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<document id="664039DA604A7AEB69F45AB6C8897765" ID-DOI="10.11646/phytotaxa.501.2.5" ID-ISSN="1179-3163" ID-Zenodo-Dep="5424848" IM.bibliography_approvedBy="guilherme" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1630392910242" checkinUser="marcus" docAuthor="Scatigna, André Vito &amp; Souza, Vinicius Castro" docDate="2021" docId="03D287E8706CFFCDF4944556FDB2F943" docLanguage="en" docName="phytotaxa.501.2.5.pdf" docOrigin="Phytotaxa 501 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.501.2.5" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Angelonia micrantha , Bentham 1846" docType="treatment" docVersion="1" lastPageNumber="298" masterDocId="FFEBFF90706DFFCCF4294013FF9FFFE2" masterDocTitle="Tying up loose ends: The synonymization of Monopera with Angelonia (Plantaginaceae), including a new combination and typifications" masterLastPageNumber="299" masterPageNumber="297" pageNumber="298" updateTime="1732203349186" updateUser="guilherme" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C06DEBE26D5E02B2C7EBA3E07A7906A2">Tying up loose ends: The synonymization of Monopera with Angelonia (Plantaginaceae), including a new combination and typifications</mods:title>
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<mods:namePart id="2DFB775C6D7F186236680B4C78C213CA">Scatigna, André Vito</mods:namePart>
<mods:affiliation id="8A14327385D2B00FBAFD5554C201ED4E">Centro de Educação, Ciências Exatas e Naturais, Departamento de Biologia, Universidade Estadual do Maranhão, Cidade Universitária Paulo VI, s / n, 65054 - 970, São Luís, Maranhão, Brazil.</mods:affiliation>
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<mods:namePart id="AC6510F6B864CA9CCDEFB18F50FEBBC5">Souza, Vinicius Castro</mods:namePart>
<mods:affiliation id="F30AC7C4343EAB825A221FEF6EC08856">Herbário ESA, Departamento de Ciências Biológicas, Escola Superior de Agricultura Luiz de Queiroz, Universidade de São Paulo, Av. Pádua Dias, 11, Agronomia, 13418 - 400, Piracicaba, São Paulo, Brazil.</mods:affiliation>
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<mods:title id="29C12759F1BA4EC00E8F14CDA2C54068">Phytotaxa</mods:title>
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<mods:date id="209AC7DE0C96F5D7E182D08918112BC4">2021</mods:date>
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<mods:number id="0DE1B5EC52308A6712EAD17B08205032">2021-05-20</mods:number>
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<taxonomicName id="4C7B4D7D706CFFCDF4944556FD3FFABF" ID-CoL="8R6ZX" authority="Bentham (1846: 252)" authorityName=", Bentham" authorityPageNumber="252" authorityYear="1846" box="[189,672,1349,1373]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="micrantha">
<emphasis id="B90FEAEC706CFFCDF4944556FE34FABF" bold="true" box="[189,427,1349,1373]" italics="true" pageId="1" pageNumber="298">Angelonia micrantha</emphasis>
<bibRefCitation id="EFEA4B0F706CFFCDF59D4556FD3FFABF" author="Bentham, G." box="[436,672,1349,1373]" pageId="1" pageNumber="298" pagination="186 - 586" refId="ref1616" refString="Bentham, G. (1846) Scrophulariaceae. In: de Candolle, A. (1846) Prodromus systematis naturalis regni vegetabilis 10: 186 - 586." type="journal article" year="1846">Bentham (1846: 252)</bibRefCitation>
</taxonomicName>
.
</paragraph>
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<taxonomicName id="4C7B4D7D706CFFCDF6844555FB71FABF" authority="(Benth.) Barringer (1983: 112)" authorityName="Barringer" authorityPageNumber="112" authorityYear="1983" baseAuthorityName="Benth." box="[685,1262,1349,1373]" class="Magnoliopsida" family="Plantaginaceae" genus="Monopera" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="micrantha">
<emphasis id="B90FEAEC706CFFCDF6844555FC0CFABF" box="[685,915,1349,1373]" italics="true" pageId="1" pageNumber="298">Monopera micrantha</emphasis>
(Benth.)
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</taxonomicName>
.
