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<document id="6DE97F721FF5134B2EB6920813252C0F" ID-DOI="10.5281/zenodo.13158554" ID-ISSN="0068-547X" ID-Zenodo-Dep="13158554" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722539346815" checkinUser="felipe" docAuthor="Griswold, Charles, Ubick, Darrell, Ledford, Joel &amp; Polotow, Daniele" docDate="2022" docId="FB2A53331861FFFC47343371A9D8EDFA" docLanguage="en" docName="ProcCalifAcadSci.67.suppl2.1-193.pdf" docOrigin="Proceedings of the California Academy of Sciences 67" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Uduba goodmani Griswold, Ubick, Ledford &amp; Polotow, 2022, new species" docType="treatment" docUuid="775E6518-0481-40E3-A112-A0519FA0B787" docUuidSource="ZooBank" docVersion="2" lastPageNumber="39" masterDocId="07132B4B1844FFDA474F310FAB37E94C" masterDocTitle="A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar" masterLastPageNumber="193" masterPageNumber="1" pageNumber="38" updateTime="1722553396936" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="76F5FDA49AFB7101A5F6FDB0E7AA7C32">
<mods:title id="D97444211300D26C9487B8059CD4E865">A Revision of the Malagasy Crack-Leg Spiders of the Genus Uduba Simon, 1880 (Araneae, Udubidae), with Description of 35 New Species from Madagascar</mods:title>
</mods:titleInfo>
<mods:name id="985DC80AE481CC6FFFBE94A334E3F518" type="personal">
<mods:role id="A934F9B10BE02DED9C22D6A1A8826805">
<mods:roleTerm id="029D7F6D097F39D4CCE240CC728E2B90">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B486A9A03AD12FC7F8CA2227E88C946C">Griswold, Charles</mods:namePart>
<mods:affiliation id="338DBA5B70705C5E32F8F30F2F1BEE05">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="FFB33C6AE4A2CDB58E838F3681CE3C18" type="personal">
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</mods:role>
<mods:namePart id="7406D7C54B64A9BFB7C721D3342441C8">Ubick, Darrell</mods:namePart>
<mods:affiliation id="06BC5B0BDEA3E6DFEBC8B8D3B76C1909">Arachnology Lab, California Academy of Sciences, 55 Music Concourse Drive, San Francisco, CA 94118, USA. E-mail: cgriswold @ calacademy. org; dubick @ calacademy. org (https: // orcid. org / 0000 - 0002 - 3813 - 8174);</mods:affiliation>
</mods:name>
<mods:name id="DF50B72827ED86D2D2963AC3704E3B2B" type="personal">
<mods:role id="AED3E9C63C632172287B0A7DB3679AF9">
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<mods:namePart id="BCA1AFF4E1E1DBC9B32610FC271042E0">Ledford, Joel</mods:namePart>
<mods:affiliation id="96A4EAF5C48CA1DCE0031DF2F4AC53A8">Department of Plant Biology, University of California, Davis, Davis, CA 95616 - 5270, USA. E-mail: jmledford @ ucdavis. edu;</mods:affiliation>
</mods:name>
<mods:name id="72463E686F0BA8802EB1D2154CDDFCF8" type="personal">
<mods:role id="194630C7A8022966FA50D08AA5277823">
<mods:roleTerm id="58C95B7C9C8B6E7489C6566FB5707601">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2F90565EFA9995E15E4BD6513A55E42">Polotow, Daniele</mods:namePart>
<mods:affiliation id="DF37C549C5C07907657110575C30D812">Departamento de Ecologia e Biologia Evolutiva (DEBE), Universidade Federal de São Carlos (UFSCar), Campus São Paulo, Rodovia Washington Luís, Km 235, CEP 13565 - 905 São Carlos, SP, Brazil. E-mail: danielepolotow @ gmail. com; https: // orcid. org / 0000 - 0002 - 1069 - 1808</mods:affiliation>
</mods:name>
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<mods:title id="9AB9465DD5795A5F21B350374E5CD4A1">Proceedings of the California Academy of Sciences</mods:title>
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<mods:part id="C18FBE12664E3181ACD058421B5B89B2">
<mods:date id="1BDB2E1DA496360F76D24EACB9ECF808">2022</mods:date>
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<mods:number id="F5215AB58FCFA14FA660129C0028AD3C">2022-12-15</mods:number>
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<mods:number id="55E4EDF753F532EFE849290BBC68D1E1">67</mods:number>
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<mods:start id="7BD92EEF4624118CC1410D7391696DCD">1</mods:start>
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<treatment id="FB2A53331861FFFC47343371A9D8EDFA" LSID="urn:lsid:zoobank.org:act:775E6518-0481-40E3-A112-A0519FA0B787" httpUri="http://treatment.plazi.org/id/FB2A53331861FFFC47343371A9D8EDFA" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<subSubSection id="3B99B1AE1861FFFF47343371AAFBEBD9" box="[123,460,638,661]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="733CE2251861FFFF47343371AAFBEBD9" blockId="37.[123,589,638,691]" box="[123,460,638,661]" pageId="37" pageNumber="38">
