303 lines
44 KiB
XML
303 lines
44 KiB
XML
<document id="B7E9EC0801A0A1036359C4E9FFD9762A" ID-DOI="10.1186/s13358-024-00303-y" ID-ISSN="1664-2384" ID-Zenodo-Dep="12006446" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1718655268858" checkinUser="felipe" docAuthor="Fuchs, Dirk, Weis, Robert & Thuy, Ben" docDate="2024" docId="03BAB207EC37EC3AFCC766757990816E" docLanguage="en" docName="SwissJPalaeontol.143.6.pdf" docOrigin="Swiss Journal of Palaeontology (6) 143 (1)" docSource="http://dx.doi.org/10.1186/s13358-024-00303-y" docStyle="DocumentStyle:DBDC680DC95E08776BDF6354C30A315B.1:SwissJPalaeontol.2022-.journal_article" docStyleId="DBDC680DC95E08776BDF6354C30A315B" docStyleName="SwissJPalaeontol.2022-.journal_article" docStyleVersion="1" docTitle="Simoniteuthis michaelyi Fuchs & Weis & Thuy 2024, n. sp." docType="treatment" docVersion="3" lastPageNumber="6" masterDocId="FF83CA7FEC35EC3FFFEA61747A3A824D" masterDocTitle="SimoniteUthiS, a new vampyromorph coleoid with prey in its arms from the Early Jurassic of Luxembourg" masterLastPageNumber="10" masterPageNumber="1" pageNumber="3" updateTime="1721659835942" updateUser="julia" zenodo-license-document="CC-BY-4.0">
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<mods:title id="912FAF136B28123C0DC97D1133289AC4">SimoniteUthiS, a new vampyromorph coleoid with prey in its arms from the Early Jurassic of Luxembourg</mods:title>
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<mods:name id="D8613BEB692D2D09A764EC1B78DDE820" type="personal">
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<mods:namePart id="A72EC375BD06D7131B0A3670A702BBEB">Fuchs, Dirk</mods:namePart>
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<mods:affiliation id="ED5DF95CABBC343B064B1386B0682391">Bayerische Staatssammlung FÜr Paläontologie Und Geologie, Richard-Wagner-Strasse 10, 80333 Munich, Germany</mods:affiliation>
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<mods:namePart id="1FD4D0291DAE64144923AEDA5C285C57">Weis, Robert</mods:namePart>
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<mods:affiliation id="785F6230EC6C719297875752AE22B25A">Musée National d’histoire Naturelle, Luxembourg, 25 Rue MÜnster,</mods:affiliation>
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<mods:namePart id="C9842E6821884E11C32180DBE9060139">Thuy, Ben</mods:namePart>
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<mods:affiliation id="697589A5570026840BA457D9F5C31226">Musée National d’histoire Naturelle, Luxembourg, 25 Rue MÜnster,</mods:affiliation>
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<mods:title id="8E39126E749DBB073903DCC3C6E9A567">Swiss Journal of Palaeontology</mods:title>
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<mods:date id="A46ECCC1F6BD21B91B5054057AD982BC">2024</mods:date>
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<mods:title id="3727294023E2F6BC7431BD8BD82AB222">6</mods:title>
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<mods:number id="025FD315FE5F5A2CF94D2C4362B0BF93">2024-02-14</mods:number>
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<mods:number id="3117496CA7C9852AFCE4090890FD4F68">143</mods:number>
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<mods:url id="A173196DDE530D11D8DA063445B0BD30">http://dx.doi.org/10.1186/s13358-024-00303-y</mods:url>
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<treatment id="03BAB207EC37EC3AFCC766757990816E" LSID="urn:lsid:plazi:treatment:03BAB207EC37EC3AFCC766757990816E" httpUri="http://treatment.plazi.org/id/03BAB207EC37EC3AFCC766757990816E" lastPageId="5" lastPageNumber="6" pageId="2" pageNumber="3">
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<subSubSection id="C309509AEC37EC3DFCC766757E648555" box="[813,1118,1793,1816]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8BAC0311EC37EC3DFCC766757E648555" blockId="2.[813,1437,1793,1912]" box="[813,1118,1793,1816]" pageId="2" pageNumber="3">
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<heading id="D0E4B47DEC37EC3DFCC766757E648555" box="[813,1118,1793,1816]" fontSize="9" level="2" pageId="2" pageNumber="3" reason="8">
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<emphasis id="B967DF03EC37EC3DFCC766757E648555" bold="true" box="[813,1118,1793,1816]" pageId="2" pageNumber="3">
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<taxonomicName id="4C137892EC37EC3DFCC766757E188555" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[813,1058,1793,1816]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="michaelyi" status="sp. nov.">
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<emphasis id="B967DF03EC37EC3DFCC766757E188555" bold="true" box="[813,1058,1793,1816]" italics="true" pageId="2" pageNumber="3">Simoniteuthis michaelyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2546278EC37EC3DFBCD66757E648555" box="[1063,1118,1793,1816]" pageId="2" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C309509AEC37EC3DFCC7665479968575" box="[813,940,1824,1848]" pageId="2" pageNumber="3" type="description">