</paragraph>
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<typeStatus id="54C0885C706CFFCDF0D24555FAF9FABC" box="[1275,1382,1350,1374]" pageId="1" pageNumber="298" type="lectotype">Lectotype</typeStatus>
(firststep designated by
<bibRefCitation id="EFEA4B0F706CFFCDF57E457AFE62FA63" author="Barringer, K." box="[343,509,1385,1409]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer 1983</bibRefCitation>
, second-step here designated):—
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. “Prov. Piauhy [
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]”, 1839,
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<collectorName id="268E5328706CFFCDF13A457AFAF1FA63" box="[1299,1390,1385,1409]" pageId="1" pageNumber="298">Gardner</collectorName>
2274
</emphasis>
(
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barcode
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</materialsCitation>
;
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<typeStatus id="54C0885C706CFFCDF580459EFDA6FA47" box="[425,569,1421,1445]" pageId="1" pageNumber="298">isolectotypes</typeStatus>
:
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barcode
<specimenCode id="DBDD9E85706CFFCDF6FE459EFCE3FA47" box="[727,892,1421,1445]" pageId="1" pageNumber="298">BM000953335</specimenCode>
,
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<collectionCode id="ED6AAE3B706CFFCDF7A3459EFC2EFA47" box="[906,945,1421,1445]" pageId="1" pageNumber="298">GH</collectionCode>
barcode
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,
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<collectionCode id="ED6AAE3B706CFFCDF0BD459DFB3CFA47" box="[1172,1187,1422,1445]" pageId="1" pageNumber="298">F</collectionCode>
barcode
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,
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barcodes
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,
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<materialsCitation id="3B133CA3706CFFCDF55F45A2FE6CFA2B" box="[374,499,1457,1481]" collectionCode="G" collectorName="Gardner" country="Brazil" pageId="1" pageNumber="298" specimenCode="G00366027" specimenCount="1" stateProvince="Piaui" typeStatus="isolectotype">
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and
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,
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<collectionCode id="ED6AAE3B706CFFCDF68745A1FD5EFA2B" box="[686,705,1458,1481]" pageId="1" pageNumber="298">K</collectionCode>
barcode
<specimenCode id="DBDD9E85706CFFCDF70945A2FC31FA2B" box="[800,942,1457,1481]" pageId="1" pageNumber="298">K000528881</specimenCode>
,
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<materialsCitation id="3B133CA3706CFFCDF79E45A2FB32FA2B" box="[951,1197,1457,1481]" collectionCode="OXF" collectorName="Gardner" country="Brazil" pageId="1" pageNumber="298" specimenCount="1" stateProvince="Piaui" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF79E45A2FC73FA2B" box="[951,1004,1457,1481]" collectionName="OXF" pageId="1" pageNumber="298">OXF</collectionCode>
[personally seen],
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<collectionCode id="ED6AAE3B706CFFCDF09B45A1FB47FA2B" box="[1202,1240,1458,1481]" pageId="1" pageNumber="298">NY</collectionCode>
barcodes
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and
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<specimenCode id="DBDD9E85706CFFCDF49C45C6FEBDFA0F" box="[181,290,1493,1517]" pageId="1" pageNumber="298">00067835</specimenCode>
,
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<materialsCitation id="3B133CA3706CFFCDF50445C5FDBEFA0F" box="[301,545,1493,1517]" collectionCode="P" collectorName="Gardner" country="Brazil" pageId="1" pageNumber="298" specimenCode="P00584257" specimenCount="1" stateProvince="Piaui" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF50445C5FEA4FA0F" box="[301,315,1494,1517]" pageId="1" pageNumber="298">P</collectionCode>
barcodes
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and
<materialsCitation id="3B133CA3706CFFCDF67C45C6FD4AFA0F" box="[597,725,1493,1517]" collectionCode="P" collectorName="Gardner" country="Brazil" pageId="1" pageNumber="298" specimenCode="P00584256" specimenCount="1" stateProvince="Piaui" typeStatus="isolectotype">
<specimenCode id="DBDD9E85706CFFCDF67C45C6FD4EFA0F" box="[597,721,1493,1517]" pageId="1" pageNumber="298">P00584256</specimenCode>
,
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<materialsCitation id="3B133CA3706CFFCDF6F245C5FC70FA0F" box="[731,1007,1493,1517]" collectionCode="W" collectorName="Gardner" country="Brazil" pageId="1" pageNumber="298" specimenCode="W 0013988" specimenCount="1" stateProvince="Piaui" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF6F245C5FD6BFA0F" box="[731,756,1494,1517]" pageId="1" pageNumber="298">W</collectionCode>
accession
<specimenCode id="DBDD9E85706CFFCDF74E45C5FC77FA0F" box="[871,1000,1493,1517]" pageId="1" pageNumber="298">W 0013988</specimenCode>
)
</materialsCitation>
.
