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<document id="CABA7F31C70A40FCE757B618C2F5454B" ID-CLB-Dataset="45970" ID-DOI="10.11646/zootaxa.3790.1.6" ID-GBIF-Dataset="bc52b393-d1b1-43c3-9491-d3b9d47ef9b0" ID-ISSN="1175-5326" ID-Zenodo-Dep="226715" ID-ZooBank="87AB27EC-DC05-48F3-8AB7-5C317B275AF5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460410436114" checkinUser="plazi" docAuthor="Sueyoshi, Masahiro" docDate="2014" docId="49168794FFE0564737ADFAFE239F372F" docLanguage="en" docName="zt03790p164.pdf" docOrigin="Zootaxa 3790 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoempheria ferruginea Brunetti 1912" docType="treatment" docVersion="9" lastPageNumber="141" masterDocId="B52FFFECFFE15644373AFFCC222B360A" masterDocTitle="Taxonomy of fungus gnats allied to Neoempheria ferruginea (Brunetti, 1912) (Diptera: Mycetophilidae), with descriptions of 11 new species from Japan and adjacent areas" masterLastPageNumber="164" masterPageNumber="139" pageNumber="140" updateTime="1698300419586" updateUser="ExternalLinkService">
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<mods:title id="CCF2B6F7181598C952A1E3C86900BC49">Taxonomy of fungus gnats allied to Neoempheria ferruginea (Brunetti, 1912) (Diptera: Mycetophilidae), with descriptions of 11 new species from Japan and adjacent areas</mods:title>
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<mods:namePart id="A103826E9F4A3035E7A13AE164B65FE4">Sueyoshi, Masahiro</mods:namePart>
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<mods:date id="24CFE47B33F2E5ECA8C1294CA8809694">2014</mods:date>
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<mods:identifier id="775825C76E7424F86AF95CB45D4D29FB" type="DOI">10.11646/zootaxa.3790.1.6</mods:identifier>
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<treatment id="49168794FFE0564737ADFAFE239F372F" ID-DOI="http://doi.org/10.5281/zenodo.6138784" ID-GBIF-Taxon="119423263" ID-Zenodo-Dep="6138784" LSID="urn:lsid:plazi:treatment:49168794FFE0564737ADFAFE239F372F" httpUri="http://treatment.plazi.org/id/49168794FFE0564737ADFAFE239F372F" lastPageId="3" lastPageNumber="141" pageId="1" pageNumber="140">
<subSubSection id="89A56509FFE0564537ADFAFE23253365" pageId="1" pageNumber="140" type="nomenclature">
<paragraph id="C1003682FFE0564537ADFAFE20523341" blockId="1.[151,633,1330,1391]" box="[151,633,1330,1355]" pageId="1" pageNumber="140">
<heading id="9A4881EEFFE0564537ADFAFE20523341" bold="true" box="[151,633,1330,1355]" fontSize="10" level="2" pageId="1" pageNumber="140" reason="2">
<taxonomicName id="06BF4D01FFE0564537ADFAFE20523341" ID-CoL="46GVZ" authority="Brunetti, 1912" authorityName="Brunetti" authorityYear="1912" box="[151,633,1330,1355]" class="Insecta" family="Mycetophilidae" genus="Neoempheria" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="F3CBEA90FFE0564537ADFAFE20523341" bold="true" box="[151,633,1330,1355]" pageId="1" pageNumber="140">
<emphasis id="F3CBEA90FFE0564537ADFAFE239F3341" bold="true" box="[151,436,1330,1355]" italics="true" pageId="1" pageNumber="140">Neoempheria ferruginea</emphasis>
(
<bibRefCitation id="A52E4B73FFE0564536F9FAFE205A3341" author="Brunetti" box="[451,625,1330,1355]" pageId="1" pageNumber="140" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 581, pls 1 - 12." type="book chapter" year="1912">Brunetti, 1912</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="C1003682FFE0564537ADFA9A23253365" blockId="1.[151,633,1330,1391]" box="[151,270,1366,1391]" pageId="1" pageNumber="140">(Fig. 1a–f)</paragraph>
</subSubSection>
<subSubSection id="89A56509FFE0564537ADFA6C200533D9" pageId="1" pageNumber="140" type="reference_group">