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<paragraph id="8BAC0311EC37EC3DFCC7665479968575" blockId="2.[813,1437,1793,1912]" box="[813,940,1824,1848]" pageId="2" pageNumber="3">
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<figureCitation id="13281F94EC37EC3DFCC7665479968575" box="[813,940,1824,1848]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[167,198,1794,1814]" captionStartId-1="4.[167,198,1676,1696]" captionTargetBox-0="[168,1419,778,1780]" captionTargetBox-1="[285,1301,238,1663]" captionTargetId-0="figure-249@3.[166,1421,776,1782]" captionTargetId-1="figure-17@4.[283,1303,236,1664]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 2 Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian, Serpentinum Chronozone, Exaratum Subchronozone, Bascharage.A–D slab; E–G counter-slab. A overview; B camera lucida drawing of A; C close-up of the head–arm complex; D same under UV-light showing the weakly illuminating arm musculature; E overview; F close-up of the preyed fishes, red colour Specimen 1 (op=opercle; sop =subopercle),blue colour Specimen 2 (caud= caudal fin; sop =subopercle; centra =central vertebra); G same under UV-light. Scale bars =10 mm" captionText-1="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi-0="http://doi.org/10.5281/zenodo.12006472" figureDoi-1="http://doi.org/10.5281/zenodo.12006484" httpUri-0="https://zenodo.org/record/12006472/files/figure.png" httpUri-1="https://zenodo.org/record/12006484/files/figure.png" pageId="2" pageNumber="3">Figures 2–3</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C309509AEC37EC3DFCA8663479818535" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="8BAC0311EC37EC3DFCA8663479818535" blockId="2.[813,1437,1793,1912]" pageId="2" pageNumber="3">
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<materialsCitation id="3B7B094CEC37EC3DFCA8663479818535" collectionCode="MNHNL" pageId="2" pageNumber="3" specimenCode="TI024" specimenCount="1" typeStatus="holotype">
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<emphasis id="B967DF03EC37EC3DFCA86634798F8515" bold="true" box="[834,949,1856,1880]" pageId="2" pageNumber="3">
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<typeStatus id="54A8BDB3EC37EC3DFCA86634798A8515" box="[834,944,1856,1880]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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<collectionCode id="ED029BD4EC37EC3DFC2C66347E10851A" box="[966,1066,1856,1879]" pageId="2" pageNumber="3">MNHNL</collectionCode>
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<specimenCode id="DBB5AB6AEC37EC3DFBD066347E448515" box="[1082,1150,1856,1880]" collectionCode="TI" country="Japan" name="Herbarium of the Department of Botany, University of Tokyo" pageId="2" pageNumber="3">TI024</specimenCode>
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, consisting of slab and counter-slab.
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</materialsCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C309509AEC36EC3CFF47619F788483AE" pageId="3" pageNumber="4" type="distribution">
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<paragraph id="8BAC0311EC36EC3CFF47619F7BCE830E" blockId="3.[151,775,235,675]" pageId="3" pageNumber="4">
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<emphasis id="B967DF03EC36EC3CFF47619F7B68834E" bold="true" box="[173,338,235,259]" pageId="3" pageNumber="4">Type locality:</emphasis>
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Bascharage, Industrial Zone Edward Steichen (
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<figureCitation id="13281F94EC36EC3CFEE2607F7B73836E" box="[264,329,267,291]" captionStart="Fig" captionStartId="2.[167,198,1359,1379]" captionTargetBox="[283,1305,237,1345]" captionTargetId="figure-172@2.[282,1305,236,1346]" captionTargetPageId="2" captionText="Fig. 1 Detailed map of the Edward Steichen Industrial Zone, northeast of the town of Bascharage in southwest Luxembourg.The area marked by a black asterisk corresponds to the place of finding (© Carte topographique: Adm. Du Cadastre et de la Topographie,Luxembourg)" figureDoi="http://doi.org/10.5281/zenodo.12006463" httpUri="https://zenodo.org/record/12006463/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
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; formerly “Bommelscheier”) (
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<bibRefCitation id="EF827EE0EC36EC3CFD75607F7B31830E" author="Henrotay, M. & Marques, D. & Paicheler, J. - C. & Gall, J. - C. & Nel, A." pageId="3" pageNumber="4" refId="ref5814" refString="Henrotay, M., Marques, D., Paicheler, J. - C., Gall, J. - C., & Nel, A. (1998). Le Toarcien inferieur des regions de Bascharage et de Bettembourg (Grand-Duche" type="book" year="1998">Henrotay et al., 1998</bibRefCitation>
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;