<figureCitation id="13402A7B706CFFCDF7D045C6FBD4FA0F" box="[1017,1099,1493,1517]" captionStart="FIGURE 1" captionStartId="1.[136,229,848,870]" captionTargetBox="[151,1435,244,824]" captionTargetId="figure-38@1.[151,1435,244,824]" captionTargetPageId="1" captionText="FIGURE 1. A. Flower of Angelonia perennis in front view showing a bisaccate corolla (white arrow). B. Flower of A. micrantha in lateral view. Photos: A by André V. Scatigna, B by Ianne C.S. Nobre." figureDoi="http://doi.org/10.5281/zenodo.5424850" httpUri="https://zenodo.org/record/5424850/files/figure.png" pageId="1" pageNumber="298">Fig. 1B</figureCitation>
.
</paragraph>
</subSubSection>
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<paragraph id="8BC436FE706CFFCDF49445E9FDB2F943" blockId="1.[136,1452,953,1697]" pageId="1" pageNumber="298">
In the protologue of
<taxonomicName id="4C7B4D7D706CFFCDF58845EAFCD9F9F3" authority=", Bentham (1846)" authorityName=", Bentham" authorityYear="1846" box="[417,838,1529,1553]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="micrantha">
<emphasis id="B90FEAEC706CFFCDF58845EAFD19F9F3" box="[417,646,1529,1553]" italics="true" pageId="1" pageNumber="298">Angelonia micrantha</emphasis>
,
<bibRefCitation id="EFEA4B0F706CFFCDF6BC45EAFCD9F9F3" author="Bentham, G." box="[661,838,1529,1553]" pageId="1" pageNumber="298" pagination="186 - 586" refId="ref1616" refString="Bentham, G. (1846) Scrophulariaceae. In: de Candolle, A. (1846) Prodromus systematis naturalis regni vegetabilis 10: 186 - 586." type="journal article" year="1846">Bentham (1846)</bibRefCitation>
</taxonomicName>
cited
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,
<emphasis id="B90FEAEC706CFFCDF01345EAFB4CF9F3" box="[1082,1235,1529,1553]" italics="true" pageId="1" pageNumber="298">Gardner 2273</emphasis>
and
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, without designating any herbarium. Later,
<bibRefCitation id="EFEA4B0F706CFFCDF66D460EFD69F9D7" author="Barringer, K." box="[580,758,1565,1589]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
selected a specimen of
<emphasis id="B90FEAEC706CFFCDF7C3460EFBE1F9D7" box="[1002,1150,1565,1589]" italics="true" pageId="1" pageNumber="298">Gardner 2274</emphasis>
housed at K as the
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. However, there are
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<emphasis id="B90FEAEC706DFFCCF71346D7FBA5F93E" box="[826,1082,1732,1756]" italics="true" pageId="0" pageNumber="297">Angelonia salicariifolia</emphasis>
Bonpl.
