283 lines
37 KiB
XML
283 lines
37 KiB
XML
<document id="D0C872D3C02E0CBED6FE18CDA9979489" ID-DOI="10.5281/zenodo.174098" ID-GBIF-Dataset="3bf47564-67d5-43f5-bdf3-1872a93fa8dd" ID-ISSN="11755326" ID-Zenodo-Dep="174098" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459790480129" checkinUser="plazi" docAuthor="Hoggett, Anne K." docDate="2006" docId="4909A10BFFCC5968FEDCF9AF478CB06C" docLanguage="en" docName="zt01326p024.pdf" docOrigin="Zootaxa 1326" docStyle="DocumentStyle:FA7E419B012A62B0FC3AC15A186C3DAF.3:Zootaxa.2001-2006.journal_article" docStyleId="FA7E419B012A62B0FC3AC15A186C3DAF" docStyleName="Zootaxa.2001-2006.journal_article" docStyleVersion="3" docTitle="Macrophiothrix caenosa Hoggett, 2006, n. sp." docType="treatment" docVersion="9" lastPageNumber="22" masterDocId="B530D973FFCD596DFFD4FFD64620B56C" masterDocTitle="A new species of Macrophiothrix (Ophiuroidea: Ophiotrichidae) common in northern Australia" masterLastPageNumber="24" masterPageNumber="17" pageNumber="18" updateTime="1698219165799" updateUser="plazi">
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<mods:mods id="E613823638B5DA668BEEB3051AA3D4D7" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="661BC14D08595F6B92FA890C6285DDB9">
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<mods:title id="61AE72C70544511BEF6E6333A0FBAAB3">A new species of Macrophiothrix (Ophiuroidea: Ophiotrichidae) common in northern Australia</mods:title>
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</mods:titleInfo>
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<mods:name id="9519C8894081CE2FDC3D540E09F90FF4" type="personal">
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<mods:roleTerm id="0F5EAC328E74EABB3061E3EC69D2DABF">Author</mods:roleTerm>
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<mods:namePart id="10A21ECD2CC090AD070EAC82D8F2A422">Hoggett, Anne K.</mods:namePart>
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<mods:typeOfResource id="3BF42BE339279D1E6BEF65DA90088FC8">text</mods:typeOfResource>
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<mods:title id="84750A9F92B468DA6AC517D23719A3AA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="87DBC73EA1A430CFB2174FE0DF4B105C">
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<mods:date id="D2FA1FDD07F8D70A246C672B15F63C71">2006</mods:date>
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<mods:detail id="8C4BF7AC3C9552016BE4056E31C7B727" type="volume">
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<mods:number id="B521A4976E208C8CBB10A2B19406CE1F">1326</mods:number>
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<mods:extent id="2856968365C4451F49889141D13DFDC2" unit="page">
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<mods:start id="7C2D950933014A79FAE9F4819034F6E5">17</mods:start>
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<mods:end id="652DA579E2FB98EB91B9C17EDE5B8CEE">24</mods:end>
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<mods:classification id="59235334A8F25D9EC4F2D28CCFE445F9">journal article</mods:classification>
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<mods:identifier id="3A8F84BB28C36F6BEC264B43011C34C7" type="DOI">10.5281/zenodo.174098</mods:identifier>
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<mods:identifier id="C62E3FF607A75F36BB7E06A8F5CA3A6B" type="GBIF-Dataset">3bf47564-67d5-43f5-bdf3-1872a93fa8dd</mods:identifier>
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<mods:identifier id="556BC08D8294DBECB6018D5E629ECC08" type="ISSN">11755326</mods:identifier>
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<mods:identifier id="49D20B698CF1C87A91F01E2423FCDF17" type="Zenodo-Dep">174098</mods:identifier>
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<treatment id="4909A10BFFCC5968FEDCF9AF478CB06C" ID-DOI="http://doi.org/10.5281/zenodo.6262639" ID-GBIF-Taxon="119350299" ID-Zenodo-Dep="6262639" LSID="urn:lsid:plazi:treatment:4909A10BFFCC5968FEDCF9AF478CB06C" httpUri="http://treatment.plazi.org/id/4909A10BFFCC5968FEDCF9AF478CB06C" lastPageId="5" lastPageNumber="22" pageId="1" pageNumber="18">
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<subSubSection id="89BA4396FFCC596CFEDCF9AF47B0B3DA" pageId="1" pageNumber="18" type="nomenclature">
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<paragraph id="C11F101DFFCC596CFEDCF9AF445BB3FF" blockId="1.[264,635,1657,1718]" box="[264,635,1657,1683]" pageId="1" pageNumber="18">
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<heading id="9A57A771FFCC596CFEDCF9AF445BB3FF" bold="true" box="[264,635,1657,1683]" fontSize="11" level="1" pageId="1" pageNumber="18" reason="1">
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<emphasis id="F3D4CC0FFFCC596CFEDCF9AF445BB3FF" bold="true" box="[264,635,1657,1683]" pageId="1" pageNumber="18">
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<taxonomicName id="06A06B9EFFCC596CFEDCF9AF4411B3FF" box="[264,561,1657,1683]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="1" pageNumber="18" phylum="Echinodermata" rank="species" species="caenosa" status="sp. nov.">