<paragraph id="C1003682FFE0564537ADFA6C200533D9" blockId="1.[151,805,1438,1491]" pageId="1" pageNumber="140">
<treatmentCitationGroup id="E1AF11ACFFE0564537ADFA6C200533D9" pageId="1" pageNumber="140">
<taxonomicName id="06BF4D01FFE0564537ADFA6C200733BE" ID-CoL="44X2W" authority="Brunetti, 1912: 74" authorityName="Brunetti" authorityPageNumber="74" authorityYear="1912" box="[151,556,1438,1461]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="140" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="F3CBEA90FFE0564537ADFA6C234133BE" box="[151,362,1438,1461]" italics="true" pageId="1" pageNumber="140">Mycomya ferruginea</emphasis>
<treatmentCitation id="401E1093FFE05645364BFA52200733BE" author="Brunetti" box="[369,556,1438,1460]" page="74" pageId="1" pageNumber="140" year="1912">
<bibRefCitation id="A52E4B73FFE05645364BFA52202D33BE" author="Brunetti" box="[369,518,1438,1460]" pageId="1" pageNumber="140" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 581, pls 1 - 12." type="book chapter" year="1912">Brunetti, 1912</bibRefCitation>
: 74
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="401E1093FFE05645350DFA52210A33BE" author="Senior-White" box="[567,801,1438,1460]" page="84" pageId="1" pageNumber="140" year="1922">
<bibRefCitation id="A52E4B73FFE05645350DFA5220D033BE" author="Senior-White" box="[567,763,1438,1460]" pageId="1" pageNumber="140" refString="Senior-White, R. (1922) Notes on Indian Diptera. Memoirs of the Department of Agriculture in India (Entomological series), 7 (9), 83 - 169." type="journal article" year="1922">Senior-White 1922</bibRefCitation>
: 84
</treatmentCitation>
. [Japanese name: nagamado-kinoko-bae]
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="89A56509FFE0564637ADF9CC229530D1" lastPageId="2" lastPageNumber="141" pageId="1" pageNumber="140" type="description">
<paragraph id="C1003682FFE0564537ADF9CC268230A3" blockId="1.[151,1437,1536,2031]" pageId="1" pageNumber="140">
<emphasis id="F3CBEA90FFE0564537ADF9CC230A3013" bold="true" box="[151,289,1536,1561]" pageId="1" pageNumber="140">Description</emphasis>
.
<emphasis id="F3CBEA90FFE056453608F9CC23443013" box="[306,367,1536,1561]" italics="true" pageId="1" pageNumber="140">Head</emphasis>
(Fig. 1a): yellow in ground color, with dark brown mark on occiput and dark brown stripes along occipital and frontal furrows. Frons bare. Lateral ocelli on black well-developed ocellar prominence. Ocellar setae absent. Face yellow. Mouthparts short. Clypeus yellow, triangular, with scattered setae at apical margin. Palpus dark brown. Antenna 4.5 times as long as head, shorter than head and thorax together. Scape and pedicel yellow. Pedicel with single seta slightly shorter than 1st flagellomere. Flagellum dark brown.
</paragraph>
<paragraph id="C1003682FFE0564537FDF97821C330FA" blockId="1.[151,1437,1536,2031]" pageId="1" pageNumber="140">
<emphasis id="F3CBEA90FFE0564537FDF978233330C7" box="[199,280,1716,1741]" italics="true" pageId="1" pageNumber="140">Thorax</emphasis>
(Fig. 1b): yellow in ground color. Scutum with 5 dark brown stripes (1 medial, 2 sublateral and 2 lateral). Setae present on each stripe. Scutellum with 2 pairs of lateral setae.
</paragraph>
<paragraph id="C1003682FFE0564537FDF93120C43155" blockId="1.[151,1437,1536,2031]" pageId="1" pageNumber="140">
<emphasis id="F3CBEA90FFE0564537FDF93122D4311E" box="[199,255,1789,1812]" italics="true" pageId="1" pageNumber="140">Wing</emphasis>
(Fig. 1c): hyaline, tinged with dark brown on apical 1/4 and posterior margin, distinct dark brown mark on each of veins Rs and R4. Vein sc-r ending in basal 1/4 of anterior margin of cell r1. Cell r1 4 times as long as wide. Halter yellow, tinged with dark yellow in knob.