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=
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(Benth.) Barringer
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(≡
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<emphasis id="B90FEAEC706DFFCCF1AC471DFF7AF8A5" italics="true" pageId="0" pageNumber="297">syn. nov.</emphasis>
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<taxonomicName id="4C7B4D7D706DFFCCF4A14766FC31F86C" ID-CoL="8RTL9" authority="(Chodat &amp; Hassler 1904: 285) Scatigna &amp; V. C. Souza" authorityName="Scatigna &amp; V. C. Souza" authorityYear="2021" baseAuthorityName="Chodat &amp; Hassler" baseAuthorityPageNumber="285" baseAuthorityYear="1904" box="[136,942,1909,1934]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="perennis" status="comb. nov.">
<emphasis id="B90FEAEC706DFFCCF4A14766FEFEF86F" bold="true" box="[136,353,1909,1933]" italics="true" pageId="0" pageNumber="297">Angelonia perennis</emphasis>
(
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,
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<emphasis id="B90FEAEC706DFFCCF7934766FBB1F86F" box="[954,1070,1909,1933]" italics="true" pageId="0" pageNumber="297">comb. nov.</emphasis>
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<taxonomicName id="4C7B4D7D706DFFCCF01F4766FE57F852" authority="Chodat &amp; Hassler (1904: 285)" authorityName=", Chodat &amp; Hassler" authorityPageNumber="285" authorityYear="1904" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="variety" species="micrantha" variety="perennis">
<emphasis id="B90FEAEC706DFFCCF01F4766FA86F86F" box="[1078,1305,1909,1933]" italics="true" pageId="0" pageNumber="297">Angelonia micrantha</emphasis>
var.
<emphasis id="B90FEAEC706DFFCCF1794765FA34F86F" box="[1360,1451,1910,1933]" italics="true" pageId="0" pageNumber="297">perennis</emphasis>
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.
</paragraph>
<paragraph id="8BC436FE706DFFCCF5F8478AFBC7F853" blockId="0.[136,1452,1909,2074]" box="[465,1112,1945,1969]" pageId="0" pageNumber="297">
<taxonomicName id="4C7B4D7D706DFFCCF5F8478AFBCCF853" authority="(Chodat &amp; Hassl.) Barringer (1983: 113)" authorityName="Barringer" authorityPageNumber="113" authorityYear="1983" baseAuthorityName="Chodat &amp; Hassl." box="[465,1107,1945,1969]" class="Magnoliopsida" family="Plantaginaceae" genus="Monopera" kingdom="Plantae" order="Lamiales" pageId="0" pageNumber="297" phylum="Tracheophyta" rank="species" species="perennis">
<emphasis id="B90FEAEC706DFFCCF5F8478AFD3FF852" box="[465,672,1945,1968]" italics="true" pageId="0" pageNumber="297">Monopera perennis</emphasis>
(Chodat &amp; Hassl.)
<bibRefCitation id="EFEA4B0F706DFFCCF742478AFBCCF853" author="Barringer, K." box="[875,1107,1945,1969]" pageId="0" pageNumber="297" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983: 113)</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3616575706DFFCDF074478AFEDCFF38" lastPageId="1" lastPageNumber="298" pageId="0" pageNumber="297" type="materials_examined">
<paragraph id="8BC436FE706DFFCDF074478AFEDCFF38" blockId="0.[136,1452,1909,2074]" lastBlockId="1.[136,1451,159,218]" lastPageId="1" lastPageNumber="298" pageId="0" pageNumber="297">
<materialsCitation id="3B133CA3706DFFCCF074478AFDB0F815" collectionCode="G" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="G00366028" specimenCount="1" typeStatus="lectotype">
<typeStatus id="54C0885C706DFFCCF074478AFB57F853" box="[1117,1224,1945,1969]" pageId="0" pageNumber="297" type="lectotype">Lectotype</typeStatus>
(first-step designated by
<bibRefCitation id="EFEA4B0F706DFFCCF48247AFFECDF836" author="Barringer, K." box="[171,338,1980,2004]" pageId="0" pageNumber="297" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer 1983</bibRefCitation>
, second-step here designated):—
<collectingCountry id="F36C766E706DFFCCF69B47AFFCD7F836" box="[690,840,1980,2004]" name="Paraguay" pageId="0" pageNumber="297">PARAGUAY</collectingCountry>
. “