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<emphasis id="F3D4CC0FFFCC596CFEDCF9AF4411B3FF" bold="true" box="[264,561,1657,1683]" italics="true" pageId="1" pageNumber="18">Macrophiothrix caenosa</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="E8E77174FFCC596CFDEDF9AF445BB3FF" box="[569,635,1657,1683]" pageId="1" pageNumber="18" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph id="C11F101DFFCC596CFEDCF94A47B0B3DA" blockId="1.[264,635,1657,1718]" box="[264,400,1692,1718]" pageId="1" pageNumber="18">
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(
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<figureCitation id="599B0C98FFCC596CFEC4F94A47A7B3DA" box="[272,391,1692,1718]" captionStart-0="FIGURES 1, 2" captionStart-1="FIGURES 3 – 9" captionStart-2="FIGURES 10 – 12" captionStartId-0="3.[264,383,1035,1059]" captionStartId-1="4.[264,382,1596,1620]" captionStartId-2="5.[264,382,1019,1043]" captionTargetBox-0="[264,1323,566,1011]" captionTargetBox-1="[264,1322,310,1572]" captionTargetBox-2="[299,1279,328,1018]" captionTargetId-0="figure@3.[264,1323,566,1011]" captionTargetId-1="figure@4.[264,1323,310,1573]" captionTargetId-2="figure@5.[264,1323,298,1031]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURES 1, 2. Macrophiothrix caenosa n. sp., holotype, disc diameter 19 mm. 1. Dorsal view. 2. Ventral view." captionText-1="FIGURES 3 – 9. Dental plates of Macrophiothrix caenosa and M. longipeda, ventral ends uppermost. 3 – 6. M. caenosa. 3. Specimen from Darwin, Northern Territory (NTM Q 4928, d. d. 16 mm), papillae and teeth removed. 4. Same specimen (NTM Q 4928), papillae and teeth intact. 5. Specimen from Japan (NTM Q 5423, d. d. 17 mm), papillae and teeth intact. 6. Specimen from Darwin, Northern Territory (NTM Q 4849, d. d. 11 mm), papillae and teeth intact. 7 – 9. M. longipeda. 7. Specimen from Lizard Island, Queensland (NTM Q 5421, d. d. 16 mm), papillae and teeth removed. 8. Same specimen (Q 5421), papillae and teeth intact. 9. Specimen from MacGillivray’s Reef near Lizard Island, Queensland, d. d. 11 mm, papillae and two of four teeth intact (dorsalmost tooth removed, second dorsalmost broken). Scale bar 1 mm." captionText-2="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri-0="https://zenodo.org/record/174099/files/figure.png" httpUri-1="https://zenodo.org/record/174100/files/figure.png" httpUri-2="https://zenodo.org/record/174101/files/figure.png" pageId="1" pageNumber="18">Figs 1–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="89BA4396FFCC596CFEDCF93A432AB24F" pageId="1" pageNumber="18" type="reference_group">
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<paragraph id="C11F101DFFCC596CFEDCF93A42A8B26F" blockId="1.[264,1290,1772,1827]" box="[264,1160,1772,1795]" pageId="1" pageNumber="18">
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<treatmentCitationGroup id="E1B03733FFCC596CFEDCF93A42A8B26F" box="[264,1160,1772,1795]" pageId="1" pageNumber="18">
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<taxonomicName id="06A06B9EFFCC596CFEDCF93A44CEB26F" authority="Hoggett, 1990: 103" authorityName="Hoggett" authorityPageNumber="103" authorityYear="1990" box="[264,750,1772,1795]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="1" pageNumber="18" phylum="Echinodermata" rank="species" species="caenosa">
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<emphasis id="F3D4CC0FFFCC596CFEDCF93A442DB26F" box="[264,525,1772,1795]" italics="true" pageId="1" pageNumber="18">Macrophiothrix caenosa</emphasis>
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:
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<treatmentCitation id="4001360CFFCC596CFDC8F93A44CEB26F" author="Hoggett" box="[540,750,1772,1795]" page="103" pageId="1" pageNumber="18" year="1990">
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<bibRefCitation id="A5316DECFFCC596CFDC8F93A4498B26F" author="Hoggett" box="[540,696,1772,1795]" pageId="1" pageNumber="24" refString="Hoggett, A. K. (1990) Taxonomy and systematic position of the brittlestar genus Macrophiothrix H. L. Clark (Echinodermata: Ophiuroidea). PhD thesis, University of Queensland, Australia." type="book" year="1990">Hoggett, 1990</bibRefCitation>
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: 103
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</treatmentCitation>
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</taxonomicName>
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;
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<bibRefCitation id="A5316DECFFCC596CFD2DF93A45A0B26F" author="Miller" box="[761,896,1772,1795]" pageId="1" pageNumber="24" refString="Miller, R. L. (1997) Specificity of sperm chemotaxis among Great Barrier Reef shallow-water holothurians and ophiuroids. Journal of Experimental Zoology, 279, 189 - 200." type="journal article" year="1997">Miller, 1997</bibRefCitation>
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;
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<bibRefCitation id="A5316DECFFCC596CFC5FF93A42A3B26F" author="Hart" box="[907,1155,1772,1795]" pageId="1" pageNumber="24" refString="Hart, M. W. & R. D. Podolsky (2005) Mitochondrial DNA phylogeny and rates of larval evolution in Macrophiothrix brittlestars. Molecular Phylogenetics and Evolution, 34, 438 - 447." type="journal article" year="2005">Hart & Podolsky, 2005</bibRefCitation>
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||
.
