]" pageId="18" pageNumber="351" pagination="1 - 92" refId="ref20648" refString="Allwood, A. J., Chinajariyawong A., Drew R. A. I., Hamacek, E. L., Hancock, D. L., Hengsawad, C., Jipanin, J. C., Jirasurat, M., Kong Krong, C., Kritsaneepaiboon, S., Leong, C. T. S. &amp; Vijaysegaran, S. (1999) Host plant records for fruit flies (Diptera: Tephritidae) in South East Asia. The Raffles Bulletin of Zoology, Supplement 7, 1 - 92." type="journal article" year="1999">
Allwood
<emphasis id="25827E3AF82DB300FB8A5C8AFBDEF4E5" box="[1058,1107,958,984]" italics="true" pageId="18" pageNumber="351">et al</emphasis>
., 1999
</bibRefCitation>
).
</paragraph>
<paragraph id="1749A228F82DB300FF6F5CD7FE73F4C1" blockId="18.[151,1437,382,1272]" box="[199,510,994,1020]" pageId="18" pageNumber="351">Attractant: Methyl eugenol.</paragraph>
<paragraph id="1749A228F82DB300FF6F5B33FC3AF3C5" blockId="18.[151,1437,382,1272]" pageId="18" pageNumber="351">
Comments: Detailed discussion on the specific status of
<taxonomicName id="D0F6D9ABF82DB300FCF15B32FC40F31D" box="[857,973,1031,1056]" class="Magnoliopsida" family="Caricaceae" genus="Carica" kingdom="Plantae" order="Brassicales" pageId="18" pageNumber="351" phylum="Tracheophyta" rank="species" species="papaya">
<emphasis id="25827E3AF82DB300FCF15B32FC40F31D" box="[857,973,1031,1056]" italics="true" pageId="18" pageNumber="351">B. papaya</emphasis>
</taxonomicName>
e has been presented in
<bibRefCitation id="7367DFD9F82DB300FB405B33FF56F379" author="Drew, R. A. I. &amp; Romig, M. C." pageId="18" pageNumber="351" refId="ref22177" refString="Drew, R. A. I. &amp; Romig, M. C. (2013) Tropical fruit flies (Tephritidae: Dacinae) of South-East Asia, Indomalaya to North-West Australasia. CABI, Wallingford, 653 pp. https: // doi. org / 10.1079 / 9781780640358.0000" type="book" year="2013">Drew &amp; Romig (2013</bibRefCitation>
,
<bibRefCitation id="7367DFD9F82DB300FF405B1FFEAEF379" author="Drew, R. A. I. &amp; Romig, M. C." box="[232,291,1066,1092]" pageId="18" pageNumber="351" refId="ref22229" refString="Drew, R. A. I. &amp; Romig, M. C. (2016) Keys to the tropical fruit flies (Tephritidae: Dacinae) of South-East Asia, Indomalaya to North-West Australasia. CABI, Wallingford, 485 pp. https: // doi. org / 10.1079 / 9781780644196.0000" type="book" year="2016">2016</bibRefCitation>
,
<bibRefCitation id="7367DFD9F82DB300FE985B1FFEE0F379" author="Drew, R. A. I. &amp; Romig, M. C." box="[304,365,1066,1092]" pageId="18" pageNumber="351" refId="ref22284" refString="Drew, R. A. I. &amp; Romig, M. C. (2022) The fruit fly fauna (Diptera: Tephritidae: Dacinae) of Papua New Guinea, Indonesian Papua, associated islands and Bougainville. CABI, Wallingford, 124 pp. https: // doi. org / 10.1079 / 9781789249514.0000" type="book" year="2022">2022</bibRefCitation>
). The major host preferences of this species make it a serious biosecurity risk, particularly for specific export industries such as bananas. The closest species morphologically is
<taxonomicName id="D0F6D9ABF82DB300FBE95B7AFB36F355" authorityName="Hendel" authorityYear="1912" box="[1089,1211,1102,1128]" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="25827E3AF82DB300FBE95B7AFB36F355" box="[1089,1211,1102,1128]" italics="true" pageId="18" pageNumber="351">B. dorsalis</emphasis>
