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<document id="FB41C11CDC7E67847796D83B88EDC0D1" ID-DOI="10.11646/phytotaxa.501.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="5424848" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1630392910242" checkinUser="marcus" docAuthor="Scatigna, André Vito &amp; Souza, Vinicius Castro" docDate="2021" docId="03D287E8706DFFCDF4A14766FD06FADB" docLanguage="en" docName="phytotaxa.501.2.5.pdf" docOrigin="Phytotaxa 501 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.501.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Angelonia perennis Scatigna &amp; V. C. Souza 2021, comb. nov." docType="treatment" docVersion="1" lastPageNumber="298" masterDocId="FFEBFF90706DFFCCF4294013FF9FFFE2" masterDocTitle="Tying up loose ends: The synonymization of Monopera with Angelonia (Plantaginaceae), including a new combination and typifications" masterLastPageNumber="299" masterPageNumber="297" pageNumber="297" updateTime="1732203349186" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="230FCFA2D76274FE774C73AB96F560BD">Tying up loose ends: The synonymization of Monopera with Angelonia (Plantaginaceae), including a new combination and typifications</mods:title>
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<mods:namePart id="1A51A0BCA015E03B340AD6548DD7D092">Scatigna, André Vito</mods:namePart>
<mods:affiliation id="371AAC4E49F7361E1C72EE310CE658BE">Centro de Educação, Ciências Exatas e Naturais, Departamento de Biologia, Universidade Estadual do Maranhão, Cidade Universitária Paulo VI, s / n, 65054 - 970, São Luís, Maranhão, Brazil.</mods:affiliation>
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<mods:namePart id="BEB28717000EFC683D65595E67332559">Souza, Vinicius Castro</mods:namePart>
<mods:affiliation id="7DC62DF2D03AA45C33B39E16E0D595D8">Herbário ESA, Departamento de Ciências Biológicas, Escola Superior de Agricultura Luiz de Queiroz, Universidade de São Paulo, Av. Pádua Dias, 11, Agronomia, 13418 - 400, Piracicaba, São Paulo, Brazil.</mods:affiliation>
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<emphasis id="B90FEAEC706DFFCCF4A14766FEFEF86F" bold="true" box="[136,353,1909,1933]" italics="true" pageId="0" pageNumber="297">Angelonia perennis</emphasis>
(
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,
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<emphasis id="B90FEAEC706DFFCCF7934766FBB1F86F" box="[954,1070,1909,1933]" italics="true" pageId="0" pageNumber="297">comb. nov.</emphasis>
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<taxonomicName id="4C7B4D7D706DFFCCF01F4766FE57F852" authority="Chodat &amp; Hassler (1904: 285)" authorityName=", Chodat &amp; Hassler" authorityPageNumber="285" authorityYear="1904" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="variety" species="micrantha" variety="perennis">
<emphasis id="B90FEAEC706DFFCCF01F4766FA86F86F" box="[1078,1305,1909,1933]" italics="true" pageId="0" pageNumber="297">Angelonia micrantha</emphasis>
var.
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.
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(Chodat &amp; Hassl.)
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.