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</treatmentCitationGroup>
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||
</paragraph>
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<paragraph id="C11F101DFFCC596CFEDCF8DA432AB24F" blockId="1.[264,1290,1772,1827]" box="[264,1290,1804,1827]" pageId="1" pageNumber="18">
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<treatmentCitationGroup id="E1B03733FFCC596CFEDCF8DA432AB24F" box="[264,1290,1804,1827]" pageId="1" pageNumber="18">
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||
<taxonomicName id="06A06B9EFFCC596CFEDCF8DA452AB24F" authority="Hoggett, 1991: 1103" authorityName="Hoggett" authorityPageNumber="1103" authorityYear="1991" box="[264,778,1804,1827]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="1" pageNumber="18" phylum="Echinodermata" rank="species" species="longipeda">
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<emphasis id="F3D4CC0FFFCC596CFEDCF8DA4401B24F" box="[264,545,1804,1827]" italics="true" pageId="1" pageNumber="18">Macrophiothrix longipeda</emphasis>
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:
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<treatmentCitation id="4001360CFFCC596CFDFBF8DA452AB24F" author="Hoggett" box="[559,778,1804,1827]" page="1103" pageId="1" pageNumber="18" year="1991">
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||
<bibRefCitation id="A5316DECFFCC596CFDFBF8DA44ECB24F" author="Hoggett" box="[559,716,1804,1827]" pageId="1" pageNumber="24" refString="Hoggett, A. K. (1991) The genus Macrophiothrix (Ophiuroidea: Ophiotrichidae) in Australian waters. Invertebrate Taxonomy, 4, 1077 - 146." type="journal article" year="1991">Hoggett, 1991</bibRefCitation>
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: 1103
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</treatmentCitation>
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</taxonomicName>
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(part), fig. 14B (not figs. 14A, 14C–F, 15A–C).
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="89BA4396FFCC596FFEDCF88247ABB6B2" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18" type="materials_examined">
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<paragraph id="C11F101DFFCC596FFEDCF882459DB4EA" blockId="1.[264,1322,1876,1902]" lastBlockId="2.[264,1325,284,1870]" lastPageId="2" lastPageNumber="19" pageId="1" pageNumber="18">
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||
<emphasis id="F3D4CC0FFFCC596CFEDCF88247E8B202" bold="true" box="[264,456,1876,1902]" pageId="1" pageNumber="18">
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||
<typeStatus id="1E1BAEBFFFCC596CFEDCF8824765B202" box="[264,325,1876,1902]" pageId="1" pageNumber="18">Type</typeStatus>
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material:
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</emphasis>
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<typeStatus id="1E1BAEBFFFCC596CFE03F8824467B202" box="[471,583,1876,1902]" pageId="1" pageNumber="18" type="holotype">Holotype</typeStatus>
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,
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<collectionCode id="A7B188D8FFCC596CFD8FF88244BCB202" box="[603,668,1876,1902]" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="1" pageNumber="18">NTM</collectionCode>
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Q4925, 8
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<typeStatus id="1E1BAEBFFFCC596CFCE4F8824583B202" box="[816,931,1876,1902]" pageId="1" pageNumber="18" type="paratype">paratypes</typeStatus>
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,
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<collectionCode id="A7B188D8FFCC596CFC62F88245D7B202" box="[950,1015,1876,1902]" httpUri="http://grbio.org/cool/jg13-7fbc" name="Northern Territory Museum of Arts and Sciences" pageId="1" pageNumber="18">NTM</collectionCode>
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Q4926, Q4927, Q4928, Q4929, Q4930, Q4931, Q4932, Q4934. All
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<typeStatus id="1E1BAEBFFFCF596FFCE3FECA4549B45A" box="[823,873,284,310]" pageId="2" pageNumber="19">type</typeStatus>
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specimens from East Arm, Darwin Harbour, Northern Territory,
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<collectingCountry id="B9B7508DFFCF596FFDB6FE9244F0B432" box="[610,720,324,350]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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, muddy sand and rocky reef, low intertidal, under rock, collected by A. Hoggett and C. Johnson,
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<date id="B51E36DDFFCF596FFCF8FEBA4599B4EA" box="[812,953,364,390]" pageId="2" pageNumber="19" value="1986-10-09">9 Oct. 1986</date>
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.
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</paragraph>
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<paragraph id="C11F101DFFCF596FFEECFE4547ABB6B2" blockId="2.[264,1325,284,1870]" pageId="2" pageNumber="19">
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<emphasis id="F3D4CC0FFFCF596FFEECFE454420B4C1" bold="true" box="[312,512,403,429]" pageId="2" pageNumber="19">Other material:</emphasis>
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<specimenCount id="D7A6DB94FFCF596FFDD8FE424491B4C2" box="[524,689,404,430]" pageId="2" pageNumber="19" type="generic">79 specimens</specimenCount>
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.
