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<document id="3B31F7A64AF4C4BBA33B282A8877BB4B" ID-DOI="10.3897/zookeys.1208.116083" ID-ZooBank="0215503D-E8B4-4179-89F6-FB2CCC59F37E" ID-publisher-id="116083" URI-arpha="CF5F342A-04A4-57D6-860A-D48A80ACC24A" URI-zoobank="http://zoobank.org/0215503D-E8B4-4179-89F6-FB2CCC59F37E" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.taxonomicNames_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1722361871078" checkinUser="pensoft" docAuthor="Man, Nem Sian, Ablett, Jonathan D., Lwin, Ngwe, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2024" docId="E2CE7E50CA86543985F056C195CB4AB9" docLanguage="en" docName="ZooKeys 1208: 173-239" docOrigin="ZooKeys 1208" docSource="https://zookeys.pensoft.net/article/116083/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Glessula woodthorpei Godwin-Austen 1920" docType="treatment" docVersion="3" dtd-version="3.0" lastPageNumber="239" masterDocId="CF5F342A04A457D6860AD48A80ACC24A" masterDocTitle="Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar" masterLastPageNumber="239" masterPageNumber="173" pageNumber="173" updateTime="1722370176574" updateUser="admin">
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<mods:title id="1A532200BB33FE4D43B82639F5FE1749">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:name id="3A5BB7F75AF4423DAEF15E90D950A232" type="personal">
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<mods:namePart id="11EB382078E5C12E1B0E352DD914D1A6">Man, Nem Sian</mods:namePart>
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<mods:affiliation id="ADAEC331E5D7724166E31894B4BB5862">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="B76FC080A7C7AD1FC09F8E29E5DD93AC">Ablett, Jonathan D.</mods:namePart>
<mods:nameIdentifier id="E0F439F7BC761FA0E57F32099685DA65" type="ORCID">0000-0002-7277-1934</mods:nameIdentifier>
<mods:affiliation id="8578ADD302E69BA0F9CF66F27CA169E8">Department of Life Sciences, The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
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<mods:namePart id="9BFCBFDA643B4C93EA6A4DACBDD27683">Lwin, Ngwe</mods:namePart>
<mods:nameIdentifier id="036969F69B54E8A6E45474FF060BE1D9" type="ORCID">0000-0002-4394-3741</mods:nameIdentifier>
<mods:affiliation id="6E86DB7D6594214D61D553D546E50F5C">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
</mods:name>
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<mods:namePart id="7C1CF8C9768EBF5A76B5F8042B8E00B3">Sutcharit, Chirasak</mods:namePart>
<mods:nameIdentifier id="6EE98595DB4EFD4190F566BD2E22E1A3" type="ORCID">0000-0001-7670-9540</mods:nameIdentifier>
<mods:affiliation id="E3E6266C995B19500E0BB8697B82CD2A">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand</mods:affiliation>
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<mods:namePart id="8C8F2D3D8692E77B680C4100CF978BF5">Panha, Somsak</mods:namePart>
<mods:nameIdentifier id="EC18E632BEA15D1F2E8F4BF0B7D24532" type="ORCID">0000-0002-4431-2458</mods:nameIdentifier>
<mods:affiliation id="57DC7018454E83AD137D9D3845BF682C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:typeOfResource id="99ED2D4F7183E2FFAF7B8E7D6EA5C1C1">text</mods:typeOfResource>
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<mods:date id="450557178E32D6A66FCA836C0FBF99EE">2024</mods:date>
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<mods:number id="C90C012B2A50A31BE2D432B6E867F138">1208</mods:number>
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<treatment id="E2CE7E50CA86543985F056C195CB4AB9" ID-DOI="http://doi.org/10.5281/zenodo.13137126" ID-Zenodo-Dep="13137126" ID-arpha="E2CE7E50-CA86-5439-85F0-56C195CB4AB9" LSID="urn:lsid:plazi:treatment:E2CE7E50CA86543985F056C195CB4AB9" httpUri="http://treatment.plazi.org/id/E2CE7E50CA86543985F056C195CB4AB9">
<subSubSection id="CE2BAC4091A2A1DE23DC9C2A74824C31" type="nomenclature">
<paragraph id="0018F872C86624737847C4A2704CE6CC">
<heading id="A1BFED996CEEE959F37650C4E4FB28BB" reason="label">19</heading>
<taxonomicName id="8B812B951A52E76EC8B6EC7E16B87802" authority="Godwin-Austen, 1920" authorityName="Godwin-Austen" authorityYear="1920" class="Gastropoda" family="Subulinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="woodthorpei">