</taxonomicName>
and the two can be separated on the basis of aedeagus and ovipositor measurements, particularly those of host-reared specimens. In
<taxonomicName id="D0F6D9ABF82DB300FF3F5BA2FE9BF38D" box="[151,278,1175,1200]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="papayae">
<emphasis id="25827E3AF82DB300FF3F5BA2FE9BF38D" box="[151,278,1175,1200]" italics="true" pageId="18" pageNumber="351">B. papayae</emphasis>
</taxonomicName>
the aedeagus length range is
<quantity id="D00E0FCDF82DB300FDC95BA3FD7BF38D" box="[609,758,1174,1201]" metricMagnitude="-3" metricUnit="m" metricValue="2.9699999999999998" metricValueMax="3.4" metricValueMin="2.54" pageId="18" pageNumber="351" unit="mm" value="2.9699999999999998" valueMax="3.4" valueMin="2.54">2.54–3.4 mm</quantity>
and in
<taxonomicName id="D0F6D9ABF82DB300FCE15BA2FC4EF38D" authorityName="Hendel" authorityYear="1912" box="[841,963,1174,1200]" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="25827E3AF82DB300FCE15BA2FC4EF38D" box="[841,963,1174,1200]" italics="true" pageId="18" pageNumber="351">B. dorsalis</emphasis>
</taxonomicName>
<quantity id="D00E0FCDF82DB300FC625BA3FBECF38D" box="[970,1121,1174,1200]" metricMagnitude="-3" metricUnit="m" metricValue="2.58" metricValueMax="2.7" metricValueMin="2.46" pageId="18" pageNumber="351" unit="mm" value="2.58" valueMax="2.7" valueMin="2.46">2.46–2.7 mm</quantity>
, and in
<taxonomicName id="D0F6D9ABF82DB300FB125BA2FAB5F38D" box="[1210,1336,1175,1200]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="papayae">
<emphasis id="25827E3AF82DB300FB125BA2FAB5F38D" box="[1210,1336,1175,1200]" italics="true" pageId="18" pageNumber="351">B. papayae</emphasis>
</taxonomicName>
the ratio of the length of the oviscape to length of tergum V is 1: 1 to 1.5: 1, whereas in
<taxonomicName id="D0F6D9ABF82DB300FBA65B8EFB0AF3E9" authorityName="Hendel" authorityYear="1912" box="[1038,1159,1210,1236]" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="25827E3AF82DB300FBA65B8EFB0AF3E9" box="[1038,1159,1210,1236]" italics="true" pageId="18" pageNumber="351">B. dorsalis</emphasis>
</taxonomicName>
it is 0.7: 1 to 0.8: 1. The glans is elongate and tubular in
<taxonomicName id="D0F6D9ABF82DB300FE535BEAFDF4F3C5" box="[507,633,1247,1272]" class="Insecta" family="Tephritidae" genus="Bactrocera" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="papayae">
<emphasis id="25827E3AF82DB300FE535BEAFDF4F3C5" box="[507,633,1247,1272]" italics="true" pageId="18" pageNumber="351">B. papayae</emphasis>
</taxonomicName>
and sobovate in
<taxonomicName id="D0F6D9ABF82DB300FC9F5BEAFC3DF3C5" authorityName="Hendel" authorityYear="1912" box="[823,944,1246,1272]" class="Insecta" family="Tephritidae" genus="Dacus" kingdom="Animalia" order="Diptera" pageId="18" pageNumber="351" phylum="Arthropoda" rank="species" species="dorsalis">
<emphasis id="25827E3AF82DB300FC9F5BEAFC3DF3C5" box="[823,944,1246,1272]" italics="true" pageId="18" pageNumber="351">B. dorsalis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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