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<emphasis id="F3D4CC0FFFCF596FFD6BFE424589B4C1" box="[703,937,404,429]" italics="true" pageId="2" pageNumber="19">Australian Museum</emphasis>
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: Capricorn Group, Queensland,
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<collectingCountry id="B9B7508DFFCF596FFEDCFE6A4753B4BA" box="[264,371,444,470]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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— J2340 (1), J7046 (1).
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<emphasis id="F3D4CC0FFFCF596FFD65FE6A45DDB4B9" box="[689,1021,444,469]" italics="true" pageId="2" pageNumber="19">Northern Territory Museum</emphasis>
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: Darwin, Port Essington and Orontes Reef, Northern Territory,
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<collectingCountry id="B9B7508DFFCF596FFD37FE324572B492" box="[739,850,484,510]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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; Lizard Island, Queensland,
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<collectingCountry id="B9B7508DFFCF596FFB61FE324304B492" box="[1205,1316,484,510]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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; Broome, Western
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<collectingCountry id="B9B7508DFFCF596FFE3CFDDA4477B74A" box="[488,599,524,550]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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;
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<collectingCountry id="B9B7508DFFCF596FFDBDFDDA448FB74A" box="[617,687,524,550]" name="Japan" pageId="2" pageNumber="19">Japan</collectingCountry>
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; intertidal to 18 metres — Q3853 (6), Q3872 (1), Q3877 (1), Q3899 (1), Q3902 (1),Q3903 (1),Q3904 (1), Q3907 (1), Q4531 (1), Q4540 (2), Q4541 (1), Q4706 (1), Q4735 (1), Q4740 (1), Q4769 (1), Q4770 (1), Q4771 (1), Q4779 (1), Q4825 (1), Q4826 (1), Q4827 (1), Q4828 (1),Q4830 (1), Q4831 (1), Q4834 (1), Q4848 (1), Q4849 (1), Q4850 (1), Q4851 (1), Q4857 (1), Q4882 (1), Q4884 (2), Q4889 (1), Q4890 (1), Q4897 (1), Q4912 (1), Q4960 (1), Q4961 (1), Q4962 (1),Q5260 (8), Q5423 (1), Q5477 (1), Q5669 (1), Q6171 (1), Q6172 (1).
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<emphasis id="F3D4CC0FFFCF596FFC05FD2A4302B679" box="[977,1314,764,789]" italics="true" pageId="2" pageNumber="19">Western Australian Museum</emphasis>
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: Walsh Point, Point Cloates, North West Cape, Coral Bay, Abrolhos Islands, Warroora, and off Port Hedland, Western
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<collectingCountry id="B9B7508DFFCF596FFD93FC9A4496B60A" box="[583,694,844,870]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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; Thursday Island, Queensland,
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<collectingCountry id="B9B7508DFFCF596FFBF9FC9A42BCB60A" box="[1069,1180,844,870]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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;
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<collectingCountry id="B9B7508DFFCF596FFB7EFC9A4304B60A" box="[1194,1316,844,870]" name="Singapore" pageId="2" pageNumber="19">Singapore</collectingCountry>
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; intertidal to 30 metres — 94976 (1), 8777 (1), 32377 (1), 44777 (1), 36378 (1), 65678 (2), 81078 (2), 86381 (1), 86781 (1), 87181 (arms), 87781 (2), 19684 (1), 37787 (1), 37887 (2).
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</paragraph>
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</subSubSection>
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<subSubSection id="89BA4396FFCF596FFEECFC3A4263B112" pageId="2" pageNumber="19" type="distribution">
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<paragraph id="C11F101DFFCF596FFEECFC3A420BB142" blockId="2.[264,1325,284,1870]" pageId="2" pageNumber="19">
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<emphasis id="F3D4CC0FFFCF596FFEECFC3A47FAB16A" bold="true" box="[312,474,1004,1030]" pageId="2" pageNumber="19">Distribution:</emphasis>
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Northern
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<collectingCountry id="B9B7508DFFCF596FFD86FC3A44E0B16A" box="[594,704,1004,1030]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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, from the Abrolhos Islands, Western
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<collectingCountry id="B9B7508DFFCF596FFBA4FC3A42FFB16A" box="[1136,1247,1004,1030]" name="Australia" pageId="2" pageNumber="19">Australia</collectingCountry>
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||
, to the Capricorn Islands, Queensland;
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<collectingCountry id="B9B7508DFFCF596FFD54FBC244DBB142" box="[640,763,1044,1070]" name="Singapore" pageId="2" pageNumber="19">Singapore</collectingCountry>
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||
;
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<collectingCountry id="B9B7508DFFCF596FFCD2FBC2456CB142" box="[774,844,1044,1070]" name="Japan" pageId="2" pageNumber="19">Japan</collectingCountry>
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||
. Intertidal to
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<specimenCount id="D7A6DB94FFCF596FFC3CFBC2420BB142" box="[1000,1067,1044,1070]" pageId="2" pageNumber="19" type="generic">
|
||
<quantity id="0658BDF8FFCF596FFC3CFBC24206B142" box="[1000,1062,1044,1070]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="2" pageNumber="19" unit="m" value="30.0">30 m</quantity>
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||
.