</paragraph>
<paragraph id="C1003682FFE0564537FDF8A02641318E" blockId="1.[151,1437,1536,2031]" box="[199,1130,1899,1924]" pageId="1" pageNumber="140">
<emphasis id="F3CBEA90FFE0564537FDF8A022D73189" box="[199,252,1900,1923]" italics="true" pageId="1" pageNumber="140">Legs</emphasis>
(Fig. 1d): yellow. Tibia with erect setae slightly shorter than diameter of tibia.
</paragraph>
<paragraph id="C1003682FFE0564537FDF84223F231E5" blockId="1.[151,1437,1536,2031]" pageId="1" pageNumber="140">
<emphasis id="F3CBEA90FFE0564537FDF842231A31AD" box="[199,305,1934,1959]" italics="true" pageId="1" pageNumber="140">Abdomen</emphasis>
(Fig. 1b): yellow in ground color, with dark brown stripes on each abdominal tergite. Male genitalia (Fig. 1e): yellow in ground color. T9 with dorsal projection flattened dorsoventrally, bearing numerous setae on ventral side, tapered to apex.
</paragraph>
<paragraph id="C1003682FFE3564637ADFA10263A305A" blockId="2.[151,1436,1499,1620]" pageId="2" pageNumber="141">
<emphasis id="F3CBEA90FFE3564637ADFA10211633F8" bold="true" box="[151,829,1499,1522]" pageId="2" pageNumber="141">
FIGURE 1.
<typeStatus id="1E048820FFE356463622FA1023A833F8" box="[280,387,1500,1522]" pageId="2" pageNumber="141" type="lectotype">Lectotype</typeStatus>
of
<taxonomicName id="06BF4D01FFE35646369EFA10213433F8" authority="Brunetti, 1912" authorityName="Brunetti" authorityYear="1912" box="[420,799,1499,1522]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="ferruginea">
<emphasis id="F3CBEA90FFE35646369EFA10205533FB" bold="true" box="[420,638,1499,1521]" italics="true" pageId="2" pageNumber="141">Mycomya ferruginea</emphasis>
<bibRefCitation id="A52E4B73FFE3564635BEFA10213433F8" author="Brunetti" box="[644,799,1500,1522]" pageId="2" pageNumber="141" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 581, pls 1 - 12." type="book chapter" year="1912">Brunetti, 1912</bibRefCitation>
</taxonomicName>
. a,
</emphasis>
head in anterior view.
<emphasis id="F3CBEA90FFE356463318FA10261C33F8" bold="true" box="[1058,1079,1500,1522]" pageId="2" pageNumber="141">b.</emphasis>
thorax in dorsal view.
<emphasis id="F3CBEA90FFE356463226FA10270533F8" bold="true" box="[1308,1326,1500,1522]" pageId="2" pageNumber="141">c.</emphasis>
right wing in ventral view.
<emphasis id="F3CBEA90FFE356463603FA362366301A" bold="true" box="[313,333,1530,1552]" pageId="2" pageNumber="141">d.</emphasis>
imago in right lateral view.
<emphasis id="F3CBEA90FFE356463553FA362051301A" bold="true" box="[617,634,1530,1552]" pageId="2" pageNumber="141">e.</emphasis>
apex of abdomen in posterior view.
<emphasis id="F3CBEA90FFE3564634D3FA3621D2301A" bold="true" box="[1001,1017,1530,1552]" pageId="2" pageNumber="141">f.</emphasis>
labels. Note that the original photo of the wing was horizontally flipped. Abbreviations: dp, dorsal projection of of tergite 9; R4 and R5, 4th and 5th radial veins; r1 and r4, 1st and 4th radial cells; cu
<emphasis id="F3CBEA90FFE35646369AF9F72386305A" box="[416,429,1595,1616]" italics="true" pageId="2" pageNumber="141">p</emphasis>
, posterior branch of cubital vein; a1, 1st branch of anal vein.