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<figureCitation id="13402A7B706CFFCDF4C240D1FEA0FF38" box="[235,319,194,218]" captionStart="FIGURE 1" captionStartId="1.[136,229,848,870]" captionTargetBox="[151,1435,244,824]" captionTargetId="figure-38@1.[151,1435,244,824]" captionTargetPageId="1" captionText="FIGURE 1. A. Flower of Angelonia perennis in front view showing a bisaccate corolla (white arrow). B. Flower of A. micrantha in lateral view. Photos: A by André V. Scatigna, B by Ianne C.S. Nobre." figureDoi="http://doi.org/10.5281/zenodo.5424850" httpUri="https://zenodo.org/record/5424850/files/figure.png" pageId="1" pageNumber="298">Fig. 1A</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF046676706CFFCDF4A14343FD42FC68" ID-DOI="http://doi.org/10.5281/zenodo.5424850" ID-Zenodo-Dep="5424850" httpUri="https://zenodo.org/record/5424850/files/figure.png" pageId="1" pageNumber="298" startId="1.[136,229,848,870]" targetBox="[151,1435,244,824]" targetPageId="1">
<paragraph id="8BC436FE706CFFCDF4A14343FD42FC68" blockId="1.[136,1451,848,906]" pageId="1" pageNumber="298">
<emphasis id="B90FEAEC706CFFCDF4A14343FF62FC87" bold="true" box="[136,253,848,870]" pageId="1" pageNumber="298">FIGURE 1.</emphasis>
A. Flower of
<taxonomicName id="4C7B4D7D706CFFCDF5A84343FDA4FC84" authorityName="Scatigna &amp; V. C. Souza" authorityYear="2021" baseAuthorityName="Chodat &amp; Hassler" baseAuthorityYear="1904" box="[385,571,848,870]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="perennis">
<emphasis id="B90FEAEC706CFFCDF5A84343FDA4FC84" box="[385,571,848,870]" italics="true" pageId="1" pageNumber="298">Angelonia perennis</emphasis>
</taxonomicName>
in front view showing a bisaccate corolla (white arrow). B. Flower of
<taxonomicName id="4C7B4D7D706CFFCDF0FF4343FACCFC87" authorityName=", Bentham" authorityYear="1846" box="[1238,1363,848,869]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="micrantha">
<emphasis id="B90FEAEC706CFFCDF0FF4343FACCFC87" box="[1238,1363,848,869]" italics="true" pageId="1" pageNumber="298">A. micrantha</emphasis>
</taxonomicName>
in lateral view. Photos: A by André V. Scatigna, B by Ianne C.S. Nobre.
</paragraph>
</caption>
<subSubSection id="C3616575706CFFCDF49443A9FD06FADB" pageId="1" pageNumber="298" type="description">
<paragraph id="8BC436FE706CFFCDF49443A9FD06FADB" blockId="1.[136,1452,953,1697]" pageId="1" pageNumber="298">
In the protologue of
<taxonomicName id="4C7B4D7D706CFFCDF58B43AAFBA1FC33" authority=", Chodat &amp; Hassler (1904)" authorityName=", Chodat &amp; Hassler" authorityYear="1904" box="[418,1086,953,977]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="variety" species="micrantha" variety="perennis">
<emphasis id="B90FEAEC706CFFCDF58B43AAFD19FC33" box="[418,646,953,977]" italics="true" pageId="1" pageNumber="298">Angelonia micrantha</emphasis>
var.
<emphasis id="B90FEAEC706CFFCDF69643A9FC85FC33" box="[703,794,954,977]" italics="true" pageId="1" pageNumber="298">perennis</emphasis>
,
<bibRefCitation id="EFEA4B0F706CFFCDF70343AAFBA1FC33" author="Chodat, R. &amp; Hassler, E." box="[810,1086,953,977]" pageId="1" pageNumber="298" pagination="475 - 489" refId="ref1646" refString="Chodat, R. &amp; Hassler, E. (1904) Plantae hasslerianae. Bulletin de l'Herbier Boissier 2: 475 - 489." type="journal article" year="1904">Chodat &amp; Hassler (1904)</bibRefCitation>
</taxonomicName>
cited the specimen
<emphasis id="B90FEAEC706CFFCDF13343AAFA34FC33" box="[1306,1451,953,977]" italics="true" pageId="1" pageNumber="298">Hassler 7828</emphasis>
without assigning it to any herbarium. Later,
<bibRefCitation id="EFEA4B0F706CFFCDF64343CEFC80FC17" author="Barringer, K." box="[618,799,989,1013]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