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</specimenCount>
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</paragraph>
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||
<paragraph id="C11F101DFFCF596FFEECFBEA4263B112" blockId="2.[264,1325,284,1870]" pageId="2" pageNumber="19">
|
||
<emphasis id="F3D4CC0FFFCF596FFEECFBEA4781B13A" bold="true" box="[312,417,1084,1110]" pageId="2" pageNumber="19">Habitat:</emphasis>
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Under slabs of rock or coral rubble that overlie sandy or muddy substrates. The animals move swiftly into deeper substrate when the slab is lifted.
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</paragraph>
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</subSubSection>
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<subSubSection id="89BA4396FFCF596FFEECFB5A44A2B1A2" pageId="2" pageNumber="19" type="etymology">
|
||
<paragraph id="C11F101DFFCF596FFEECFB5A44A2B1A2" blockId="2.[264,1325,284,1870]" pageId="2" pageNumber="19">
|
||
<emphasis id="F3D4CC0FFFCF596FFEECFB5A47E9B1CA" bold="true" box="[312,457,1164,1190]" pageId="2" pageNumber="19">Etymology:</emphasis>
|
||
From the Latin
|
||
<emphasis id="F3D4CC0FFFCF596FFD53FB5A44D3B1C9" box="[647,755,1164,1189]" italics="true" pageId="2" pageNumber="19">caenosus</emphasis>
|
||
(= muddy), referring to the habitat in which this species lives at the
|
||
<typeStatus id="1E1BAEBFFFCF596FFE3EFB62443DB1A2" box="[490,541,1204,1230]" pageId="2" pageNumber="19">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89BA4396FFCF596FFEECFB0A440CB392" pageId="2" pageNumber="19" type="diagnosis">
|
||
<paragraph id="C11F101DFFCF596FFEECFB0A440CB392" blockId="2.[264,1325,284,1870]" pageId="2" pageNumber="19">
|
||
<emphasis id="F3D4CC0FFFCF596FFEECFB0A479AB19A" bold="true" box="[312,442,1244,1270]" pageId="2" pageNumber="19">Diagnosis:</emphasis>
|
||
Based on all material examined. A species of
|
||
<taxonomicName id="06A06B9EFFCF596FFC2DFB0A4292B199" box="[1017,1202,1244,1269]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="2" pageNumber="19" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="F3D4CC0FFFCF596FFC2DFB0A4292B199" box="[1017,1202,1244,1269]" italics="true" pageId="2" pageNumber="19">Macrophiothrix</emphasis>
|
||
</taxonomicName>
|
||
with disc diameter up to
|
||
<quantity id="0658BDF8FFCF596FFE17FAD2443CB072" box="[451,540,1284,1310]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="2" pageNumber="19" unit="mm" value="24.0">24 mm</quantity>
|
||
; arms 9 to 20 times disc diameter. Wide trapeziform dorsal arm plates (2.0 to 2.6 times wider than long;
|
||
<figureCitation id="599B0C98FFCF596FFD31FAFA457AB02A" box="[741,858,1324,1350]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="2" pageNumber="19">Figure 10</figureCitation>
|
||
). Multifid disc stumps (1.5 to 2.5 times higher than wide). Radial shields bearing short multifid stumps or rugose granules. Seven to 10 dorsoventrally flattened arm spines, the upper ones with thorns along whole length of both sides, terminating in a truncate or slightly expanded, but not clavate, tip. Longest arm spine second or third from uppermost, 0.9–1.3 times corresponding dorsal arm plate width. Ventral disc armament sparse or absent near genital slits. Genital plates usually lack armament. Oral shields without spinelets (
|
||
<figureCitation id="599B0C98FFCF596FFCDEF9CA45A4B35A" box="[778,900,1564,1590]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="2" pageNumber="19">Figure 12</figureCitation>
|
||
). Dental plate with distinct central constriction, adoral part broader than aboral part (
|
||
<figureCitation id="599B0C98FFCF596FFCB2F99245DBB332" box="[870,1019,1604,1630]" captionStart="FIGURES 3 – 9" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[264,1322,310,1572]" captionTargetId="figure@4.[264,1323,310,1573]" captionTargetPageId="4" captionText="FIGURES 3 – 9. Dental plates of Macrophiothrix caenosa and M. longipeda, ventral ends uppermost. 3 – 6. M. caenosa. 3. Specimen from Darwin, Northern Territory (NTM Q 4928, d. d. 16 mm), papillae and teeth removed. 4. Same specimen (NTM Q 4928), papillae and teeth intact. 5. Specimen from Japan (NTM Q 5423, d. d. 17 mm), papillae and teeth intact. 6. Specimen from Darwin, Northern Territory (NTM Q 4849, d. d. 11 mm), papillae and teeth intact. 7 – 9. M. longipeda. 7. Specimen from Lizard Island, Queensland (NTM Q 5421, d. d. 16 mm), papillae and teeth removed. 8. Same specimen (Q 5421), papillae and teeth intact. 9. Specimen from MacGillivray’s Reef near Lizard Island, Queensland, d. d. 11 mm, papillae and two of four teeth intact (dorsalmost tooth removed, second dorsalmost broken). Scale bar 1 mm." httpUri="https://zenodo.org/record/174100/files/figure.png" pageId="2" pageNumber="19">Figures 3–6</figureCitation>