<location id="8EA46025706DFFCCF77747AFFABBF836" LSID="urn:lsid:plazi:treatment:03D287E8706DFFCDF4A14766FD06FADB:8EA46025706DFFCCF77747AFFABBF836" box="[862,1316,1980,2004]" country="Paraguay" name="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297">in regione cursus superioris fluminis Apa</location>
”, November,
<emphasis id="B90FEAEC706DFFCCF4BF47CCFEB9F815" box="[150,294,2015,2039]" italics="true" pageId="0" pageNumber="297">
<collectorName id="268E5328706DFFCCF4BF47CCFF76F815" box="[150,233,2015,2039]" pageId="0" pageNumber="297">Hassler</collectorName>
7828
</emphasis>
(
<collectionCode id="ED6AAE3B706DFFCCF51F47CCFED5F815" box="[310,330,2015,2039]" pageId="0" pageNumber="297">G</collectionCode>
barcode
<specimenCode id="DBDD9E85706DFFCCF58747CCFDB0F815" box="[430,559,2015,2039]" pageId="0" pageNumber="297">G00366028</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B133CA3706DFFCCF61247CCFC48F815" box="[571,983,2015,2039]" collectionCode="G" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="G00366023" specimenCount="1" typeStatus="isolectotype">
<typeStatus id="54C0885C706DFFCCF61247CCFD54F815" box="[571,715,2015,2039]" pageId="0" pageNumber="297">isolectotypes</typeStatus>
:
<collectionCode id="ED6AAE3B706DFFCCF6FC47CCFD77F815" box="[725,744,2015,2039]" pageId="0" pageNumber="297">G</collectionCode>
barcodes
<specimenCode id="DBDD9E85706DFFCCF77F47CCFC48F815" box="[854,983,2015,2039]" pageId="0" pageNumber="297">G00366023</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706DFFCCF7CB47CCFBFCF815" box="[994,1123,2015,2039]" collectionCode="G" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="G00366029" specimenCount="1" typeStatus="isolectotype">
<specimenCode id="DBDD9E85706DFFCCF7CB47CCFBFCF815" box="[994,1123,2015,2039]" pageId="0" pageNumber="297">G00366029</specimenCode>
</materialsCitation>
, and
<materialsCitation id="3B133CA3706DFFCCF0B447CCFA81F815" box="[1181,1310,2015,2039]" collectionCode="G" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="G00366033" specimenCount="1" typeStatus="isolectotype">
<specimenCode id="DBDD9E85706DFFCCF0B447CCFA81F815" box="[1181,1310,2015,2039]" pageId="0" pageNumber="297">G00366033</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706DFFCCF10047CCFF6AF7F8" collectionCode="GH" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="00077891" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706DFFCCF10047CCFACFF815" box="[1321,1360,2015,2039]" pageId="0" pageNumber="297">GH</collectionCode>
barcode
<specimenCode id="DBDD9E85706DFFCCF4A14811FF6AF7F8" box="[136,245,2050,2074]" pageId="0" pageNumber="297">00077891</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706DFFCCF52D4810FD90F7F8" box="[260,527,2050,2074]" collectionCode="K" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="K000528880" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706DFFCCF52D4810FE88F7F8" box="[260,279,2051,2074]" pageId="0" pageNumber="297">K</collectionCode>
barcode
<specimenCode id="DBDD9E85706DFFCCF5A84811FD90F7F8" box="[385,527,2050,2074]" pageId="0" pageNumber="297">K000528880</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706DFFCCF6344811FCD9F7F8" box="[541,838,2050,2074]" collectionCode="MO" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="MO-2267514" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706DFFCCF6344811FDD7F7F8" box="[541,584,2050,2074]" pageId="0" pageNumber="297">MO</collectionCode>
barcode
<specimenCode id="DBDD9E85706DFFCCF6984811FCD9F7F8" box="[689,838,2050,2074]" pageId="0" pageNumber="297">MO-2267514</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706DFFCCF77D4810FB1CF7F8" box="[852,1155,2050,2074]" collectionCode="MPU" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="MPU018189" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706DFFCCF77D4810FC11F7F8" box="[852,910,2051,2074]" collectionName="MPU" pageId="0" pageNumber="297">MPU</collectionCode>
barcode