<heading id="287455491861FFFF47343371AAFBEBD9" box="[123,460,638,661]" fontSize="10" level="2" pageId="37" pageNumber="38" reason="4">
<emphasis id="41F73E371861FFFF47343371AAFBEBD9" bold="true" box="[123,460,638,661]" pageId="37" pageNumber="38">
<taxonomicName id="B48399A61861FFFF47343371AA0DEBD9" ID-CoL="C8KJ9" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[123,314,638,661]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="goodmani" status="sp. nov.">
<emphasis id="41F73E371861FFFF47343371AA0DEBD9" bold="true" box="[123,314,638,661]" italics="true" pageId="37" pageNumber="38">Uduba goodmani</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="5AC4834C1861FFFF46073371AAFBEBD9" box="[328,460,638,661]" pageId="37" pageNumber="38" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47343391A97AEBFF" box="[123,589,670,691]" pageId="37" pageNumber="38" type="description">
<paragraph id="733CE2251861FFFF47343391A97AEBFF" blockId="37.[123,589,638,691]" box="[123,589,670,691]" pageId="37" pageNumber="38">
<figureCitation id="EBB8FEA01861FFFF47343391ABD8EBFF" box="[123,239,670,691]" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="37" pageNumber="38">Figures 7 D</figureCitation>
,
<figureCitation id="EBB8FEA01861FFFF47B53391AA11EBFF" box="[250,294,670,691]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="37" pageNumber="38">20 F</figureCitation>
,
<figureCitation id="EBB8FEA01861FFFF467F3391AA4AEBFF" box="[304,381,670,691]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="37" pageNumber="38">77 A–C</figureCitation>
,
<figureCitation id="EBB8FEA01861FFFF46C73391AAF8EBFF" box="[392,463,670,691]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">80 E, F</figureCitation>
, Maps 6, 16.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E433CCA8D7EA37" pageId="37" pageNumber="38" type="materials_examined">
<paragraph id="733CE2251861FFFF47E433CCA8D7EA37" blockId="37.[123,1174,707,1725]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E433CCAA67EB96" bold="true" box="[171,336,707,730]" pageId="37" pageNumber="38">Type material.</emphasis>
<materialsCitation id="C3EBE8781861FFFF461833CCAFA4EA57" collectingDate="1995-11-27" collectingDateMax="1995-12-05" collectingDateMin="1995-11-27" collectionCode="FMNH" collectorName="S. Goodman" country="Madagascar" elevation="1875" location="Reserve Naturelle Integrale d'Andohahela" pageId="37" pageNumber="38" specimenCode="CASENT9006074" specimenCount="1" stateProvince="Toliara" typeStatus="holotype">
<typeStatus id="AC385C871861FFFF461833CCAA8DEB96" box="[343,442,707,730]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
female from camp 5 at
<quantity id="B47B4FC01861FFFF458E33CCA83BEB96" box="[705,780,707,730]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="37" pageNumber="38" unit="m" value="1875.0">
<elevation id="F8AE05161861FFFF458E33CCA83BEB96" box="[705,780,707,730]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="37" pageNumber="38" unit="m" value="1875.0">1875m</elevation>
</quantity>
elevation in
<location id="765CB4FE1861FFFF44D533CCAA66EBB6" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1861FFFF44D533CCAA66EBB6" country="Madagascar" name="Reserve Naturelle Integrale d'Andohahela" pageId="37" pageNumber="38" stateProvince="Toliara">Réserve Naturelle Intégrale d’Andohahela</location>
(24º33.7ʹS, 46º43.3ʹE),
<collectingRegion id="B1472CC71861FFFF451333ECA826EBB6" box="[604,785,739,762]" country="Madagascar" name="Toliara" pageId="37" pageNumber="38">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B51861FFFF445033ECA8A9EBB6" box="[799,926,739,762]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, collected
<date id="073DC4E51861FFFF435633ECAA58EA57" pageId="37" pageNumber="38" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">
<collectingDate id="17793D0D1861FFFF435633ECAA58EA57" pageId="37" pageNumber="38" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">27 November–5 December 1995</collectingDate>
</date>
by
<collectorName id="DE7687F31861FFFF46D0320BA91FEA57" box="[415,552,772,795]" pageId="37" pageNumber="38">S. Goodman</collectorName>
(SG1995-01) (
<specimenCode id="23254A5E1861FFFF459E320BA8ABEA57" box="[721,924,772,795]" collectionCode="CASENT" pageId="37" pageNumber="38">CASENT9006074</specimenCode>
), deposited in
<collectionCode id="15927AE01861FFFF430D320BAFA4EA57" box="[1090,1171,772,795]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="37" pageNumber="38" type="Museum">FMNH</collectionCode>
</materialsCitation>
.
<materialsCitation id="C3EBE8781861FFFF4734322BA8ECEA37" collectingDate="2002-01-21" collectingDateMax="2002-01-25" collectingDateMin="2002-01-21" collectionCode="CAS" country="Madagascar" elevation="900" latitude="-24.76389" location="Col du Sedro in Parc National d'Andohahela" longLatPrecision="21" longitude="46.751667" municipality="Fisher-Griswold Arthropod Team" pageId="37" pageNumber="38" specimenCode="CASENT9009538" specimenCount="1" stateProvince="Toliara" typeStatus="paratype">
<typeStatus id="AC385C871861FFFF4734322BABEFEA77" box="[123,216,804,827]" pageId="37" pageNumber="38" type="paratype">Paratype</typeStatus>
female (
<specimenCode id="23254A5E1861FFFF4677322BA936EA77" box="[312,513,804,827]" collectionCode="CASENT" pageId="37" pageNumber="38">CASENT9009538</specimenCode>
) from montane rainforest at
<quantity id="B47B4FC01861FFFF4474322BA84FEA77" box="[827,888,804,827]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="38" unit="m" value="900.0">
<elevation id="F8AE05161861FFFF4474322BA84FEA77" box="[827,888,804,827]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="37" pageNumber="38" unit="m" value="900.0">900m</elevation>
</quantity>
elevation at
<location id="765CB4FE1861FFFF434A322BAAFCEA17" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1861FFFF434A322BAAFCEA17" country="Madagascar" latitude="-24.76389" longLatPrecision="21" longitude="46.751667" municipality="Fisher-Griswold Arthropod Team" name="Col du Sedro in Parc National d'Andohahela" pageId="37" pageNumber="38" stateProvince="Toliara">Col du Sedro in Parc National d’Andohahela</location>
(
<geoCoordinate id="16B784E21861FFFF4694324BA961EA17" box="[475,598,836,859]" degrees="24" direction="south" minutes="45" orientation="latitude" pageId="37" pageNumber="38" precision="15" seconds="50" value="-24.76389">24°45ʹ50ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21861FFFF452B324BA9E3EA17" box="[612,724,836,859]" degrees="46" direction="east" minutes="45" orientation="longitude" pageId="37" pageNumber="38" precision="15" seconds="6" value="46.751667">46°45ʹ6ʺE</geoCoordinate>
),
<collectingRegion id="B1472CC71861FFFF45A7324BA8ABEA17" box="[744,924,836,859]" country="Madagascar" name="Toliara" pageId="37" pageNumber="38">Toliara Province</collectingRegion>
,
<collectingCountry id="0B94A2B51861FFFF44E8324BAF11EA17" box="[935,1062,836,859]" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
, collected
<date id="073DC4E51861FFFF4734326BAA62EA37" box="[123,341,868,891]" pageId="37" pageNumber="38" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">
<collectingDate id="17793D0D1861FFFF4734326BAA62EA37" box="[123,341,868,891]" pageId="37" pageNumber="38" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">21–25 January 2002</collectingDate>
</date>
by the
<collectingMunicipality id="9358785F1861FFFF46E6326BA839EA37" box="[425,782,868,891]" pageId="37" pageNumber="38">Fisher-Griswold Arthropod Team</collectingMunicipality>
, deposited in
<collectionCode id="15927AE01861FFFF44EB326BA8ECEA37" box="[932,987,868,891]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="37" pageNumber="38" type="Museum">CAS</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E4328BAA7DEA97" pageId="37" pageNumber="38" type="etymology">
<paragraph id="733CE2251861FFFF47E4328BAA7DEA97" blockId="37.[123,1174,707,1725]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E4328BAA1DEAD7" bold="true" box="[171,298,900,923]" pageId="37" pageNumber="38">Etymology.</emphasis>
The species epithet is a patronym in honor of Steve Goodman, naturalist, author, collector and mammologist who has made substantial contributions to our knowledge of
<collectingCountry id="0B94A2B51861FFFF431F32ABAB88EA97" name="Madagascar" pageId="37" pageNumber="38">Madagascar</collectingCountry>
biodiversity.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFF47E432EBA8CFEDF0" pageId="37" pageNumber="38" type="diagnosis">
<paragraph id="733CE2251861FFFF47E432EBA8CFEDF0" blockId="37.[123,1174,707,1725]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E432EBAADEEAB7" bold="true" box="[171,489,996,1019]" pageId="37" pageNumber="38">
Diagnosis.
<taxonomicName id="B48399A61861FFFF466732EBAADEEAB7" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[296,489,996,1019]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371861FFFF466732EBAADEEAB7" bold="true" box="[296,489,996,1019]" italics="true" pageId="37" pageNumber="38">Uduba goodmani</emphasis>
</taxonomicName>
</emphasis>
belong to Group II, the Epigynum lateral projection group, or
<taxonomicName id="B48399A61861FFFF4734350BAA76ED57" authorityName=", Polotow et al." authorityYear="2015" baseAuthorityName="Dahl" baseAuthorityYear="1901" box="[123,321,1028,1051]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="evanescens">Uduba evanescens</taxonomicName>
group (Map 16). They are ecribellate. Females can be distinguished from those of other
<taxonomicName id="B48399A61861FFFF479A352BAA44ED77" box="[213,371,1060,1083]" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF479A352BAA28ED77" bold="true" box="[213,287,1060,1083]" italics="true" pageId="37" pageNumber="38">Uduba</emphasis>
species
</taxonomicName>
by the form of the female genitalia (
<figureCitation id="EBB8FEA01861FFFF45B2352BA8BAED77" box="[765,909,1060,1083]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="37" pageNumber="38">Figs. 77 A–C</figureCitation>
.