|
||
). Lateral edges of adoral part of dental plate with a single row of conspicuously taller and more pointed dental papillae, enclosing a group of smaller rounded papillae (
|
||
<figureCitation id="599B0C98FFCF596FFC6FF942426EB3C2" box="[955,1102,1684,1710]" captionStart="FIGURES 3 – 9" captionStartId="4.[264,382,1596,1620]" captionTargetBox="[264,1322,310,1572]" captionTargetId="figure@4.[264,1323,310,1573]" captionTargetPageId="4" captionText="FIGURES 3 – 9. Dental plates of Macrophiothrix caenosa and M. longipeda, ventral ends uppermost. 3 – 6. M. caenosa. 3. Specimen from Darwin, Northern Territory (NTM Q 4928, d. d. 16 mm), papillae and teeth removed. 4. Same specimen (NTM Q 4928), papillae and teeth intact. 5. Specimen from Japan (NTM Q 5423, d. d. 17 mm), papillae and teeth intact. 6. Specimen from Darwin, Northern Territory (NTM Q 4849, d. d. 11 mm), papillae and teeth intact. 7 – 9. M. longipeda. 7. Specimen from Lizard Island, Queensland (NTM Q 5421, d. d. 16 mm), papillae and teeth removed. 8. Same specimen (Q 5421), papillae and teeth intact. 9. Specimen from MacGillivray’s Reef near Lizard Island, Queensland, d. d. 11 mm, papillae and two of four teeth intact (dorsalmost tooth removed, second dorsalmost broken). Scale bar 1 mm." httpUri="https://zenodo.org/record/174100/files/figure.png" pageId="2" pageNumber="19">Figures 4–6</figureCitation>
|
||
,
|
||
<figureCitation id="599B0C98FFCF596FFB8BF94242A0B3C2" box="[1119,1152,1684,1710]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="2" pageNumber="19">12</figureCitation>
|
||
). Ripe female gonads golden yellow and ripe male gonads pale orange in living specimens, both fading to white on preservation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="89BA4396FFCF5968FEECF8DA478CB06C" lastPageId="5" lastPageNumber="22" pageId="2" pageNumber="19" type="description">
|
||
<paragraph id="C11F101DFFCF596EFEECF8DA45BAB492" blockId="2.[264,1325,284,1870]" lastBlockId="3.[264,1325,284,510]" lastPageId="3" lastPageNumber="20" pageId="2" pageNumber="19">
|
||
<emphasis id="F3D4CC0FFFCF596FFEECF8DA4449B24A" bold="true" box="[312,617,1804,1830]" pageId="2" pageNumber="19">
|
||
Description of
|
||
<typeStatus id="1E1BAEBFFFCF596FFE20F8DA4444B24A" box="[500,612,1804,1830]" pageId="2" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="1E1BAEBFFFCF596FFD7EF8DA4531B24A" box="[682,785,1804,1830]" pageId="2" pageNumber="19" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="599B0C98FFCF596FFCF6F8DA4592B24A" box="[802,946,1804,1830]" captionStart="FIGURES 1, 2" captionStartId="3.[264,383,1035,1059]" captionTargetBox="[264,1323,566,1011]" captionTargetId="figure@3.[264,1323,566,1011]" captionTargetPageId="3" captionText="FIGURES 1, 2. Macrophiothrix caenosa n. sp., holotype, disc diameter 19 mm. 1. Dorsal view. 2. Ventral view." httpUri="https://zenodo.org/record/174099/files/figure.png" pageId="2" pageNumber="19">Figures 1–2</figureCitation>
|
||
) is a female. Maximum oocyte diameter is
|
||
<quantity id="0658BDF8FFCF596FFE44F8E2442BB222" box="[400,523,1844,1870]" metricMagnitude="-4" metricUnit="m" metricValue="2.04" pageId="2" pageNumber="19" unit="mm" value="0.204">0.204 mm</quantity>
|
||
. Disc diameter is
|
||
<quantity id="0658BDF8FFCF596FFD0DF8E2450EB222" box="[729,814,1844,1870]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="2" pageNumber="19" unit="mm" value="19.0">19 mm</quantity>
|
||
, arm length
|
||
<quantity id="0658BDF8FFCF596FFC6AF8E24203B222" box="[958,1059,1844,1870]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="2" pageNumber="19" unit="mm" value="290.0">290 mm</quantity>
|
||
. Dorsal disc armament is of short (about twice as high as wide), cylindrical, multifid stumps, density 18 per square mm. Ventrally, disc stumps become shorter and have fewer points as they approach the genital plates and jaw. Stumps do not extend onto the long genital plates that border the genital slits, or onto the winglike genital plates distal to the oral shields (
|
||
<figureCitation id="599B0C98FFCE596EFB75FE42433DB4C2" box="[1185,1309,404,430]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="3" pageNumber="20">Figure 12</figureCitation>
|
||
). Radial shields are 6.0 mm long,
|
||
<quantity id="0658BDF8FFCE596EFD49FE6A44DBB4BA" box="[669,763,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide, widest
|
||
<quantity id="0658BDF8FFCE596EFC7EFE6A4228B4BA" box="[938,1032,444,470]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="20" unit="mm" value="2.5">2.5 mm</quantity>
|
||
from distal end. Radial shields bear rugose granules, density 25 per square mm.