<specimenCode id="DBDD9E85706DFFCCF7DE4811FB1CF7F8" box="[1015,1155,2050,2074]" pageId="0" pageNumber="297">MPU018189</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706DFFCCF0BB4810FA10F7F8" box="[1170,1423,2050,2074]" collectionCode="NY" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="00067836" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706DFFCCF0BB4810FB27F7F8" box="[1170,1208,2051,2074]" pageId="0" pageNumber="297">NY</collectionCode>
barcode
<specimenCode id="DBDD9E85706DFFCCF1084811FA10F7F8" box="[1313,1423,2050,2074]" pageId="0" pageNumber="297">00067836</specimenCode>
</materialsCitation>
;
<materialsCitation id="3B133CA3706DFFCDF1B44810FEF9FF55" collectionCode="P" collectorName="Hassler" country="Paraguay" lastPageId="1" lastPageNumber="298" location="in regione cursus superioris fluminis Apa" pageId="0" pageNumber="297" specimenCode="P00584259" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706DFFCCF1B44810FA34F7F8" box="[1437,1451,2051,2074]" pageId="0" pageNumber="297">P</collectionCode>
barcodes
<specimenCode id="DBDD9E85706CFFCDF4C5408CFEF9FF55" box="[236,358,159,183]" pageId="1" pageNumber="298">P00584259</specimenCode>
</materialsCitation>
and
<materialsCitation id="3B133CA3706CFFCDF5BE408CFD8BFF55" box="[407,532,159,183]" collectionCode="P" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="1" pageNumber="298" specimenCode="P00584258" specimenCount="1" typeStatus="isolectotype">
<specimenCode id="DBDD9E85706CFFCDF5BE408CFD8BFF55" box="[407,532,159,183]" pageId="1" pageNumber="298">P00584258</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706CFFCDF63440B3FC95FF55" box="[541,778,159,183]" collectionCode="PH" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="1" pageNumber="298" specimenCode="00003211" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF63440B3FDA0FF55" box="[541,575,160,183]" pageId="1" pageNumber="298">PH</collectionCode>
barcode
<specimenCode id="DBDD9E85706CFFCDF6B7408CFC95FF55" box="[670,778,159,183]" pageId="1" pageNumber="298">00003211</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706CFFCDF73D408CFB9AFF55" box="[788,1029,159,183]" collectionCode="S" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="1" pageNumber="298" specimenCode="S10-19369" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF73D408CFCBCFF55" box="[788,803,159,183]" pageId="1" pageNumber="298">S</collectionCode>
Herb. N°
<specimenCode id="DBDD9E85706CFFCDF7A4408CFB9AFF55" box="[909,1029,159,183]" pageId="1" pageNumber="298">S10-19369</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706CFFCDF027408CFA95FF55" box="[1038,1290,159,183]" collectionCode="UC" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="1" pageNumber="298" specimenCode="UC944846" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF027408CFBACFF55" box="[1038,1075,159,183]" pageId="1" pageNumber="298">UC</collectionCode>
barcode
<specimenCode id="DBDD9E85706CFFCDF0BA408CFA95FF55" box="[1171,1290,159,183]" pageId="1" pageNumber="298">UC944846</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B133CA3706CFFCDF13A40B3FF7EFF38" collectionCode="W" collectorName="Hassler" country="Paraguay" location="in regione cursus superioris fluminis Apa" pageId="1" pageNumber="298" specimenCode="622" specimenCount="1" typeStatus="isolectotype">
<collectionCode id="ED6AAE3B706CFFCDF13A40B3FAB3FF55" box="[1299,1324,160,183]" pageId="1" pageNumber="298">W</collectionCode>
Acqu. 1904 N°
<specimenCode id="DBDD9E85706CFFCDF48440D1FF46FF38" box="[173,217,194,218]" pageId="1" pageNumber="298">622</specimenCode>
)
</materialsCitation>
.
<figureCitation id="13402A7B706CFFCDF4C240D1FEA0FF38" box="[235,319,194,218]" captionStart="FIGURE 1" captionStartId="1.[136,229,848,870]" captionTargetBox="[151,1435,244,824]" captionTargetId="figure-38@1.[151,1435,244,824]" captionTargetPageId="1" captionText="FIGURE 1. A. Flower of Angelonia perennis in front view showing a bisaccate corolla (white arrow). B. Flower of A. micrantha in lateral view. Photos: A by André V. Scatigna, B by Ianne C.S. Nobre." figureDoi="http://doi.org/10.5281/zenodo.5424850" httpUri="https://zenodo.org/record/5424850/files/figure.png" pageId="1" pageNumber="298">Fig. 1A</figureCitation>
.