</paragraph>
<paragraph id="C1003682FFE3564637FDF9B6229530D1" blockId="2.[151,1437,1658,2007]" pageId="2" pageNumber="141">
<emphasis id="F3CBEA90FFE3564637FDF9B623CC3099" bold="true" box="[199,487,1658,1683]" pageId="2" pageNumber="141">Photographs examined.</emphasis>
<typeStatus id="1E048820FFE3564636CEF9B7204C3099" box="[500,615,1659,1683]" pageId="2" pageNumber="141" type="lectotype">Lectotype</typeStatus>
, here designated.
<collectingCountry id="B9A87612FFE356463479F9B621BC3098" box="[835,919,1658,1682]" name="India" pageId="2" pageNumber="141">INDIA</collectingCountry>
:
<specimenCount id="D7B9FD0BFFE356463492F9B621E73098" box="[936,972,1658,1682]" pageId="2" pageNumber="141" type="generic">1♂</specimenCount>
, “Kurseong/ 5000 Feet [=
<quantity id="06479B67FFE356463222F9B6274C3099" box="[1304,1383,1658,1683]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="141" unit="m" value="1500.0">1500m</quantity>
]/ E. Himalayas/
<date id="B5011042FFE356463627F95323AF30BD" box="[285,388,1695,1719]" pageId="2" pageNumber="141" value="2008-07-03">3-VII-08</date>
”, “Myco/
<taxonomicName id="06BF4D01FFE3564636CCF952204230BD" box="[502,617,1694,1719]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="ferruginea">ferruginea</taxonomicName>
/ Brun.
<typeStatus id="1E048820FFE356463587F95320DE30BD" box="[701,757,1695,1719]" pageId="2" pageNumber="141">Type</typeStatus>
♂”, “Myco/
<taxonomicName id="06BF4D01FFE3564634BCF95221D230BD" box="[902,1017,1694,1719]" class="Insecta" family="Mycetophilidae" genus="Mycomya" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="141" phylum="Arthropoda" rank="species" species="ferruginea">ferruginea</taxonomicName>
/ ♂”, “
<typeStatus id="1E048820FFE35646337AF95326A730BD" box="[1088,1164,1695,1719]" pageId="2" pageNumber="141">TYPE</typeStatus>
”, “1904/ 20” (ZSI) (Fig. 1f).
</paragraph>
</subSubSection>
<subSubSection id="89A56509FFE3564637FDF92A20373128" pageId="2" pageNumber="141" type="distribution">
<paragraph id="C1003682FFE3564637FDF92A20373128" blockId="2.[151,1437,1658,2007]" pageId="2" pageNumber="141">
<emphasis id="F3CBEA90FFE3564637FDF92A237230F5" bold="true" box="[199,345,1766,1791]" pageId="2" pageNumber="141">Distribution</emphasis>
.
<collectingCountry id="B9A87612FFE35646364EF92A238530F5" box="[372,430,1766,1791]" name="India" pageId="2" pageNumber="141">India</collectingCountry>
(Darjeeling, Kolkata, Shillong) (
<bibRefCitation id="A52E4B73FFE35646346EF92A262A30F5" author="Brunetti" box="[852,1025,1766,1791]" pageId="2" pageNumber="141" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 581, pls 1 - 12." type="book chapter" year="1912">Brunetti 1912</bibRefCitation>
;
<bibRefCitation id="A52E4B73FFE356463320F92A272A30F5" author="Senior-White" box="[1050,1281,1766,1791]" pageId="2" pageNumber="141" refString="Senior-White, R. (1922) Notes on Indian Diptera. Memoirs of the Department of Agriculture in India (Entomological series), 7 (9), 83 - 169." type="journal article" year="1922">Senior-White 1922</bibRefCitation>
),
<collectingCountry id="B9A87612FFE35646321BF92A27B730F5" box="[1313,1436,1766,1791]" name="Sri Lanka" pageId="2" pageNumber="141">Sri Lanka</collectingCountry>
(Peradeniya) (
<bibRefCitation id="A52E4B73FFE35646360EF8C6203B3128" author="Senior-White" box="[308,528,1802,1827]" pageId="2" pageNumber="141" refString="Senior-White, R. (1922) Notes on Indian Diptera. Memoirs of the Department of Agriculture in India (Entomological series), 7 (9), 83 - 169." type="journal article" year="1922">Senior-White 1922</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="89A56509FFE3564737FDF8E2239F372F" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141" type="discussion">
<paragraph id="C1003682FFE3564637FDF8E2202831B9" blockId="2.[151,1437,1658,2007]" pageId="2" pageNumber="141">
<emphasis id="F3CBEA90FFE3564637FDF8E2231F314D" bold="true" box="[199,308,1838,1863]" pageId="2" pageNumber="141">Remarks</emphasis>
.