|
||
</paragraph>
|
||
<caption id="95DF4095FFCE596EFEDCFBDD47B5B12A" httpUri="https://zenodo.org/record/174099/files/figure.png" pageId="3" pageNumber="20" targetBox="[264,1323,566,1011]" targetPageId="3">
|
||
<paragraph id="C11F101DFFCE596EFEDCFBDD47B5B12A" blockId="3.[264,1323,1035,1094]" pageId="3" pageNumber="20">
|
||
<emphasis id="F3D4CC0FFFCE596EFEDCFBDD478EB14F" bold="true" box="[264,430,1035,1059]" pageId="3" pageNumber="20">FIGURES 1, 2</emphasis>
|
||
.
|
||
<taxonomicName id="06A06B9EFFCE596EFE68FBDA44E2B14F" box="[444,706,1036,1059]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="3" pageNumber="20" phylum="Echinodermata" rank="species" species="caenosa" status="sp. nov.">
|
||
<emphasis id="F3D4CC0FFFCE596EFE68FBDA44E2B14F" box="[444,706,1036,1059]" italics="true" pageId="3" pageNumber="20">Macrophiothrix caenosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="E8E77174FFCE596EFD1EFBDA4525B14F" box="[714,773,1036,1059]" pageId="3" pageNumber="20" rank="species">n. sp.</taxonomicNameLabel>
|
||
, holotype, disc diameter 19 mm. 1. Dorsal view. 2. Ventral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C11F101DFFCE596EFEECFBAC444AB060" blockId="3.[264,1325,1146,1852]" pageId="3" pageNumber="20">
|
||
Dorsal arm plates are trapeziform (
|
||
<figureCitation id="599B0C98FFCE596EFD36FBAC457BB1F8" box="[738,859,1146,1172]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="3" pageNumber="20">Figure 10</figureCitation>
|
||
), some fractured near the longitudinal midline. The proximal one or two plates bear a few stumps similar to, but smaller than, those on the disc. All other dorsal arm plates lack stumps. The 17th dorsal arm plate is
|
||
<quantity id="0658BDF8FFCE596EFEDCFB244751B060" box="[264,369,1266,1292]" metricMagnitude="-3" metricUnit="m" metricValue="2.12" pageId="3" pageNumber="20" unit="mm" value="2.12">2.12 mm</quantity>
|
||
wide,
|
||
<quantity id="0658BDF8FFCE596EFE6BFB244408B060" box="[447,552,1266,1292]" metricMagnitude="-4" metricUnit="m" metricValue="8.4" pageId="3" pageNumber="20" unit="mm" value="0.84">0.84 mm</quantity>
|
||
long.
|
||
</paragraph>
|
||
<paragraph id="C11F101DFFCE596EFEECFACC47C4B318" blockId="3.[264,1325,1146,1852]" pageId="3" pageNumber="20">
|
||
Arm spines are dorsoventrally flattened with small thorns along the lateral edges of the uppermost few spines. Lower spines are smooth proximally with thorns only along the distal edges. The spines have parallel edges and terminate in an abruptly rounded tip, which has slightly longer thorns on the aboral side; they are not clavate. First and second free segments each have 9 spines, 3rd to 5th segments each have 8 spines, and 10th and 20th segments each have 7 spines. Longest spine in each series is the second or third from uppermost; longest spine is
|
||
<quantity id="0658BDF8FFCE596EFD8CF9DC44E4B348" box="[600,708,1546,1572]" metricMagnitude="-3" metricUnit="m" metricValue="2.33" pageId="3" pageNumber="20" unit="mm" value="2.33">2.33 mm</quantity>
|
||
on segment 17, 1.1 times the corresponding dorsal arm plate width. Ventralmost arm spine is the smallest in the series and has a few short thorns near the tip.
|
||
</paragraph>
|
||
<paragraph id="C11F101DFFCE596EFEECF95444EBB3A8" blockId="3.[264,1325,1146,1852]" pageId="3" pageNumber="20">
|
||
Ventral arm plates are wider than long, with rounded distal corners (
|
||
<figureCitation id="599B0C98FFCE596EFB75F954433DB3F0" box="[1185,1309,1666,1692]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="3" pageNumber="20">Figure 11</figureCitation>
|
||
). Tentacle scales are small and rounded.
|
||
</paragraph>
|
||
<paragraph id="C11F101DFFCE596EFEECF90442B2B250" blockId="3.[264,1325,1146,1852]" pageId="3" pageNumber="20">
|
||
Jaw structure is shown in
|
||
<figureCitation id="599B0C98FFCE596EFDB0F90444F8B380" box="[612,728,1746,1772]" captionStart="FIGURES 10 – 12" captionStartId="5.[264,382,1019,1043]" captionTargetBox="[299,1279,328,1018]" captionTargetId="figure@5.[264,1323,298,1031]" captionTargetPageId="5" captionText="FIGURES 10 – 12. Macrophiothrix caenosa, holotype. 10. Dorsal arm plates, 16 th and 17 th from disc. 11. Ventral arm plates, 19 th and 20 th from mouth. 12. Single jaw. Scale bar 1 mm." httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="3" pageNumber="20">Figure 12</figureCitation>
|
||
. All oral shields are bare. Dental papillae on lateral edges of the dental plate are conspicuously and abruptly taller than those in the centre and at the adoral end of the plate. The jaw of the
|
||
<typeStatus id="1E1BAEBFFFCE596EFCC3F8F4455EB250" box="[791,894,1826,1852]" pageId="3" pageNumber="20" type="holotype">holotype</typeStatus>
|
||
has not been dissected.