</paragraph>
</subSubSection>
<caption id="DF046676706CFFCDF4A14343FD42FC68" ID-DOI="http://doi.org/10.5281/zenodo.5424850" ID-Zenodo-Dep="5424850" httpUri="https://zenodo.org/record/5424850/files/figure.png" pageId="1" pageNumber="298" startId="1.[136,229,848,870]" targetBox="[151,1435,244,824]" targetPageId="1">
<paragraph id="8BC436FE706CFFCDF4A14343FD42FC68" blockId="1.[136,1451,848,906]" pageId="1" pageNumber="298">
<emphasis id="B90FEAEC706CFFCDF4A14343FF62FC87" bold="true" box="[136,253,848,870]" pageId="1" pageNumber="298">FIGURE 1.</emphasis>
A. Flower of
<taxonomicName id="4C7B4D7D706CFFCDF5A84343FDA4FC84" authorityName="Scatigna &amp; V. C. Souza" authorityYear="2021" baseAuthorityName="Chodat &amp; Hassler" baseAuthorityYear="1904" box="[385,571,848,870]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="perennis">
<emphasis id="B90FEAEC706CFFCDF5A84343FDA4FC84" box="[385,571,848,870]" italics="true" pageId="1" pageNumber="298">Angelonia perennis</emphasis>
</taxonomicName>
in front view showing a bisaccate corolla (white arrow). B. Flower of
<taxonomicName id="4C7B4D7D706CFFCDF0FF4343FACCFC87" authorityName=", Bentham" authorityYear="1846" box="[1238,1363,848,869]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="species" species="micrantha">
<emphasis id="B90FEAEC706CFFCDF0FF4343FACCFC87" box="[1238,1363,848,869]" italics="true" pageId="1" pageNumber="298">A. micrantha</emphasis>
</taxonomicName>
in lateral view. Photos: A by André V. Scatigna, B by Ianne C.S. Nobre.
</paragraph>
</caption>
<subSubSection id="C3616575706CFFCDF49443A9FD06FADB" pageId="1" pageNumber="298" type="description">
<paragraph id="8BC436FE706CFFCDF49443A9FD06FADB" blockId="1.[136,1452,953,1697]" pageId="1" pageNumber="298">
In the protologue of
<taxonomicName id="4C7B4D7D706CFFCDF58B43AAFBA1FC33" authority=", Chodat &amp; Hassler (1904)" authorityName=", Chodat &amp; Hassler" authorityYear="1904" box="[418,1086,953,977]" class="Magnoliopsida" family="Plantaginaceae" genus="Angelonia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="298" phylum="Tracheophyta" rank="variety" species="micrantha" variety="perennis">
<emphasis id="B90FEAEC706CFFCDF58B43AAFD19FC33" box="[418,646,953,977]" italics="true" pageId="1" pageNumber="298">Angelonia micrantha</emphasis>
var.
<emphasis id="B90FEAEC706CFFCDF69643A9FC85FC33" box="[703,794,954,977]" italics="true" pageId="1" pageNumber="298">perennis</emphasis>
,
<bibRefCitation id="EFEA4B0F706CFFCDF70343AAFBA1FC33" author="Chodat, R. &amp; Hassler, E." box="[810,1086,953,977]" pageId="1" pageNumber="298" pagination="475 - 489" refId="ref1646" refString="Chodat, R. &amp; Hassler, E. (1904) Plantae hasslerianae. Bulletin de l'Herbier Boissier 2: 475 - 489." type="journal article" year="1904">Chodat &amp; Hassler (1904)</bibRefCitation>
</taxonomicName>
cited the specimen
<emphasis id="B90FEAEC706CFFCDF13343AAFA34FC33" box="[1306,1451,953,977]" italics="true" pageId="1" pageNumber="298">Hassler 7828</emphasis>
without assigning it to any herbarium. Later,
<bibRefCitation id="EFEA4B0F706CFFCDF64343CEFC80FC17" author="Barringer, K." box="[618,799,989,1013]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
cited “
<typeStatus id="54C0885C706CFFCDF74343CEFC4CFC17" box="[874,979,989,1013]" pageId="1" pageNumber="298" type="holotype">Holotype</typeStatus>
: G!; Isotypes: K!, MO!, NY!, P!, UC!, W!”. More recently,