<emphasis id="CAC726CF47970EE0CEB8A4D8F6CD84A7" italics="true">Glessula woodthorpei</emphasis>
<bibRefCitation id="54E7F26E69ED6619F2D8F94E85225D51" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="8218BF9E9E97133EA6848C1F0AAD369E" type="description">
<paragraph id="9A9C055C195599D4501A811FFA53BF2E">
<figureCitation id="7CD92F3A1A6A902F5498136E06C8DD4A" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">Fig. 11 C E</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="6981EAABE71905DD0A64C8A9B33137DB" type="reference_group">
<paragraph id="74724A4EB06F960AD416EB6CB903B575">
<treatmentCitationGroup id="0FBFB6773CBC91C8544517149CD6FDAE">
<treatmentCitation id="5FB2409568F9B12B7D278A9374710A97" author="Godwin-Austen HH" page="58" year="1920">
<taxonomicName id="7C01383A3233860D9D41A8428D50BC3E" ID-CoL="7RFLT" authority="Godwin-Austen, 1920: 58" authorityName="Godwin-Austen" authorityPageNumber="58" authorityYear="1920" class="Gastropoda" family="Achatinidae" genus="Glessula" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="woodthorpei">
<emphasis id="F687A077FA47F77DC66B03739F2F9B42" italics="true">Glessula woodthorpei</emphasis>
<comment id="D259C6B7D82DE512BE468C6870F45DE6">
<bibRefCitation id="B88CB3342D2805813E133CFA7B72098A" author="Godwin-Austen HH" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanley's Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 1 - 65. [pls 159 - 165] https: // www. biodiversitylibrary. org / item / 57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen, 1920: 58</bibRefCitation>
, pl. 162, fig. 19. Type locality:
<collectingRegion id="8943129366F02A5F6F1CB6092FA932D6" country="Myanmar" name="Shan">Shan</collectingRegion>
States [
<collectingRegion id="769B9DD6119552438D2C75BA0B6E41D2" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<collectingCountry id="B649C07D2DB0F86EC40D0C7F2F38DD2C" name="Myanmar">Myanmar</collectingCountry>
].
</comment>
</taxonomicName>
</treatmentCitation>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="SECID0EZCBK" type="Type specimens">
<paragraph id="7B506C2E96C38F137A5FBD0977CB5E7D">
<heading id="51C760C85A404A73AC8E5B60CE362063" reason="title">Type specimens.</heading>
</paragraph>
<paragraph id="BCE1F1E5EF43FB910FD1D63B405DA9F0">
<emphasis id="CD87E8DE3341EA434C6649E1DD53B58D" bold="true" italics="true">
<typeStatus id="E226BEB04B03F5FDCE8A5713139F4FC5" type="syntype">Syntypes</typeStatus>
</emphasis>
<named-content id="5FF000387F7A428394C99B0326ED5052" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="2330440B86BE7FA83977AB5299958074" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.1628 [re-registered in error as 1986051] (9 shells; Fig.
<figureCitation id="4B22D48466F6272496FD9CB56B23AB93" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 C E</figureCitation>
) collected by Woodthorpe ex. Godwin-Austen collection from
<collectingRegion id="B7BBD5EBB340C72EDBF85C8D3E8B71D3" country="Myanmar" name="Shan">Shan</collectingRegion>
States.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ENDBK" type="material">
<paragraph id="E7369D3109C0ED5C3F87D221C36047FC">
<heading id="531A792CA1CC656DA46F99CDB6229DFC" reason="title">Other material.</heading>
</paragraph>
<paragraph id="EBE0899CA8F1472A8DCDD3FC2AC117D2">
<named-content id="8DA534A5B96C7D49E76FE8F2830DEECC" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="A1E47AA8C8F5E545443D832646AF370F" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.3655 (2 shells) collected by Woodthorpe ex. Godwin-Austen collection from
<collectingCountry id="20338068FE5F67098D9FFA595553481B" name="Thailand">Siam</collectingCountry>
N. W. Boundary.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EYDBK" type="diagnosis">
<paragraph id="1C74A16EB26983B4573C6090EDCA2353">
<heading id="4F9AF1BBB50AE05832155A0E61FF82C9" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="851E80E36AC534F150294DA26D56697B">Shell oblong conical and regularly attenuated; spire high conical; embryonic whorls rounded; subsequent whorls nearly smooth and equally spaced radial ridges appeared near suture. Suture impressed and whorls slightly convex. Aperture ovate; columella strong, concave, and truncated.</paragraph>
</subSubSection>
<subSubSection id="SECID0E4DBK" type="distribution">
<paragraph id="63237EFFF27811E446E8F3871E03F2BA">
<heading id="38A23602C1AE90A273BC342F259A2507" reason="title">Distribution.</heading>
</paragraph>
<paragraph id="EBDCCD56E7E56B68B21B3B3F090A8D4D">
This species is known from
<collectingRegion id="68B47E66B2C7483723AE62056A7B4E1A" country="Myanmar" name="Shan">Shan State</collectingRegion>
,
<collectingCountry id="A498E643E5C5DB552085F76E675EDDD5" name="Myanmar">Myanmar</collectingCountry>
with an additional record from
<collectingCountry id="94CC8B7E3CE5FCFB9FC5A8D6058D75D5" name="Thailand">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ECEBK" type="remarks">
<paragraph id="615BACEDBD96979C4B90B2F1F6796C12">
<heading id="CDC6EA45252FEFE552487267962EB77F" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="BB85E62937D556D4B8483BABC0427D4A">
<bibRefCitation id="1C2524457F5AD7D24C371591717AD431" author="Godwin-Austen HH" firstAuthor="Godwin-Austen" refId="B39" refString="Godwin-Austen HH (1920) Land and freshwater Mollusca of India, including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other islands of the Indian Ocean. Supplementary to Messrs. Theobald and Hanleys Conchologia Indica. London, Taylor &amp; Francis, Volume 3, 165. [pls 159165] https://www.biodiversitylibrary.org/item/57585" url="https://www.biodiversitylibrary.org/item/57585" year="1920">Godwin-Austen (1920)</bibRefCitation>
indicated that
<specimenCount id="C8347DC4F9DAC9685F969233707C3871" count="9" type="generic">nine specimens</specimenCount>
with the catalogue number Type No. 1682
<abbrev content-type="institution" id="ABBRID0EMEBK" xlink_title="used in old labels for the British Museum">
<collectionCode id="539E644D4E120BBC61A5609995BBFEC1">B</collectionCode>
. M
</abbrev>
. were examined. The species description included one set of measurements and an illustration of one shell. The
<named-content id="5AA00750D95B1E72191499AC03D6BAD5" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="70FDF3BCE0CB2D870D9E52CDD831B896" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
type collections contain a lot of
<specimenCount id="E12BD58C35F9BC8321457F6E4780E461" count="9" type="generic">nine specimens</specimenCount>
from the Woodthorpe collection with a label in Godwin-Austens handwriting stating Type. The measurements of the specimen with red wool inside the shell are close to the measurements given in the original description. It likely corresponds to the illustration of the species provided in the original description and it herein figured (Fig.
<figureCitation id="D47B4DAD67BE77BAEDC1B8FBFC94CA60" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 C</figureCitation>
) along with other shells (Fig.
<figureCitation id="9C72C06156E000CE85127F05D7C5DC58" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. A, B Glessula ponsiensis A syntypes NHMUK 1989043 from Ponsee and B specimen NHMUK 1912.4. 16.747 from Ponsee, Yunnan C E Glessula woodthorpei, syntypes NHMUK 1903.7. 1.1628 from Shan States F Glessula yuangensis, holotype NHMUK 1903.7. 1.1312 from Yuang Ha, Shan States." figureDoi="10.3897/zookeys.1208.116083.figure11" httpUri="https://binary.pensoft.net/fig/1104509">11 D, E</figureCitation>
) from the same
<typeStatus id="70E08BC56A18E4B8CBF6B9707399E8A4" type="syntype">syntype</typeStatus>
lot.
</paragraph>
<paragraph id="B1066F9227F7A4258802EC7542A2F036">
Godwin-Austen separately mentioned (1920: 58) another bleached specimen lot
<named-content id="583C8872D00C1355A40BFE6D694342B7" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/natural-history-museum-london" xlink_title="The Natural History Museum, London">
<collectionCode id="972114BD19B6CF612DC4EEDA2FFEDC85" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" type="Museum">NHMUK</collectionCode>
</named-content>
1903.7. 1.3655 (2 shells) from
<collectingCountry id="DA8A40881AA389FA31EBEE6528146E0F" name="Thailand">Thailand</collectingCountry>
. However, this specimen lot does not form part of the type series of this species.
</paragraph>
</subSubSection>
</treatment>
</document>