<bibRefCitation id="A52E4B73FFE35646367DF8E223D1314D" author="Brunetti" box="[327,506,1838,1863]" pageId="2" pageNumber="141" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 581, pls 1 - 12." type="book chapter" year="1912">Brunetti (1912)</bibRefCitation>
described this species based on two males and one female, without explicitly designating a
<typeStatus id="1E048820FFE35646360CF89E23B03161" box="[310,411,1874,1899]" pageId="2" pageNumber="141" type="holotype">holotype</typeStatus>
, though the specimen from Kurseong was labeled as
<typeStatus id="1E048820FFE35646333BF89F261A3161" box="[1025,1073,1875,1899]" pageId="2" pageNumber="141">type</typeStatus>
when examined later (
<bibRefCitation id="A52E4B73FFE356463202F89E22F83185" author="Edwards" pageId="2" pageNumber="141" refString="Edwards, F. W. (1924) Notes on the types of Diptera Nematocera (Mycetophilidae and Tipulidae) described by Mr. E. Brunetti. Records of Indian Museum, 26, 291 - 307." type="journal article" year="1924">Edwards 1924</bibRefCitation>
). ZSI holds only a single male from Kurseong with
<typeStatus id="1E048820FFE356463420F8BB21613185" box="[794,842,1911,1935]" pageId="2" pageNumber="141">type</typeStatus>
labels (Fig. 1f) (Dr. S. Sheela, pers. comm.) and it is designated here as the
<typeStatus id="1E048820FFE3564636AEF85623D431B9" box="[404,511,1946,1971]" pageId="2" pageNumber="141" type="lectotype">lectotype</typeStatus>
.
</paragraph>
<paragraph id="C1003682FFE3564737FDF873239F372F" blockId="2.[151,1437,1658,2007]" lastBlockId="3.[151,1436,151,293]" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141">
As
<bibRefCitation id="A52E4B73FFE3564637D5F872238F31DD" author="Edwards" box="[239,420,1982,2007]" pageId="2" pageNumber="141" refString="Edwards, F. W. (1924) Notes on the types of Diptera Nematocera (Mycetophilidae and Tipulidae) described by Mr. E. Brunetti. Records of Indian Museum, 26, 291 - 307." type="journal article" year="1924">Edwards (1924)</bibRefCitation>
noted, the wing marks of the
<typeStatus id="1E048820FFE35646343BF872214331DD" box="[769,872,1982,2007]" pageId="2" pageNumber="141" type="lectotype">lectotype</typeStatus>
(Fig. 1c) are different from figure 12 of
<bibRefCitation id="A52E4B73FFE35647327AF87222CA36BA" author="Brunetti" lastPageId="3" lastPageNumber="142" pageId="2" pageNumber="141" refString="Brunetti, E. (1912) Diptera Nematocera (excluding Chironomidae and Culicidae). In: Shipley, A. E. &amp; Marshall, G. A. K. (Eds.), Fauna of British India, including Ceylon and Burma. Taylor and Francis, London, pp. 1 - 581, pls 1 - 12." type="book chapter" year="1912">Brunetti (1912)</bibRefCitation>
in the following points: the wing anterior to veins M and R4+5 is hyaline and has distinct dark marks on veins Rs and R4; the dark brown mark in cell r4 is continuous to cell cua1 via the apex of cell r5, all of cell m1, and the apical portion of cell m; the brown mark in cell m1 fills the entire cell; and the brown mark on cell cu
<emphasis id="F3CBEA90FFE2564733C3FF2A272C36F7" box="[1273,1287,230,253]" italics="true" pageId="3" pageNumber="142">p</emphasis>
expands into the apical half of the cell.
</paragraph>
</subSubSection>
</treatment>
</document>
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