<bibRefCitation id="EFEA4B0F706CFFCDF5064412FDDEFBFB" author="Souza, V. C. &amp; Giulietti, A. M." box="[303,577,1025,1049]" pageId="1" pageNumber="298" pagination="7 - 288" refId="ref1829" refString="Souza, V. C. &amp; Giulietti, A. M. (2009) Levantamento das especies de Scrophulariaceae sensu lato nativas do Brasil. Pesquisas Botanica 60: 7 - 288." type="journal article" year="2009">Souza &amp; Giulietti (2009)</bibRefCitation>
designated the specimen at G as
<typeStatus id="54C0885C706CFFCDF79E4412FB83FBFB" box="[951,1052,1025,1049]" pageId="1" pageNumber="298" type="lectotype">lectotype</typeStatus>
. However, because no
<typeStatus id="54C0885C706CFFCDF1324412FAE7FBFB" box="[1307,1400,1025,1049]" pageId="1" pageNumber="298" type="holotype">holotype</typeStatus>
was designated in the protologue and considering that the authors had access to all extant specimens of
<emphasis id="B90FEAEC706CFFCDF0EB4436FACDFBDF" box="[1218,1362,1061,1085]" italics="true" pageId="1" pageNumber="298">Hassler 7828</emphasis>
prior to publication, since Hassler is the gathering collector, the specimens in different herbaria are
<typeStatus id="54C0885C706CFFCDF0124459FB04FB80" box="[1083,1179,1098,1122]" pageId="1" pageNumber="298" type="syntype">syntypes</typeStatus>
, and the use of “
<typeStatus id="54C0885C706CFFCDF169445AFA3DFB83" box="[1344,1442,1097,1121]" pageId="1" pageNumber="298" type="holotype">holotype</typeStatus>
” by
<bibRefCitation id="EFEA4B0F706CFFCDF480447EFEC1FB67" author="Barringer, K." box="[169,350,1133,1157]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
is an error to be corrected to “
<typeStatus id="54C0885C706CFFCDF68B447EFC97FB67" box="[674,776,1133,1157]" pageId="1" pageNumber="298" type="lectotype">lectotype</typeStatus>
” according to Art. 9.10. Moreover, this inadvertent designation is effective, fulfilling Art. 7.11, and must be followed according to Art. 10.5, rendering the subsequent lectotypification by
<bibRefCitation id="EFEA4B0F706CFFCDF4A144A6FE0AFB2F" author="Souza, V. C. &amp; Giulietti, A. M." box="[136,405,1205,1229]" pageId="1" pageNumber="298" pagination="7 - 288" refId="ref1829" refString="Souza, V. C. &amp; Giulietti, A. M. (2009) Levantamento das especies de Scrophulariaceae sensu lato nativas do Brasil. Pesquisas Botanica 60: 7 - 288." type="journal article" year="2009">Souza &amp; Giulietti (2009)</bibRefCitation>
superfluous. On the other hand, we found
<specimenCount id="9D7DFD77706CFFCDF74144A6FB94FB2F" box="[872,1035,1205,1229]" pageId="1" pageNumber="298" type="generic">four specimens</specimenCount>
of
<emphasis id="B90FEAEC706CFFCDF01844A6FB5EFB2F" box="[1073,1217,1205,1229]" italics="true" pageId="1" pageNumber="298">Hassler 7828</emphasis>
at G and believe that the lectotypification should be narrowed according to Art. 9.17. Hence, we selected the specimen G barcode G00366028 as second-step
<typeStatus id="54C0885C706CFFCDF50744EEFE0FFAF7" box="[302,400,1277,1301]" pageId="1" pageNumber="298" type="lectotype">lectotype</typeStatus>
because it bears the label of “
<typeStatus id="54C0885C706CFFCDF6C944EEFCD7FAF7" box="[736,840,1277,1301]" pageId="1" pageNumber="298" type="holotype">Holotype</typeStatus>
” signed by Barringer, indicating that this was the exact specimen he was referring to in
<bibRefCitation id="EFEA4B0F706CFFCDF5F74532FD0AFADB" author="Barringer, K." box="[478,661,1313,1337]" pageId="1" pageNumber="298" pagination="111 - 114" refId="ref1581" refString="Barringer, K. (1983) Monopera, a new genus of Scrophulariaceae from South America. Brittonia 35: 111 - 114. https: // doi. org / 10.2307 / 2805946" type="journal article" year="1983">Barringer (1983)</bibRefCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>