cited “
<typeStatus id="54C0885C706CFFCDF74343CEFC4CFC17" box="[874,979,989,1013]" pageId="1" pageNumber="298" type="holotype">Holotype</typeStatus>
: G!; Isotypes: K!, MO!, NY!, P!, UC!, W!”. More recently,
<bibRefCitation id="EFEA4B0F706CFFCDF5064412FDDEFBFB" author="Souza, V. C. &amp; Giulietti, A. M." box="[303,577,1025,1049]" pageId="1" pageNumber="298" pagination="7 - 288" refId="ref1829" refString="Souza, V. C. &amp; Giulietti, A. M. (2009) Levantamento das especies de Scrophulariaceae sensu lato nativas do Brasil. Pesquisas Botanica 60: 7 - 288." type="journal article" year="2009">Souza &amp; Giulietti (2009)</bibRefCitation>
designated the specimen at G as
<typeStatus id="54C0885C706CFFCDF79E4412FB83FBFB" box="[951,1052,1025,1049]" pageId="1" pageNumber="298" type="lectotype">lectotype</typeStatus>
. However, because no
<typeStatus id="54C0885C706CFFCDF1324412FAE7FBFB" box="[1307,1400,1025,1049]" pageId="1" pageNumber="298" type="holotype">holotype</typeStatus>
was designated in the protologue and considering that the authors had access to all extant specimens of
<emphasis id="B90FEAEC706CFFCDF0EB4436FACDFBDF" box="[1218,1362,1061,1085]" italics="true" pageId="1" pageNumber="298">Hassler 7828</emphasis>
prior to publication, since Hassler is the gathering collector, the specimens in different herbaria are
<typeStatus id="54C0885C706CFFCDF0124459FB04FB80" box="[1083,1179,1098,1122]" pageId="1" pageNumber="298" type="syntype">syntypes</typeStatus>
, and the use of “
<typeStatus id="54C0885C706CFFCDF169445AFA3DFB83" box="[1344,1442,1097,1121]" pageId="1" pageNumber="298" type="holotype">holotype</typeStatus>
” by
<bibRefCitation id="EFEA4B0F706CFFCDF480447EFEC1FB67" author="Barringer, K." box="[169,350,1133,1157]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
is an error to be corrected to “
<typeStatus id="54C0885C706CFFCDF68B447EFC97FB67" box="[674,776,1133,1157]" pageId="1" pageNumber="298" type="lectotype">lectotype</typeStatus>
” according to Art. 9.10. Moreover, this inadvertent designation is effective, fulfilling Art. 7.11, and must be followed according to Art. 10.5, rendering the subsequent lectotypification by
<bibRefCitation id="EFEA4B0F706CFFCDF4A144A6FE0AFB2F" author="Souza, V. C. &amp; Giulietti, A. M." box="[136,405,1205,1229]" pageId="1" pageNumber="298" pagination="7 - 288" refId="ref1829" refString="Souza, V. C. &amp; Giulietti, A. M. (2009) Levantamento das especies de Scrophulariaceae sensu lato nativas do Brasil. Pesquisas Botanica 60: 7 - 288." type="journal article" year="2009">Souza &amp; Giulietti (2009)</bibRefCitation>
superfluous. On the other hand, we found
<specimenCount id="9D7DFD77706CFFCDF74144A6FB94FB2F" box="[872,1035,1205,1229]" pageId="1" pageNumber="298" type="generic">four specimens</specimenCount>
of
<emphasis id="B90FEAEC706CFFCDF01844A6FB5EFB2F" box="[1073,1217,1205,1229]" italics="true" pageId="1" pageNumber="298">Hassler 7828</emphasis>
at G and believe that the lectotypification should be narrowed according to Art. 9.17. Hence, we selected the specimen G barcode G00366028 as second-step
<typeStatus id="54C0885C706CFFCDF50744EEFE0FFAF7" box="[302,400,1277,1301]" pageId="1" pageNumber="298" type="lectotype">lectotype</typeStatus>
because it bears the label of “
<typeStatus id="54C0885C706CFFCDF6C944EEFCD7FAF7" box="[736,840,1277,1301]" pageId="1" pageNumber="298" type="holotype">Holotype</typeStatus>
” signed by Barringer, indicating that this was the exact specimen he was referring to in
<bibRefCitation id="EFEA4B0F706CFFCDF5F74532FD0AFADB" author="Barringer, K." box="[478,661,1313,1337]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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