|
||
</paragraph>
|
||
<caption id="95DF4095FFC95969FEDCF9EA45ABB205" httpUri="https://zenodo.org/record/174100/files/figure.png" pageId="4" pageNumber="21" targetBox="[264,1322,310,1572]" targetPageId="4">
|
||
<paragraph id="C11F101DFFC95969FEDCF9EA45ABB205" blockId="4.[264,1323,1596,1897]" pageId="4" pageNumber="21">
|
||
<emphasis id="F3D4CC0FFFC95969FEDCF9EA479DB338" bold="true" box="[264,445,1596,1620]" pageId="4" pageNumber="21">FIGURES 3–9.</emphasis>
|
||
Dental plates of
|
||
<taxonomicName id="06A06B9EFFC95969FD4EF9EB4589B338" box="[666,937,1597,1620]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="21" phylum="Echinodermata" rank="species" species="caenosa">
|
||
<emphasis id="F3D4CC0FFFC95969FD4EF9EB4589B338" box="[666,937,1597,1620]" italics="true" pageId="4" pageNumber="21">Macrophiothrix caenosa</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="06A06B9EFFC95969FC25F9EB42A9B338" box="[1009,1161,1597,1620]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="21" phylum="Echinodermata" rank="species" species="longipeda">
|
||
<emphasis id="F3D4CC0FFFC95969FC25F9EB42A9B338" box="[1009,1161,1597,1620]" italics="true" pageId="4" pageNumber="21">M. longipeda</emphasis>
|
||
</taxonomicName>
|
||
, ventral ends uppermost. 3–6.
|
||
<taxonomicName id="06A06B9EFFC95969FE69F9B6441AB31B" box="[445,570,1632,1655]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="21" phylum="Echinodermata" rank="species" species="caenosa">
|
||
<emphasis id="F3D4CC0FFFC95969FE69F9B6441AB31B" box="[445,570,1632,1655]" italics="true" pageId="4" pageNumber="21">M. caenosa</emphasis>
|
||
</taxonomicName>
|
||
. 3. Specimen from Darwin, Northern Territory (NTM Q4928, d.d. 16 mm), papillae and teeth removed. 4. Same specimen (NTM Q4928), papillae and teeth intact. 5. Specimen from Japan (NTM Q5423, d.d. 17 mm), papillae and teeth intact. 6. Specimen from Darwin, Northern Territory (NTM Q4849, d.d.11 mm), papillae and teeth intact. 7–9.
|
||
<taxonomicName id="06A06B9EFFC95969FB42F91E4303B3B3" box="[1174,1315,1736,1759]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="4" pageNumber="21" phylum="Echinodermata" rank="species" species="longipeda">
|
||
<emphasis id="F3D4CC0FFFC95969FB42F91E4303B3B3" box="[1174,1315,1736,1759]" italics="true" pageId="4" pageNumber="21">M. longipeda</emphasis>
|
||
</taxonomicName>
|
||
. 7. Specimen from Lizard Island, Queensland (NTM Q5421, d.d. 16 mm), papillae and teeth removed. 8. Same specimen (Q5421), papillae and teeth intact. 9. Specimen from MacGillivray’s Reef near Lizard Island, Queensland, d.d. 11 mm, papillae and two of four teeth intact (dorsalmost tooth removed, second dorsalmost broken). Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="95DF4095FFC85968FEDCFC2D428EB154" httpUri="https://zenodo.org/record/174101/files/figure.png" pageId="5" pageNumber="22" targetBox="[299,1279,328,1018]" targetPageId="5">
|
||
<paragraph id="C11F101DFFC85968FEDCFC2D428EB154" blockId="5.[264,1322,1019,1080]" pageId="5" pageNumber="22">
|
||
<emphasis id="F3D4CC0FFFC85968FEDCFC2D47F0B17F" bold="true" box="[264,464,1019,1043]" pageId="5" pageNumber="22">FIGURES 10–12.</emphasis>
|
||
<taxonomicName id="06A06B9EFFC85968FE0DFC2A44C2B17F" box="[473,738,1020,1043]" class="Ophiuroidea" family="Ophiotrichidae" genus="Macrophiothrix" kingdom="Animalia" order="Ophiurida" pageId="5" pageNumber="22" phylum="Echinodermata" rank="species" species="caenosa">
|
||
<emphasis id="F3D4CC0FFFC85968FE0DFC2A44C2B17F" box="[473,738,1020,1043]" italics="true" pageId="5" pageNumber="22">Macrophiothrix caenosa</emphasis>
|
||
</taxonomicName>
|
||
, holotype. 10. Dorsal arm plates, 16th and 17th from disc. 11. Ventral arm plates, 19th and 20th from mouth. 12. Single jaw. Scale bar 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="C11F101DFFC85968FEECFBB8478CB06C" blockId="5.[264,1323,1134,1280]" pageId="5" pageNumber="22">Preserved colour is pale purple with scattered darker patches on the disc and darker transverse bands on the arms at intervals of 4–5 segments. Colour in life was shades of brown rather than purple. The gonads are white in the preserved specimen but were golden yellow in life.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |