329 lines
39 KiB
XML
329 lines
39 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.79.e62332" ID-GBIF-Dataset="c6cd0951-6d64-4b2f-b920-bb7ba439eb39" ID-GBIF-Taxon="189469056" ID-Pensoft-Pub="1864-8312-79-509" ID-Pensoft-UUID="F1742697BB0C564B9BA5036415F063D4" ID-ZooBank="951E28C6EAC0494EBC255DEF1991583B" LSID="urn:lsid:zoobank.org:act:D847F421-25EA-4FA2-8989-38837493CD19" ModsDocID="1864-8312-79-509" checkinTime="1633953041321" checkinUser="pensoft" docAuthor="Greenberg, Marleen R., Huey, Joel A., Framenau, Volker W. & Harms, Danilo" docDate="2021" docId="6294FFA5B7095301A471739A872F3A1B" docLanguage="en" docName="ArthropodSystPhyl 79: 509-533" docOrigin="Arthropod Systematics & amp; Phylogeny 79" docPubDate="2021-10-11" docSource="http://dx.doi.org/10.3897/asp.79.e62332" docTitle="Missulena manningensis Greenberg & Huey & Framenau & Harms 2021, sp. nov." docType="treatment" docUuid="D847F421-25EA-4FA2-8989-38837493CD19" docUuidSource="ZooBank" docVersion="3" id="F1742697BB0C564B9BA5036415F063D4" lastPageNumber="509" masterDocId="F1742697BB0C564B9BA5036415F063D4" masterDocTitle="Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome" masterLastPageNumber="533" masterPageNumber="509" pageId="0" pageNumber="509" updateTime="1635284731334" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Three new species of mouse spider (Araneae: Actinopodidae: Missulena Walckenaer, 1805) from Western Australia, including an assessment of intraspecific variability in a widespread species from the arid biome</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Greenberg, Marleen R.</mods:namePart>
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<mods:affiliation>Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany & Fachbereich Biologie, Universitaet Hamburg, Ohnhorststrasse 18, 22609 Hamburg, Germany</mods:affiliation>
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<mods:nameIdentifier type="email">marleen.greenberg@outlook.de</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Huey, Joel A.</mods:namePart>
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<mods:affiliation>Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Framenau, Volker W.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7724-3831</mods:nameIdentifier>
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<mods:affiliation>Harry Butler Institute, Murdoch University, 90 South Street, Murdoch 6150, Australia & Department of Terrestrial Zoology, Western Australian Museum, Welshpool, WA 6106, Australia & Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Harms, Danilo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7189-5345</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum Hamburg, Leibniz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-10-11</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>79</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>509</mods:start>
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<mods:end>533</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/asp.79.e62332</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.79.e62332</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-79-509</mods:identifier>
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<mods:identifier type="ZooBank">951E28C6EAC0494EBC255DEF1991583B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F1742697BB0C564B9BA5036415F063D4</mods:identifier>
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</mods:mods>
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<subSection pageId="0" pageNumber="509" type="3. taxonomy">
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<treatment ID-GBIF-Taxon="189469056" LSID="urn:lsid:zoobank.org:act:D847F421-25EA-4FA2-8989-38837493CD19" httpUri="http://treatment.plazi.org/id/6294FFA5B7095301A471739A872F3A1B" lastPageNumber="509" pageId="0" pageNumber="509">
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<subSubSection pageId="0" pageNumber="509" type="nomenclature">
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<paragraph pageId="0" pageNumber="509">
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3.3.
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<taxonomicName LSID="http://zoobank.org/D847F421-25EA-4FA2-8989-38837493CD19" authority="Greenberg & Huey & Framenau & Harms, 2021" authorityName="Greenberg & Huey & Framenau & Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena manningensis" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="manningensis" status="sp. nov.">Missulena manningensis</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="509">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="509" type="description">
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<paragraph pageId="0" pageNumber="509">
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Missulena manningensis sp. nov. and Missulena iugum sp. nov. in Western Australia." figureDoi="10.3897/asp.79.e62332.figure10" httpUri="https://binary.pensoft.net/fig/597116" pageId="0" pageNumber="509">Figs 10</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">, 12</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="509" type="type material">
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<paragraph pageId="0" pageNumber="509">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="509">
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<emphasis bold="true" pageId="0" pageNumber="509">Holotype</emphasis>
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: AUSTRALIA - Western Australia • ♂; Mt Manning area, site CR2;
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<geoCoordinate degrees="30" direction="south" minutes="27.95" orientation="latitude" precision="9" value="-30.465834">30°27.95′S</geoCoordinate>
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<geoCoordinate degrees="119" direction="east" minutes="58.02" orientation="longitude" precision="9" value="119.967">119°58.02′E</geoCoordinate>
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; 21 June 2008; J. Francesconi leg.; WAM T92071.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="509" type="diagnosis">
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<paragraph pageId="0" pageNumber="509">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="509">
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Males share with
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<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
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</taxonomicName>
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sp. nov.,
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<taxonomicName genus="M." lsidName="M. iugum" pageId="0" pageNumber="509" rank="species" species="iugum">
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<emphasis italics="true" pageId="0" pageNumber="509">M. iugum</emphasis>
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</taxonomicName>
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sp. nov.,
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<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
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</taxonomicName>
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,
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<taxonomicName genus="M." lsidName="M. occatoria" pageId="0" pageNumber="509" rank="species" species="occatoria">
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<emphasis italics="true" pageId="0" pageNumber="509">M. occatoria</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
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<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
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</taxonomicName>
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, the closest morphological matches, the red colouration of the chelicerae and pars cephalica. They differ from
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<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
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</taxonomicName>
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by a longer carapace (>3.00 mm;
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<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
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</taxonomicName>
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up to 2.8 mm) and the presence of strong, conical spines of the rastellum (simple in
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<taxonomicName genus="M." lsidName="M. langlandsi" pageId="0" pageNumber="509" rank="species" species="langlandsi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. langlandsi</emphasis>
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</taxonomicName>
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). They differ from
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<taxonomicName genus="M." lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
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<emphasis italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
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</taxonomicName>
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and
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<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
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<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
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</taxonomicName>
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by the lack of spines ventrally on patellae III and IV (at the most 1 thickened seta). Pars cephalica lower than in
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<taxonomicName genus="M." lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
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<emphasis italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
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</taxonomicName>
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(up to 1.96;
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<taxonomicName genus="M." lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
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<emphasis italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
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</taxonomicName>
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approx. 3.0) and carapace shorter (3.6 long, 4.61 wide;
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<taxonomicName genus="M." lsidName="M. occataria" pageId="0" pageNumber="509" rank="species" species="occataria">
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<emphasis italics="true" pageId="0" pageNumber="509">M. occataria</emphasis>
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</taxonomicName>
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approx. 5.0 long, 7.0 wide). More cuspules on the labium and maxillae than in
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<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
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<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
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</taxonomicName>
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but less than in
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<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
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</taxonomicName>
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sp. nov. (
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<taxonomicName genus="M." lsidName="M. insignis" pageId="0" pageNumber="509" rank="species" species="insignis">
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<emphasis italics="true" pageId="0" pageNumber="509">M. insignis</emphasis>
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</taxonomicName>
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: none;
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<taxonomicName genus="M." lsidName="M. manningensis" pageId="0" pageNumber="509" rank="species" species="manningensis">
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<emphasis italics="true" pageId="0" pageNumber="509">M. manningensis</emphasis>
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</taxonomicName>
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sp. nov.: 5 at labium, 30 at maxillae;
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<taxonomicName genus="M." lsidName="M. davidi" pageId="0" pageNumber="509" rank="species" species="davidi">
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<emphasis italics="true" pageId="0" pageNumber="509">M. davidi</emphasis>
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</taxonomicName>
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sp. nov.: 15-10 at labium, 35-100 at maxillae). Lacks a ridged cheliceral groove which is present in
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<taxonomicName genus="M." lsidName="M. iugum" pageId="0" pageNumber="509" rank="species" species="iugum">
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<emphasis italics="true" pageId="0" pageNumber="509">M. iugum</emphasis>
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</taxonomicName>
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sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="509" type="description">
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<paragraph pageId="0" pageNumber="509">Description.</paragraph>
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<paragraph pageId="0" pageNumber="509">
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<emphasis bold="true" pageId="0" pageNumber="509">MALE</emphasis>
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(based on holotype; WAMT92071). Total length 8.95.
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<emphasis bold="true" pageId="0" pageNumber="509">Colour</emphasis>
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: pars cephalica and chelicerae orange (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11C</figureCitation>
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); a slim, black ring surrounding the PME (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11H</figureCitation>
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); pars thoracica brown with a light, purplish sheen (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11C</figureCitation>
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); abdomen greyish with a light, metallic blue sheen on the dorsal side (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11D</figureCitation>
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), ventrally more brownish with a faint hint of purple (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11E</figureCitation>
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); sternum orange, slightly fading into olive with 8 sigilla in different shades of orange (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11G</figureCitation>
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); labium and maxillae orange with a dark olive spot on the base of labium (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11F</figureCitation>
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); legs olive fading into light brown ventrally, dorsally brown (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11A, B</figureCitation>
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); spinnerets beige (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11E</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="509">Carapace</emphasis>
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: 3.6 long, 4.61 wide and 1.96 high; clypeus 0.31; pars cephalica covers 2.25 of its length, is highly elevated and slightly granulated with very few setae (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12D</figureCitation>
|
||
); pars thoracica also granulated with bands of faint, radial fissures and with two deeply expressed notches close to the abdomen (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11C</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Eyes</emphasis>
|
||
: OQ 3.4 times wider than long; outer width of each eye pair AME 0.48, ALE 2.26, PME 1.44 and PLE 2.14; diameter of AME 0.19, ALE 0.19, PME 0.13, PLE 0.17; anterior eyes in a straight line; posterior eyes strongly recurved (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11H</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Chelicerae</emphasis>
|
||
: 2.17 long and 1.41 wide on the base; edges rounded and recurved with the widest point being 1.51 close to the chelicerae base (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11C</figureCitation>
|
||
); few setae along the inner margin and slightly more evenly spread setae along the anterior part of the chelicerae; rastellum present, slightly pronounced, consisting of a sclerotized process with 5 (left 8) strong, conical spines (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11I</figureCitation>
|
||
); over 20 setae cover the anterior base of fang of each chelicera; inner margin of cheliceral furrow with 2 rows of teeth and a general cheliceral teeth area in between those 2 clear rows (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12G</figureCitation>
|
||
); prolateral row with approx. 11 teeth; retrolateral row with 4 teeth; intermediate area with 5 small teeth.
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Maxillae</emphasis>
|
||
: 2.09 long and 1.5 wide; at least 30 extremely weakly developed cuspules along anterior margin (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11F</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Labium</emphasis>
|
||
: 0.79 long and 0.86 wide on the base; conical; at least 5 extremely weak developed cuspules anteriorly (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11F</figureCitation>
|
||
); labiosternal junction visible (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11G</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Sternum</emphasis>
|
||
: 2.62 long and 2.29 wide; ovoid (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11G</figureCitation>
|
||
); setae of various length somewhat densely but irregular along the margin and a smaller amount of setae spread unevenly over the sternum; 4 pairs of sigilla, anterior pair very small and hardly visible, second pair also hardly visible, smallest of all pairs and divided into two circles, third pair significantly larger than second (roughly 4 times bigger) in the shape of an elongated oval, and posterior pair biggest (roughly 2 times the size of the third pair) and drop-shaped, all sigilla slightly depressed.
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Abdomen</emphasis>
|
||
: 3.12 long and 2.74 wide; shape of a rounded trapezoid (but collapsed through preservation; Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11D</figureCitation>
|
||
); 4 spinnerets, PLS 0.43 long (part of it broken off), 0.35 wide; PMS 0.37 long, 0.19 wide (Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A habitus, dorsal view; B same, ventral view; C carapace, dorsal view; D abdomen, dorsal view; E abdomen, ventral view; F maxillae, labium, and chelicerae, ventral view; G sternum, ventral view; H eye region, dorsal view; I rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm." figureDoi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509">11E</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Pedipalp</emphasis>
|
||
: length of trochanter 1.64, femur 4.46, patella 1.82, tibia 3.75, tarsus 0.64; all segments with setae, tibia ventrally covered with comparably long and dense setae (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12A-C</figureCitation>
|
||
); tibia rather thin and slightly recurved, 1.00 wide on the widest point from dorsal/ventral and prolateral/retrolateral view (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12A-C</figureCitation>
|
||
); bulb roughly pyriform, two strongly sclerotized sections connected by a velar median structure (
|
||
<normalizedToken originalValue="“haematodocha”">"haematodocha"</normalizedToken>
|
||
); embolus rather short and bend with an intumescence in proximal region; tip of embolus triangular with a small lamella, best visible retrolateral (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12I</figureCitation>
|
||
), and a tooth, best visible prolateral (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12H</figureCitation>
|
||
).
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Legs</emphasis>
|
||
: brown setae of various sizes on all sides of the legs and bent towards the exterior with the exception of the femur setae on ventral position which are mostly vertically; ventral preening comb on tarsi and metatarsi III and IV.
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Leg spination</emphasis>
|
||
: leg I: tibia rl0, v17, pl0, d0; metatarsus rl0, v9, pl0, d0; tarsus rl0, v154-8-3, pl0, d0; leg II: tibia rl1, v12, pl0, d0; metatarsus rl0, v8, pl0, d0; tarsus rl0, v10, pl2, d0; leg III: tibia rl3, v13, pl0, d7; metatarsus rl3, v6, pl3, d9; tarsus rl3, v11, pl4, d5; leg IV: tibia rl0, v12, pl0, d0; metatarsus rl0, v12, pl5, d1; tarsus rl8, v13, pl7, d5; patella I with 8 spines spread out prolaterally and one spine ventrally close to tibia; patella II with one spine prolaterally and one ventrally both close to tibia; patella III with 26 spines spread out dorsally and prolaterally and one spine retrolaterally (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12E</figureCitation>
|
||
); patella IV with approx. 14 spines on the dorsal side (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Missulena manningensis sp. nov. Male holotype (WAMT 92071): A right pedipalp, retrolateral view; B ventral view; C prolateral view; D carapace, lateral view; E patella III, dorsal view; F patella IV, dorsal view; G pattern of cheliceral teeth in cheliceral groove; H left pedipalp, embolus with embolar tooth, prolateral-dorsal view; I left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40 µm" figureDoi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509">12F</figureCitation>
|
||
) and approx. 15 spines on the prolateral side, all very small.
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Leg measurement</emphasis>
|
||
: Leg I: femur 4.0, patella 1.37, tibia 2.97, metatarsus 2.69, tarsus 1.56, total 12.59. Leg II: femur 3.4, patella 1.12, tibia 2.5, metatarsus 2.38, tarsus 1.47, total 10.87. Leg III: femur 2.81, patella 1.05, tibia 1.94, metatarsus 2.26, tarsus 1.54, total 9.6. Leg IV: femur 3.28, patella 1.13, tibia 2.83, metatarsus 2.41, tarsus 1.59, total 11.24. Formula: 1>4>2>3.
|
||
</paragraph>
|
||
<caption doi="10.3897/asp.79.e62332.figure11" httpUri="https://binary.pensoft.net/fig/597117" pageId="0" pageNumber="509" start="Figure 11" startId="F11">
|
||
<paragraph pageId="0" pageNumber="509">
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Figure 11.</emphasis>
|
||
<taxonomicName authorityName="Greenberg & Huey & Framenau & Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena manningensis" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="manningensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="509">Missulena manningensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Male holotype (WAMT92071):
|
||
<emphasis bold="true" pageId="0" pageNumber="509">A</emphasis>
|
||
habitus, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">B</emphasis>
|
||
same, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">C</emphasis>
|
||
carapace, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">D</emphasis>
|
||
abdomen, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">E</emphasis>
|
||
abdomen, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">F</emphasis>
|
||
maxillae, labium, and chelicerae, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">G</emphasis>
|
||
sternum, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">H</emphasis>
|
||
eye region, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">I</emphasis>
|
||
rastellum, anterior view. Scale bars: A, B 5.0 mm; C-I 2.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/asp.79.e62332.figure12" httpUri="https://binary.pensoft.net/fig/597118" pageId="0" pageNumber="509" start="Figure 12" startId="F12">
|
||
<paragraph pageId="0" pageNumber="509">
|
||
<emphasis bold="true" pageId="0" pageNumber="509">Figure 12.</emphasis>
|
||
<taxonomicName authorityName="Greenberg & Huey & Framenau & Harms" authorityYear="2021" class="Arachnida" family="Actinopodidae" genus="Missulena" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Missulena manningensis" order="Araneae" pageId="0" pageNumber="509" phylum="Arthropoda" rank="species" species="manningensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="509">Missulena manningensis</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. Male holotype (WAMT92071):
|
||
<emphasis bold="true" pageId="0" pageNumber="509">A</emphasis>
|
||
right pedipalp, retrolateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">B</emphasis>
|
||
ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">C</emphasis>
|
||
prolateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">D</emphasis>
|
||
carapace, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">E</emphasis>
|
||
patella III, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">F</emphasis>
|
||
patella IV, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">G</emphasis>
|
||
pattern of cheliceral teeth in cheliceral groove;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">H</emphasis>
|
||
left pedipalp, embolus with embolar tooth, prolateral-dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="509">I</emphasis>
|
||
left pedipalp, embolus with lamella, retrolateral-ventral view. Scale bars: A-D 2.0 mm; E-G 1.0 mm; H, I 40
|
||
<normalizedToken originalValue="µm">µm</normalizedToken>
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="509" type="etymology">
|
||
<paragraph pageId="0" pageNumber="509">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="509">The specific epithet refers to the type locality, Mt Manning, in the Goldfields region of Western Australia.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="509" type="distribution">
|
||
<paragraph pageId="0" pageNumber="509">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="509">
|
||
Known only from the Mt Manning area approximately 47 km northwest of Boorabbin in the Goldfields region of Western Australia (Fig.
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Distribution records of Missulena manningensis sp. nov. and Missulena iugum sp. nov. in Western Australia." figureDoi="10.3897/asp.79.e62332.figure10" httpUri="https://binary.pensoft.net/fig/597116" pageId="0" pageNumber="509">10</figureCitation>
|
||
). The habitat of the holotype comprises open tall eucalypt woodland with mixed shrubs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</subSection>
|
||
</document> |