<figureCitation id="EBB8FEA01861FFFF44D7352BA8D1ED77" box="[920,998,1060,1083]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">80 E, F</figureCitation>
<emphasis id="41F73E371861FFFF44A9352BA8CFED77" bold="true" box="[998,1016,1060,1083]" pageId="37" pageNumber="38">):</emphasis>
epigynal plate a broad oval, width 1.50 times length, lateral lobes with convex edges forming ridges on each side (
<figureCitation id="EBB8FEA01861FFFF47CC356BABDEED37" box="[131,233,1124,1147]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">Fig. 80 E</figureCitation>
); vulva broad, width one and-a-half times length with spermathecal ducts characteristic, forming one and-a-half loose, transverse loops (
<figureCitation id="EBB8FEA01861FFFF45CB358AA9DBEDD0" box="[644,748,1157,1180]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">Fig. 80 F</figureCitation>
), copulatory duct lateral and extending mesally to lead to posterior, widely spaced fertilization ducts. Males are unknown.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1861FFFC47E435CAAA53EA19" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38" type="description">
<paragraph id="733CE2251861FFFC47E435CAA96FE8DF" blockId="37.[123,1174,707,1725]" lastBlockId="38.[123,1174,220,1206]" lastPageId="38" lastPageNumber="39" pageId="37" pageNumber="38">
<emphasis id="41F73E371861FFFF47E435CAAA40ED90" bold="true" box="[171,375,1221,1244]" pageId="37" pageNumber="38">Description. Male</emphasis>
: Unknown.
<emphasis id="41F73E371861FFFF46B135CAA966ED90" bold="true" box="[510,593,1221,1244]" pageId="37" pageNumber="38">Female</emphasis>
(
<typeStatus id="AC385C871861FFFF452F35CAA9FDED90" box="[608,714,1221,1244]" pageId="37" pageNumber="38" type="holotype">Holotype</typeStatus>
): Total length 16.43. Markings as in
<figureCitation id="EBB8FEA01861FFFF431935CAABA6EDB0" captionStart="FIGURE" captionStartId="111.[155,167,1458,1477]" captionTargetBox="[182,1114,216,1438]" captionTargetId="figure-16@111.[182,1114,216,1438]" captionTargetPageId="111" captionText="FIGURE 7. Habitus photos of Uduba spp., females, dorsal views. A. U. pseudoevanescens, CASENT9006007, from Forêt Marovato, MNHN Paris. B. U. evanescens (Dahl, 1901), CASENT9006090 from Ranomafana National Park. C. U. volana, Holotype (CASENT9006072) from 1400m elevation at Ambohimanga. D. U. goodmani, Holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela. E. U. salegy, Paratype (CASENT9006045) from Parc National Montagne d’Ambre. F. U. ibonia, CASENT9006033 from Ranomafana National Park. G. U. schlingeri, CASENT9006075 from Parc National Ranomafana. H. U. barbarae, paratype (CASENT9006064), from Lokobe Forest on the island of Nosy Be. I. U. kavanaughi, paratype (CASENT9003517) from Parc National Ranomafana. J. U. woodae, paratype (CASENT9017834) from spiny forest at Lake Ranobe. K. U. woodae, CASENT9006092 from Ambovombe, Carougeau, Muséum Paris. L. U. madagascariensis (Vinson, 1863), CASENT9006029, from forest near village of Ambohimanga. Scale bars (see A, I) = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158572" httpUri="https://zenodo.org/record/13158572/files/figure.png" pageId="37" pageNumber="38">Fig. 7 D</figureCitation>
. Carapace 8.50 long, 5.71 wide, 3.50 high; clypeus 0.43 high. Eye diameters: AME 0.27, ALE and PME 0.36, PLE 0.38. Chelicerae 3.79 long; sternum 3.71 long, 2.79 wide; labium 1.82 long; palpal coxae 2.82 long. With a colulus (
<figureCitation id="EBB8FEA01861FFFF456A342AA9BEEC70" box="[549,649,1317,1340]" captionStart="FIGURE" captionStartId="121.[155,167,1018,1037]" captionTargetBox="[129,1168,234,986]" captionTargetId="figure-16@121.[120,1176,216,1003]" captionTargetPageId="121" captionText="FIGURE 20. Uduba spp., females: cribellum or colulus, and base of anterior lateral spinnerets surrounded by soft, flexible cuticle, anterior to top. A. U. halabe, CASENT9017873 from Parc Nacional Ranomafana, entire cribellum. B. U. ibonia, CASENT9006035 from Parc Nacional Ranomafana, divided cribellum (note small size of spinning plates). C. U. pseudoevanescens, CASENT9006008 from Parc Nacional Périnet, divided cribellum. D. U. evanescens (Dahl, 1901), CASENT9024032 from Parc Nacional Ranomafana, colulus. E. U. madagascariensis (Vinson, 1863), CASENT9006028 from Forêt Analavelona, colulus. F. U. goodmani, holotype (CASENT9006074) from Réserve Naturelle Intégrale d’Andohahela, colulus. G. U. schlingeri, CASENT9006028 from Parc Nacional Ranomafana, divided cribellum (note large size of spinning plates at posterior margin of soft cuticle). Illustrations by Charles Griswold. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.13158606" httpUri="https://zenodo.org/record/13158606/files/figure.png" pageId="37" pageNumber="38">Fig. 20 F</figureCitation>
<emphasis id="41F73E371861FFFF45C6342AA9A5EC70" bold="true" box="[649,658,1317,1340]" pageId="37" pageNumber="38">)</emphasis>
. Spination: palpus–femur d0-1-1, p0-0-1, patella p1-0, tibia p2-1, tarsus p2-1, r1-0-0; leg I–femur d1-0-0, p0-0-1, tibia v2-2-2-0, metatarsus v2- 2-3; leg II–femur d1-0-0, p0-0-1, tibia p0-0-1-0, v1-1-1-0, metatarsus v2-2-3; leg III–femur d0-0- 0-1, p1-1-0-0-2, r1-1-0-1, tibia d0-1-0, p0-1-1-0, v1-2-0, r0-1-1-0, metatarsus p1-1-2, v2-2-2, r1- 1-2; leg IV–femur d1-0-1-0, tibia v1-1-2, metatarsus p0-0-2-2, v2-1-2-2, r1-2-2. Scopulae: strong ventral beneath all tarsi and metatarsi and beneath tibiae I and II. Leg measurements (
<typeStatus id="AC385C871861FFFF437D34CAAFA5EC90" box="[1074,1170,1477,1500]" pageId="37" pageNumber="38" type="paratype">paratype</typeStatus>
, CASENT9009538): I: 4.20 + 2.00 + 3.60 + 3.10 + 2.00 = 14.90; II: 3.90 + 2.00 + 3.10 + 2.90 + 1.90 = 13.80; III: 3.20 + 1.50 + 1.90 + 2.80 + 1.55 = 10.95; IV: 4.20 + 1.80 + 3.60 + 4.10 + 1.90 = 15.60; palpus: 2.10 + 1.00 + 1.10 + NA + 1.80 = 6.00. Leg formula 4123.
<emphasis id="41F73E371861FFFF434E3729AB9AEF11" bold="true" pageId="37" pageNumber="38">Female genitalia</emphasis>
(
<typeStatus id="AC385C871861FFFF47F33749AA17EF11" box="[188,288,1606,1629]" pageId="37" pageNumber="38" type="holotype">holotype</typeStatus>
): epigynal plate without depressed atrium, lateral lobes extending to sides forming blunt, earlike lobes (
<figureCitation id="EBB8FEA01861FFFF461A3769AAD1EF31" box="[341,486,1638,1661]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="37" pageNumber="38">Figs. 77 A, C</figureCitation>
), plate width 1.63 times length; width between bases of LL 0.72 times epigynum width; lateral ears short and blunt (
<figureCitation id="EBB8FEA01861FFFF45E43789A823EFD1" box="[683,788,1670,1693]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="37" pageNumber="38">Fig. 80 E</figureCitation>
), arising near epigastric furrow, lateral ears (LL side) width 0.75 times LL side length, (LL side) width 0.16 times width epigynum; median lobe a flat plate, each side with an oblique bulge where copulatory openings are found, ML width 0.64 times epigynum width, 1.36 times ML length, CO origin at 0.485 epigynum length from epigastric furrow, distance between copulatory openings 0.41 times epigynum width. Vulva (
<figureCitation id="EBB8FEA01862FFFC47CC3033ABDEE81F" box="[131,233,316,339]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="38" pageNumber="39">Fig. 77 B</figureCitation>
) with copulatory ducts lateral, median to these spermathecal ducts forming a single longitudinal loop (
<figureCitation id="EBB8FEA01862FFFC46523053AAB7E83F" box="[285,384,348,371]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="38" pageNumber="39">Fig. 80 F</figureCitation>
), vulva length 0.53 times width, fertilization ducts widely separated, vulva width only 3.21 times distance between FD.
</paragraph>
<paragraph id="733CE2251862FFFC47E43093AA53EA19" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43093AA2DE8FF" bold="true" box="[171,282,412,435]" pageId="38" pageNumber="39">Variation.</emphasis>
(N=2): Carapace length / width = 1.49–1.51, carapace height / width = 0.61–0.64, PER / carapace width = 0.45–0.69, PER/OAL = 2.70–2.90, PER/AER = 1.30–1.32, OAL/OQL = 1.00–1.25, OQP/ OQA = 1.07–1.18, clypeus height / AME = 1.60–1.75, cheliceral length / clypeus height = 7.71–8.83, sternum length / width = 1.22–1.33, palpal coxa length / width = 2.11–2.39, femur I length / carapace width = 1.08–1.14, metatarsus I length / carapace width = 0.58–0.79, femur IV length / carapace width = 1.08–1.09, palpal tarsus length / carapace width = 0.44–0.46, palpal tibia length / palpal patella length = 1.59–1.80, palpal tarsus length / palpal tibia length = 1.40–1.64, palpal tarsus length / palpal femur length = 0.80–0.86, palpal tibia length / palpal patella length = 1.10–1.14. Epigynal plate (
<figureCitation id="EBB8FEA01862FFFC456D3392A98FEBF8" box="[546,696,669,692]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="38" pageNumber="39">Figs. 77 A, C</figureCitation>
,
<figureCitation id="EBB8FEA01862FFFC458A3392A9CDEBF8" box="[709,762,669,692]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="38" pageNumber="39">80 E</figureCitation>
) width 1.50–1.63 times length, width between bases of LL 0.72–0.75 times epigynum width, CO origin at 0.43–0.48 epigynum length from epigastric furrow, distance between copulatory openings 0.33–0.41 times epigynum width; vulva (
<figureCitation id="EBB8FEA01862FFFC478933F2AA0CEA58" box="[198,315,765,788]" captionStart="FIGURE" captionStartId="165.[155,167,1212,1231]" captionTargetBox="[120,1176,216,1196]" captionTargetId="figure-16@165.[120,1176,216,1196]" captionTargetPageId="165" captionText="FIGURE 77. Uduba spp., female genitalia drawings. A–C. U. goodmani, new species. A, B. Paratype female (CASENT9009538) from montane rainforest at 900m elevation at Col du Sedro in Parc National d’Andohahela. C. Holotype (CASENT9006074) from camp #5 at 1875m elevation in Réserve Naturelle Intégrale d’Andohahela. D–F. U. volana, new species, holotype (CASENT9006072) dug from a burrow at 1400m elevation at Ambohimanga. A, C, D, E. Epigynum, ventral. B, F. Vulva, dorsal. Illustrations C–D by Jenny Speckels, A–B, E–F by Charles Griswold. BS, base of spermatheca of vulva; CD, copulatory ducts of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.13158745" httpUri="https://zenodo.org/record/13158745/files/figure.png" pageId="38" pageNumber="39">Figs. 77 B</figureCitation>
,
<figureCitation id="EBB8FEA01862FFFC460833F2AA4DEA58" box="[327,378,765,788]" captionStart="FIGURE" captionStartId="168.[155,167,859,878]" captionTargetBox="[120,1176,219,859]" captionTargetId="figure-11@168.[120,1176,216,847]" captionTargetPageId="168" captionText="FIGURE 80. Uduba spp., ♀ genitalia. A, B. U. schlingeri, new species, CASENT9006075 from Vatoharanana, Ranomafana National Park. C. U. madagascariensis (Vinson, 1863), CASENT9006029 from forest near Ambohimanga Village. D. U. woodae, new species, CASENT9006092 from Ambovombe, Carougeau, MNHN, Muséum Paris. E, F. U. goodmani, new species, Holotype (CASENT9006074), from Réserve Naturelle Intégrale d’Andohahela. G. U. evanescens (Dahl, 1901) CASENT9006083 from Forêt Ambohimasoa. H, I. U. valiha, new species, Paratype (CASENT9006071) from Massif Andringitra. J. U. pseudoevanescens, new species, CASENT9006007 from Forêt Marovato. A, E, H. Epigynum, ventral. B–D, F, G, I, J. Vulva, dorsal. At, atrium of epigynum; BS, base of spermatheca of vulva; CO, copulatory openings of epigynum; EF, epigastric furrow of abdomen; FD, fertilization duct of vulva; HS, head of spermatheca of vulva, including spermathecal poreplate; LL, lateral lobe of epigynum; LP, lateral process of epigynum; ML, median lobe of epigynum. Scale bars = A, B = 0.5 mm, C–J = 0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.13158753" httpUri="https://zenodo.org/record/13158753/files/figure.png" pageId="38" pageNumber="39">80 F</figureCitation>
) with spermathecal ducts forming one and-a-half loose, transverse loops, vulva length 0.53–0.64 times width, fertilization ducts widely spaced, vulva width 2.58–3.21 times distance between FD.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1862FFFC47E43251ABC7ED3A" pageId="38" pageNumber="39" type="materials_examined">
<paragraph id="733CE2251862FFFC47E43251AAF5ED79" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43251AABCEA39" bold="true" box="[171,395,862,885]" pageId="38" pageNumber="39">Material examined.</emphasis>
<materialsCitation id="C3EBE8781862FFFC46D63251AF6FEAF9" collectingDate="1995-11-27" collectingDateMax="1995-12-05" collectingDateMin="1995-11-27" collectionCode="FMNH" collectorName="SE Andranondambo &amp; S. Goodman" country="Madagascar" elevation="1875" location="Reserve Naturelle Integrale d'Andohahela" pageId="38" pageNumber="39" specimenCode="CASENT9006074" specimenCount="1" specimenCount-female="1" stateProvince="Toliara" typeStatus="holotype">
<collectingCountry id="0B94A2B51862FFFC46D63251A96CEA39" box="[409,603,862,885]" name="Madagascar" pageId="38" pageNumber="39">MADAGASCAR</collectingCountry>
:
<emphasis id="41F73E371862FFFC45293251A820EA39" box="[614,791,862,885]" italics="true" pageId="38" pageNumber="39">
<collectingRegion id="B1472CC71862FFFC45293251A820EA39" box="[614,791,862,885]" country="Madagascar" name="Toliara" pageId="38" pageNumber="39">Toliara Province</collectingRegion>
</emphasis>
:
<location id="765CB4FE1862FFFC44693251ABE9EAD9" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1862FFFC44693251ABE9EAD9" country="Madagascar" name="Reserve Naturelle Integrale d'Andohahela" pageId="38" pageNumber="39" stateProvince="Toliara">Réserve Naturelle Intégrale d’Andohahela</location>
, 20.0 km
<collectorName id="DE7687F31862FFFC46093271A929EAD9" box="[326,542,894,917]" pageId="38" pageNumber="39">SE Andranondambo</collectorName>
, camp 5, 24º33.7ʹS, 46º43.3ʹE, elev.
<quantity id="B47B4FC01862FFFC44EF3271A8DAEAD9" box="[928,1005,894,917]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="38" pageNumber="39" unit="m" value="1875.0">
<elevation id="F8AE05161862FFFC44EF3271A8DAEAD9" box="[928,1005,894,917]" metricMagnitude="3" metricUnit="m" metricValue="1.875" pageId="38" pageNumber="39" unit="m" value="1875.0">1875m</elevation>
</quantity>
,
<date id="073DC4E51862FFFC44B83271AA75EAF9" pageId="38" pageNumber="39" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">
<collectingDate id="17793D0D1862FFFC44B83271AA75EAF9" pageId="38" pageNumber="39" value="1995-11-27" valueMax="1995-12-05" valueMin="1995-11-27">27 November– 5 December 1995</collectingDate>
</date>
,
<collectorName id="DE7687F31862FFFC461F3291AAEEEAF9" box="[336,473,926,949]" pageId="38" pageNumber="39">S. Goodman</collectorName>
[SG1995-01] (
<typeStatus id="AC385C871862FFFC45CE3291A9DFEAF9" box="[641,744,926,949]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
,
<specimenCode id="23254A5E1862FFFC45B83291A888EAF9" box="[759,959,926,949]" collectionCode="CASENT" pageId="38" pageNumber="39">CASENT9006074</specimenCode>
,
<specimenCount id="658529AC1862FFFC44823291A8D9EAF9" box="[973,1006,926,949]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01862FFFC44B03291AF66EAF9" box="[1023,1105,926,949]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="38" pageNumber="39" type="Museum">FMNH</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="C3EBE8781862FFFC43293291AA88ED79" collectingDate="2002-01-21" collectingDateMax="2002-01-25" collectingDateMin="2002-01-21" collectionCode="CAS" collectorName="Col du Sedro &amp; Fisher-Griswold Arthropod Team" country="Madagascar" elevation="900" latitude="-24.76389" location="Col du Sedro" longLatPrecision="21" longitude="46.751667" municipality="Parc National d'Andohahela" pageId="38" pageNumber="39" specimenCode="CASENT9009538" specimenCount="1" specimenCount-female="1" stateProvince="Toliara Province" typeStatus="paratype">
<collectingMunicipality id="9358785F1862FFFC43293291AAB5EA99" pageId="38" pageNumber="39">Parc National d’Andohahela</collectingMunicipality>
,
<collectorName id="DE7687F31862FFFC46D732B1A900EA99" box="[408,567,958,981]" pageId="38" pageNumber="39">
<location id="765CB4FE1862FFFC46D732B1A900EA99" LSID="urn:lsid:plazi:treatment:FB2A53331861FFFC47343371A9D8EDFA:765CB4FE1862FFFC46D732B1A900EA99" box="[408,567,958,981]" country="Madagascar" latitude="-24.76389" longLatPrecision="21" longitude="46.751667" municipality="Parc National d'Andohahela" name="Col du Sedro" pageId="38" pageNumber="39" stateProvince="Toliara Province">Col du Sedro</location>
</collectorName>
,
<locationDeviation id="628EFB071862FFFC450132B1A8ACEA99" box="[590,923,958,981]" location="Mahamavo" pageId="38" pageNumber="39">
<quantity id="B47B4FC01862FFFC450132B1A996EA99" box="[590,673,958,981]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="38" pageNumber="39" unit="km" value="3.8">3.8 km</quantity>
113°ESE Mahamavo
</locationDeviation>
,
<locationDeviation id="628EFB071862FFFC44FE32B1ABDFEAB9" location="Tolagnaro" pageId="38" pageNumber="39">
<quantity id="B47B4FC01862FFFC44FE32B1AF25EA99" box="[945,1042,958,981]" metricMagnitude="4" metricUnit="m" metricValue="3.76" pageId="38" pageNumber="39" unit="km" value="37.6">37.6 km</quantity>
341°NNW Tolagnaro
</locationDeviation>
,
<geoCoordinate id="16B784E21862FFFC47B532D1AA43EAB9" box="[250,372,990,1013]" degrees="24" direction="south" minutes="45" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="50" value="-24.76389">24°45ʹ50ʺS</geoCoordinate>
,
<geoCoordinate id="16B784E21862FFFC46C732D1AACFEAB9" box="[392,504,990,1013]" degrees="46" direction="east" minutes="45" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="6" value="46.751667">46°45ʹ6ʺE</geoCoordinate>
, montane rainforest at
<quantity id="B47B4FC01862FFFC444332D1A87EEAB9" box="[780,841,990,1013]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="39" unit="m" value="900.0">
<elevation id="F8AE05161862FFFC444332D1A87EEAB9" box="[780,841,990,1013]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="38" pageNumber="39" unit="m" value="900.0">900m</elevation>
</quantity>
elevation, general collecting ground spiders,
<date id="073DC4E51862FFFC461F32F1A962ED59" box="[336,597,1022,1045]" pageId="38" pageNumber="39" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">
<collectingDate id="17793D0D1862FFFC461F32F1A962ED59" box="[336,597,1022,1045]" pageId="38" pageNumber="39" value="2002-01-21" valueMax="2002-01-25" valueMin="2002-01-21">21–25 January 2002</collectingDate>
</date>
,
<collectorName id="DE7687F31862FFFC453D32F1AF3EED59" box="[626,1033,1022,1045]" pageId="38" pageNumber="39">Fisher-Griswold Arthropod Team</collectorName>
(
<typeStatus id="AC385C871862FFFC436832F1AFA6ED59" box="[1063,1169,1022,1045]" pageId="38" pageNumber="39" type="paratype">Paratype</typeStatus>
,
<specimenCode id="23254A5E1862FFFC47343511AA75ED79" box="[123,322,1054,1077]" collectionCode="CASENT" pageId="38" pageNumber="39">CASENT9009538</specimenCode>
,
<specimenCount id="658529AC1862FFFC46013511AA47ED79" box="[334,368,1054,1077]" count="1" pageId="38" pageNumber="39" type="female">1♀</specimenCount>
,
<collectionCode id="15927AE01862FFFC46313511AA81ED79" box="[382,438,1054,1077]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="38" pageNumber="39" type="Museum">CAS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="733CE2251862FFFC47E43530ABC7ED3A" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43530AA69ED1A" bold="true" box="[171,350,1087,1110]" pageId="38" pageNumber="39">Natural history.</emphasis>
Collection data suggest that the species
<taxonomicName id="B48399A61862FFFC44423530A8FBED1A" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[781,972,1087,1110]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371862FFFC44423530A8FBED1A" bold="true" box="[781,972,1087,1110]" italics="true" pageId="38" pageNumber="39">Uduba goodmani</emphasis>
</taxonomicName>
occurs in montane rain forest.
</paragraph>
</subSubSection>
<subSubSection id="3B99B1AE1862FFFC47E43570A9D8EDFA" pageId="38" pageNumber="39" type="distribution">
<paragraph id="733CE2251862FFFC47E43570A9D8EDFA" blockId="38.[123,1174,220,1206]" pageId="38" pageNumber="39">
<emphasis id="41F73E371862FFFC47E43570A931EDDA" bold="true" box="[171,518,1151,1174]" pageId="38" pageNumber="39">
Distribution.
<taxonomicName id="B48399A61862FFFC460A3570A931EDDA" authorityName="Griswold &amp; Ubick &amp; Ledford &amp; Polotow" authorityYear="2022" box="[325,518,1151,1174]" class="Arachnida" family="Udubidae" genus="Uduba" kingdom="Animalia" order="Araneae" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="goodmani">
<emphasis id="41F73E371862FFFC460A3570A931EDDA" bold="true" box="[325,518,1151,1174]" italics="true" pageId="38" pageNumber="39">Uduba goodmani</emphasis>
</taxonomicName>
</emphasis>
are known only from two nearby localities on mountains at Andohahela in far southeastern
<collectingCountry id="0B94A2B51862FFFC469C3590A963EDFA" box="[467,596,1183,1206]" name="Madagascar" pageId="38" pageNumber="39">Madagascar</collectingCountry>
(Maps 6, 16
<emphasis id="41F73E371862FFFC45903590A9DFEDFA" bold="true" box="[735,744,1183,1206]" pageId="38" pageNumber="39">)</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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