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<bibRefCitation id="EF827EE0EC36EC3CFEFD605F7BDF830E" author="Szwedo, J. & Weis, R." box="[279,485,299,323]" pageId="3" pageNumber="4" pagination="806 - 812" refId="ref7254" refString="Szwedo, J., Weis, R. & A. Nel (2017). A bizarre sternorrhynchan wing from the Lower Jurassicof Luxembourg (Hemiptera: Sternorrhyncha: Pincombeomorpha?). Historical Biology, 31 (6), 806 - 812." type="journal article" year="2017">Szwedo et al., 2017</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8BAC0311EC36EC3CFF47603F788483AE" blockId="3.[151,775,235,675]" pageId="3" pageNumber="4">
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<emphasis id="B967DF03EC36EC3CFF47603F7B6E832E" bold="true" box="[173,340,331,355]" pageId="3" pageNumber="4">Type horizon:</emphasis>
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Near-continuous level of bituminous finely laminated carbonate nodules rich in insect and teleost remains, approximately
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<quantity id="4CEBAEF4EC36EC3CFE3960FF782D83EE" box="[467,535,395,419]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="3" pageNumber="4" unit="cm" value="90.0">90 cm</quantity>
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above the base of the bituminous black shale succession; Lower Toarcian, Serpentinum Chronozone, Exaratum Subchronozone.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309509AEC36EC3CFF47609F7843800E" pageId="3" pageNumber="4" type="etymology">
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<paragraph id="8BAC0311EC36EC3CFF47609F7843800E" blockId="3.[151,775,235,675]" pageId="3" pageNumber="4">
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<emphasis id="B967DF03EC36EC3CFF47609F7BA7804E" bold="true" box="[173,413,491,515]" pageId="3" pageNumber="4">Derivation of name:</emphasis>
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Dedicated to Patrick Michaely, Director of the Natural History Museum
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<collectingCountry id="F3044381EC36EC3CFD91637F793C806E" box="[635,774,523,547]" name="Luxembourg" pageId="3" pageNumber="4">Luxembourg</collectingCountry>
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with a passion for palaeontological research.
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</paragraph>
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</subSubSection>
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<subSubSection id="C309509AEC36EC3AFF47633F7B70846E" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
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<paragraph id="8BAC0311EC36EC3CFF47633F7EB7802E" blockId="3.[151,775,235,675]" lastBlockId="3.[813,1437,235,675]" pageId="3" pageNumber="4">
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<emphasis id="B967DF03EC36EC3CFF47633F7B07802E" bold="true" box="[173,317,587,611]" pageId="3" pageNumber="4">Description:</emphasis>
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The
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<typeStatus id="54A8BDB3EC36EC3CFE65633F7BD5802E" box="[399,495,587,611]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
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exhibits structures of the gladius as well as the head–arm complex. The latter is only visible on the slab (
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<figureCitation id="13281F94EC36EC3CFE1C63FF789380EE" box="[502,681,651,675]" captionStart="Fig" captionStartId="3.[167,198,1794,1814]" captionTargetBox="[168,1419,778,1780]" captionTargetId="figure-249@3.[166,1421,776,1782]" captionTargetPageId="3" captionText="Fig. 2 Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian, Serpentinum Chronozone, Exaratum Subchronozone, Bascharage.A–D slab; E–G counter-slab. A overview; B camera lucida drawing of A; C close-up of the head–arm complex; D same under UV-light showing the weakly illuminating arm musculature; E overview; F close-up of the preyed fishes, red colour Specimen 1 (op=opercle; sop =subopercle),blue colour Specimen 2 (caud= caudal fin; sop =subopercle; centra =central vertebra); G same under UV-light. Scale bars =10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006472" httpUri="https://zenodo.org/record/12006472/files/figure.png" pageId="3" pageNumber="4">Figs. 2A–D, 2G</figureCitation>
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,
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<figureCitation id="13281F94EC36EC3CFD5D63FF78C180EE" box="[695,763,651,675]" captionStart="Fig" captionStartId="4.[167,198,1676,1696]" captionTargetBox="[285,1301,238,1663]" captionTargetId="figure-17@4.[283,1303,236,1664]" captionTargetPageId="4" captionText="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="3" pageNumber="4">3A, B</figureCitation>
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); the counter-slab is unfortunately missing the head–arm complex (
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||
<figureCitation id="13281F94EC36EC3CFC70607F7E2B836E" box="[922,1041,267,291]" captionStart="Fig" captionStartId="3.[167,198,1794,1814]" captionTargetBox="[168,1419,778,1780]" captionTargetId="figure-249@3.[166,1421,776,1782]" captionTargetPageId="3" captionText="Fig. 2 Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian, Serpentinum Chronozone, Exaratum Subchronozone, Bascharage.A–D slab; E–G counter-slab. A overview; B camera lucida drawing of A; C close-up of the head–arm complex; D same under UV-light showing the weakly illuminating arm musculature; E overview; F close-up of the preyed fishes, red colour Specimen 1 (op=opercle; sop =subopercle),blue colour Specimen 2 (caud= caudal fin; sop =subopercle; centra =central vertebra); G same under UV-light. Scale bars =10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006472" httpUri="https://zenodo.org/record/12006472/files/figure.png" pageId="3" pageNumber="4">Figs. 2E, F</figureCitation>
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,
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<figureCitation id="13281F94EC36EC3CFBF4607F7E65836E" box="[1054,1119,267,291]" captionStart="Fig" captionStartId="4.[167,198,1676,1696]" captionTargetBox="[285,1301,238,1663]" captionTargetId="figure-17@4.[283,1303,236,1664]" captionTargetPageId="4" captionText="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="3" pageNumber="4">3D, E</figureCitation>
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). The head–arm complex of the slab is represented by proximal and middle parts of the arms, eyeballs, and the above-mentioned bony fish (
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<figureCitation id="13281F94EC36EC3CFCDE601F799083CE" box="[820,938,363,387]" captionStart="Fig" captionStartId="3.[167,198,1794,1814]" captionTargetBox="[168,1419,778,1780]" captionTargetId="figure-249@3.[166,1421,776,1782]" captionTargetPageId="3" captionText="Fig. 2 Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian, Serpentinum Chronozone, Exaratum Subchronozone, Bascharage.A–D slab; E–G counter-slab. A overview; B camera lucida drawing of A; C close-up of the head–arm complex; D same under UV-light showing the weakly illuminating arm musculature; E overview; F close-up of the preyed fishes, red colour Specimen 1 (op=opercle; sop =subopercle),blue colour Specimen 2 (caud= caudal fin; sop =subopercle; centra =central vertebra); G same under UV-light. Scale bars =10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006472" httpUri="https://zenodo.org/record/12006472/files/figure.png" pageId="3" pageNumber="4">Fig. 2C, D</figureCitation>
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). UV-light reveals scarce remains of the arm musculature (
|
||
<figureCitation id="13281F94EC36EC3CFC2D60FF7E2583EE" box="[967,1055,395,419]" captionStart="Fig" captionStartId="3.[167,198,1794,1814]" captionTargetBox="[168,1419,778,1780]" captionTargetId="figure-249@3.[166,1421,776,1782]" captionTargetPageId="3" captionText="Fig. 2 Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian, Serpentinum Chronozone, Exaratum Subchronozone, Bascharage.A–D slab; E–G counter-slab. A overview; B camera lucida drawing of A; C close-up of the head–arm complex; D same under UV-light showing the weakly illuminating arm musculature; E overview; F close-up of the preyed fishes, red colour Specimen 1 (op=opercle; sop =subopercle),blue colour Specimen 2 (caud= caudal fin; sop =subopercle; centra =central vertebra); G same under UV-light. Scale bars =10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006472" httpUri="https://zenodo.org/record/12006472/files/figure.png" pageId="3" pageNumber="4">Fig. 2D</figureCitation>
|
||
). The arm tips are obviously not conserved in the concretion. One can count eight arms; evidence of a rudimentary (filamentous) fifth arm pair does not exist. Further details such as suckers or arm length are not determinable. The two fishes are situated between two greyish eyeballs and obviously take the former position of the buccal mass.
|
||
</paragraph>
|
||
<paragraph id="8BAC0311EC36EC3AFCA8631F7B70846E" blockId="3.[813,1437,235,675]" lastBlockId="5.[151,776,235,1923]" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4">
|
||
Leaked ink unfortunately hampers the reconstruction of the gladius. The preserved length of the gladius measures
|
||
<quantity id="4CEBAEF4EC30EC3AFEEC619F7B71834E" box="[262,331,235,259]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="5" pageNumber="6" unit="cm" value="23.0">23 cm</quantity>
|
||
(
|
||
<figureCitation id="13281F94EC30EC3AFEB1619F7BF4834E" box="[347,462,235,259]" captionStart="Fig" captionStartId="4.[167,198,1676,1696]" captionTargetBox="[285,1301,238,1663]" captionTargetId="figure-17@4.[283,1303,236,1664]" captionTargetPageId="4" captionText="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="5" pageNumber="6">Fig. 3A–E</figureCitation>
|
||
). Only
|
||
<quantity id="4CEBAEF4EC30EC3AFDCA619F784F834E" box="[544,629,235,259]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="5" pageNumber="6" unit="cm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 cm</quantity>
|
||
of the conus part is missing as can be assessed by well-developed diverging inner asymptotes. The maximum width of the lateral fields reaches
|
||
<quantity id="4CEBAEF4EC30EC3AFE93603F7BFB832E" box="[377,449,331,355]" metricMagnitude="-2" metricUnit="m" metricValue="8.2" pageId="5" pageNumber="6" unit="cm" value="8.2">8.2 cm</quantity>
|
||
suggesting a moderately wide gladius (gladius
|
||
<subScript id="17970154EC30EC3AFEBA601F7B8D83CE" attach="left" box="[336,439,363,392]" fontSize="7" pageId="5" pageNumber="6">widthmax-</subScript>
|
||
to-gladius length
|
||
<emphasis id="B967DF03EC30EC3AFD90601F78B683CE" bold="true" box="[634,652,363,387]" pageId="5" pageNumber="6">=</emphasis>
|
||
0.33). Taking into account a certain degree of dorsoventral compaction the overall width was slightly less. Apart from this issue, parts of the lateral field are poorly visible. In particular, their anterior extensions are hidden behind mantle musculature and thus complicate interpretations. Outer asymptotes (opening angle c.15°) as well as growth increments suggest that the transition from the parabolar lateral field to the hyperbolar zone is sharp (pointed) and distinctly projected. The hyperbolar zone is accordingly V-shaped and deeply incised. The hyperbolar zone takes
|
||
<quantity id="4CEBAEF4EC30EC3AFF7D63BF7AE080AE" box="[151,218,715,739]" metricMagnitude="-1" metricUnit="m" metricValue="2.1" pageId="5" pageNumber="6" unit="cm" value="21.0">21 cm</quantity>
|
||
of the total median field length (ratio of the hyperbolar zone length to total gladius length
|
||
<emphasis id="B967DF03EC30EC3AFD83639F7841814E" bold="true" box="[617,635,747,771]" pageId="5" pageNumber="6">=</emphasis>
|
||
0.86), which is categorized as very long. Where the hyperbolar zone joins the median field, the latter is
|
||
<quantity id="4CEBAEF4EC30EC3AFDCF625F784B810E" box="[549,625,811,835]" metricMagnitude="-2" metricUnit="m" metricValue="4.3" pageId="5" pageNumber="6" unit="cm" value="4.3">4.3 cm</quantity>
|
||
wide. This is simultaneously the widest part of the median field; anteriorly, its width is decreasing. The ratio median field width at anterior end of the hyperbolar zone to hyperbolar zone length of 0.21 suggests a slender to very slender opening angle of the median field of 11–12°. The lateral margins of the median field (adjacent to the hyperbolar zones) are plate-like reinforced. Anteriorly, these diverging reinforcements reach a maximum width of
|
||
<quantity id="4CEBAEF4EC30EC3AFDB6655F7892860E" box="[604,680,1067,1091]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="5" pageNumber="6" unit="cm" value="0.9">0.9 cm</quantity>
|
||
and are slightly shorter than the anterior gladius margin. The shape of the latter, particularly the transition from lateral reinforcements to central median field is problematic due to the lack of informative growth increments (
|
||
<figureCitation id="13281F94EC30EC3AFD4C65DF78C0868E" box="[678,762,1195,1219]" captionStart="Fig" captionStartId="4.[167,198,1676,1696]" captionTargetBox="[285,1301,238,1663]" captionTargetId="figure-17@4.[283,1303,236,1664]" captionTargetPageId="4" captionText="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="5" pageNumber="6">Fig. 3C</figureCitation>
|
||
). Anteriorly, the lateral reinforcements appear parabolar (?and projected;
|
||
<figureCitation id="13281F94EC30EC3AFEBA659F7BF8874E" box="[336,450,1259,1283]" captionStart="Fig" captionStartId="4.[167,198,1676,1696]" captionTargetBox="[285,1301,238,1663]" captionTargetId="figure-17@4.[283,1303,236,1664]" captionTargetPageId="4" captionText="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="5" pageNumber="6">Fig. 3A–C</figureCitation>
|
||
) suggesting a second zone of hyperbolar growth lines intercalated between the parabolar reinforcements and parabolar central median field. Such a “secondary hyperbolar zone” is very unusual and a series of anteriorly parabolar structures imprinted in the leaked ink (?remains of growth structures) alternatively imply an evenly convex anterior gladius margin (
|
||
<figureCitation id="13281F94EC30EC3AFD4664DF78C1878E" box="[684,763,1451,1475]" captionStart="Fig" captionStartId="4.[167,198,1676,1696]" captionTargetBox="[285,1301,238,1663]" captionTargetId="figure-17@4.[283,1303,236,1664]" captionTargetPageId="4" captionText="Fig. 3 Gladius morphology of Simoniteuthis michaelyi n. gen. n. sp., holotype (MNHNL TI024), Lower Toarcian,Serpentinum Chronozone, Exaratum Subchronozone, Bascharage. A overview of the slab; B close-up of the anterior hyperbolar zone showing the course of growth increments; C schematic morphology and measurements; D overview of the counter-slab;E close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm" figureDoi="http://doi.org/10.5281/zenodo.12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="5" pageNumber="6">Fig. 3B</figureCitation>
|
||
). The median field bears a median line, but is insufficiently preserved for a detailed description. A median keel is certainly absent.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF6C5399EC36EC3CFF4D66767B2385CC" ID-DOI="http://doi.org/10.5281/zenodo.12006472" ID-Zenodo-Dep="12006472" httpUri="https://zenodo.org/record/12006472/files/figure.png" pageId="3" pageNumber="4" startId="3.[167,198,1794,1814]" targetBox="[168,1419,778,1780]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BAC0311EC36EC3CFF4D66767B2385CC" blockId="3.[167,1397,1794,1921]" pageId="3" pageNumber="4">
|
||
<emphasis id="B967DF03EC36EC3CFF4D66767AE0855B" bold="true" box="[167,218,1794,1814]" pageId="3" pageNumber="4">Fig. 2</emphasis>
|
||
<taxonomicName id="4C137892EC36EC3CFF0E66767B9C855B" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[228,422,1794,1814]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC36EC3CFF0E66767B9C855B" box="[228,422,1794,1814]" italics="true" pageId="3" pageNumber="4">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC36EC3CFE4166767BDC855B" box="[427,486,1794,1814]" pageId="3" pageNumber="4" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp., holotype (MNHNL TI024), Lower Toarcian,
|
||
<emphasis id="B967DF03EC36EC3CFC97667679DC855B" box="[893,998,1794,1814]" italics="true" pageId="3" pageNumber="4">Serpentinum</emphasis>
|
||
Chronozone,
|
||
<emphasis id="B967DF03EC36EC3CFBB666767E91855B" box="[1116,1195,1794,1814]" italics="true" pageId="3" pageNumber="4">Exaratum</emphasis>
|
||
Subchronozone, Bascharage.
|
||
<emphasis id="B967DF03EC36EC3CFEFA66697B02857C" bold="true" box="[272,312,1821,1841]" pageId="3" pageNumber="4">A–D</emphasis>
|
||
slab;
|
||
<emphasis id="B967DF03EC36EC3CFE8C66697BB0857C" bold="true" box="[358,394,1821,1841]" pageId="3" pageNumber="4">E–G</emphasis>
|
||
counter-slab.
|
||
<emphasis id="B967DF03EC36EC3CFDE86669782A857C" bold="true" box="[514,528,1821,1841]" pageId="3" pageNumber="4">A</emphasis>
|
||
overview;
|
||
<emphasis id="B967DF03EC36EC3CFD816669784D857C" bold="true" box="[619,631,1821,1841]" pageId="3" pageNumber="4">B</emphasis>
|
||
camera lucida drawing of
|
||
<emphasis id="B967DF03EC36EC3CFCB26669795C857C" bold="true" box="[856,870,1821,1841]" pageId="3" pageNumber="4">A</emphasis>
|
||
;
|
||
<emphasis id="B967DF03EC36EC3CFC8466697941857C" bold="true" box="[878,891,1821,1841]" pageId="3" pageNumber="4">C</emphasis>
|
||
close-up of the head–arm complex;
|
||
<emphasis id="B967DF03EC36EC3CFB5966697EF8857C" bold="true" box="[1203,1218,1821,1841]" pageId="3" pageNumber="4">D</emphasis>
|
||
same under UV-light showing the weakly illuminating arm musculature;
|
||
<emphasis id="B967DF03EC36EC3CFDB3664C785E8501" bold="true" box="[601,612,1848,1868]" pageId="3" pageNumber="4">E</emphasis>
|
||
overview;
|
||
<emphasis id="B967DF03EC36EC3CFD2A664C78F18501" bold="true" box="[704,715,1848,1868]" pageId="3" pageNumber="4">F</emphasis>
|
||
close-up of the preyed fishes, red colour Specimen 1 (op=opercle; sop =subopercle),blue colour Specimen 2 (caud= caudal fin; sop =subopercle; centra =central vertebra);
|
||
<emphasis id="B967DF03EC36EC3CFBCC66267E0E852B" bold="true" box="[1062,1076,1874,1894]" pageId="3" pageNumber="4">G</emphasis>
|
||
same under UV-light. Scale bars =10 mm
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6C5399EC31EC3BFF4D67F87B8884BD" ID-DOI="http://doi.org/10.5281/zenodo.12006484" ID-Zenodo-Dep="12006484" httpUri="https://zenodo.org/record/12006484/files/figure.png" pageId="4" pageNumber="5" startId="4.[167,198,1676,1696]" targetBox="[285,1301,238,1663]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BAC0311EC31EC3BFF4D67F87B8884BD" blockId="4.[167,1395,1676,1776]" pageId="4" pageNumber="5">
|
||
<emphasis id="B967DF03EC31EC3BFF4D67F87AE084ED" bold="true" box="[167,218,1676,1696]" pageId="4" pageNumber="5">Fig. 3</emphasis>
|
||
Gladius morphology of
|
||
<taxonomicName id="4C137892EC31EC3BFE4767F8785484ED" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[429,622,1676,1696]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC31EC3BFE4767F8785484ED" box="[429,622,1676,1696]" italics="true" pageId="4" pageNumber="5">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC31EC3BFD9E67F8789484ED" box="[628,686,1676,1696]" pageId="4" pageNumber="5" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp., holotype (MNHNL TI024), Lower Toarcian,
|
||
<emphasis id="B967DF03EC31EC3BFBAF67F87E9484ED" box="[1093,1198,1676,1696]" italics="true" pageId="4" pageNumber="5">Serpentinum</emphasis>
|
||
Chronozone,
|
||
<emphasis id="B967DF03EC31EC3BFACE67F87F4984ED" box="[1316,1395,1676,1696]" italics="true" pageId="4" pageNumber="5">Exaratum</emphasis>
|
||
Subchronozone, Bascharage.
|
||
<emphasis id="B967DF03EC31EC3BFE4A67D37B9484F6" bold="true" box="[416,430,1703,1723]" pageId="4" pageNumber="5">A</emphasis>
|
||
overview of the slab;
|
||
<emphasis id="B967DF03EC31EC3BFD8C67D3784884F6" bold="true" box="[614,626,1703,1723]" pageId="4" pageNumber="5">B</emphasis>
|
||
close-up of the anterior hyperbolar zone showing the course of growth increments;
|
||
<emphasis id="B967DF03EC31EC3BFAAA67D37F7784F6" bold="true" box="[1344,1357,1703,1723]" pageId="4" pageNumber="5">C</emphasis>
|
||
schematic morphology and measurements;
|
||
<emphasis id="B967DF03EC31EC3BFDF567B67814849B" bold="true" box="[543,558,1730,1750]" pageId="4" pageNumber="5">D</emphasis>
|
||
overview of the counter-slab;
|
||
<emphasis id="B967DF03EC31EC3BFCC467B67903849B" bold="true" box="[814,825,1730,1750]" pageId="4" pageNumber="5">E</emphasis>
|
||
close-up of the posterior gladius showing the course of growth increments. Scale bars=10 mm
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C309509AEC30EC3AFF47675F7990816E" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="8BAC0311EC30EC3AFF47675F7BF1854E" blockId="5.[151,776,235,1923]" pageId="5" pageNumber="6">
|
||
<emphasis id="B967DF03EC30EC3AFF47675F7B74840E" bold="true" box="[173,334,1579,1603]" pageId="5" pageNumber="6">Comparisons:</emphasis>
|
||
The cluster analysis (
|
||
<figureCitation id="13281F94EC30EC3AFDD2675F78B3840E" box="[568,649,1579,1603]" captionStart="Fig" captionStartId="6.[167,198,1101,1121]" captionTargetBox="[170,1418,237,1087]" captionTargetId="figure-363@6.[169,1419,236,1088]" captionTargetPageId="6" captionText="Fig. 4 Comparative gladius morphology of selected octobrachiate coleoids.A Topology of the cluster analysis based on gladius outlines; B 3-reconstructions of the same gladiuses systematically sorted (after Fuchs, 2020). Where known arm configurations are indicated. Note the gladius outline of Vampyrofugies (based on Rowe et al., 2023) is approximated" figureDoi="http://doi.org/10.5281/zenodo.12006496" httpUri="https://zenodo.org/record/12006496/files/figure.png" pageId="5" pageNumber="6">Fig. 4A</figureCitation>
|
||
) of the gladius outline exposes
|
||
<taxonomicName id="4C137892EC30EC3AFE6B673F7B3684CE" authority="Rowe et al., 2023" authorityName="Rowe" authorityYear="2023" class="Cephalopoda" family="Geopeltidae" genus="Vampyrofugiens" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="atramentum">
|
||
<emphasis id="B967DF03EC30EC3AFE6B673F78F8842E" box="[385,706,1611,1635]" italics="true" pageId="5" pageNumber="6">Vampyrofugiens atramentum</emphasis>
|
||
<bibRefCitation id="EF827EE0EC30EC3AFD2167387B3684CE" author="Rowe, A. & Kruta, I. & Villier, L. & Rouget, I." pageId="5" pageNumber="6" refId="ref7043" refString="Rowe, A., Kruta, I., Villier, L., & Rouget, I. (2023). A new vampyromorph species from the Middle Jurassic La Voulte-sur-Rhone Lagerstatte. Papers in Palaeontology, 9 (3), e 1511. https: // doi. org / 10.1002 / spp 2.1511" type="journal volume" year="2023">Rowe et al., 2023</bibRefCitation>
|
||
</taxonomicName>
|
||
from the Callovian of La Voulte-sur-Rhône to be most similar to
|
||
<taxonomicName id="4C137892EC30EC3AFE8167FF784B84EE" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[363,625,1675,1699]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC30EC3AFE8167FF784B84EE" box="[363,625,1675,1699]" italics="true" pageId="5" pageNumber="6">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC30EC3AFD9067F878FE84E9" box="[634,708,1676,1700]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp. However, the gladius of this deep-sea vampyromorph is based solely on a tomographic outline reconstruction and thus still poorly understood.
|
||
</paragraph>
|
||
<paragraph id="8BAC0311EC30EC3AFF47667F7F6183AE" blockId="5.[151,776,235,1923]" lastBlockId="5.[813,1437,235,803]" pageId="5" pageNumber="6">
|
||
Besides, the gladius shape of
|
||
<taxonomicName id="4C137892EC30EC3AFDE7667F7A88850E" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC30EC3AFDE7667F7A88850E" italics="true" pageId="5" pageNumber="6">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC30EC3AFF2966587B2D8509" box="[195,279,1836,1860]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp. is most similar to contemporary geopeltid (
|
||
<emphasis id="B967DF03EC30EC3AFEFD663F78C4852E" box="[279,766,1867,1891]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C137892EC30EC3AFEFD663F7BDE852E" box="[279,484,1867,1891]" class="Cephalopoda" family="Geopeltidae" genus="Geopeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="simplex">Geopeltis simplex</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C137892EC30EC3AFE13663F78C4852E" box="[505,766,1867,1891]" class="Cephalopoda" family="Loligosepiidae" genus="Parabelopeltis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="flexuosa">Parabelopeltis flexuosa</taxonomicName>
|
||
</emphasis>
|
||
) and loligosepiid vampyromorphs (
|
||
<emphasis id="B967DF03EC30EC3AFDC9661F7E07834E" box="[547,1085,235,1923]" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C137892EC30EC3AFDC9661F793985CE" box="[547,771,1899,1923]" class="Cephalopoda" family="Loligosepiidae" genus="Loligosepia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="aalensis">Loligosepia aalensis</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C137892EC30EC3AFCC7619F7E07834E" box="[813,1085,235,259]" class="Cephalopoda" genus="Jeletzkyteuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="coriaceus">Jeletzkyteuthis coriaceus</taxonomicName>
|
||
</emphasis>
|
||
). The distinctly forward-projected lateral fields combined with the deeply incised V-shaped hyperbolar zones are typical for loligosepiids, although the relative length of the hyperbolar zones is slightly shorter in
|
||
<taxonomicName id="4C137892EC30EC3AFC11601F7F3983CE" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[1019,1283,363,387]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC30EC3AFC11601F7F3983CE" box="[1019,1283,363,387]" italics="true" pageId="5" pageNumber="6">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC30EC3AFAE660187F6D83C9" box="[1292,1367,364,388]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp. than in loligosepiids. In geopeltids, the hyperbolar zone is shorter than in
|
||
<taxonomicName id="4C137892EC30EC3AFC1060DF7F38838E" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[1018,1282,427,451]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC30EC3AFC1060DF7F38838E" box="[1018,1282,427,451]" italics="true" pageId="5" pageNumber="6">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC30EC3AFAE160D87F6D8389" box="[1291,1367,428,452]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp. and evenly arcuated (rather than sharply recurved).
|
||
</paragraph>
|
||
<paragraph id="8BAC0311EC30EC3AFCA860987990816E" blockId="5.[813,1437,235,803]" pageId="5" pageNumber="6">
|
||
The plate-like lateral reinforcements as described for
|
||
<taxonomicName id="4C137892EC30EC3AFCC7637F7E0B806E" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" box="[813,1073,523,547]" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC30EC3AFCC7637F7E0B806E" box="[813,1073,523,547]" italics="true" pageId="5" pageNumber="6">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC30EC3AFBD263787EBA8069" box="[1080,1152,524,548]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp. are unusual for loligosepiids, but known from
|
||
<emphasis id="B967DF03EC30EC3AFBB0635F79AF802E" italics="true" pageId="5" pageNumber="6">
|
||
<taxonomicName id="4C137892EC30EC3AFBB0635F7F26800E" box="[1114,1308,555,579]" class="Cephalopoda" family="Geopeltidae" genus="Geopeltis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="simplex">Geopeltis simplex</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C137892EC30EC3AFACD635F79AF802E" class="Cephalopoda" genus="Leptotheuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="gigas">Leptotheuthis gigas</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C137892EC30EC3AFC38633F7EC2802E" box="[978,1272,587,611]" class="Cephalopoda" family="Plesioteuthididae" genus="Paraplesioteuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="sagittata">
|
||
<emphasis id="B967DF03EC30EC3AFC38633F7EC2802E" box="[978,1272,587,611]" italics="true" pageId="5" pageNumber="6">Paraplesioteuthis sagittata</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13281F94EC30EC3AFAE0633F7F66802E" box="[1290,1372,587,611]" captionStart="Fig" captionStartId="6.[167,198,1101,1121]" captionTargetBox="[170,1418,237,1087]" captionTargetId="figure-363@6.[169,1419,236,1088]" captionTargetPageId="6" captionText="Fig. 4 Comparative gladius morphology of selected octobrachiate coleoids.A Topology of the cluster analysis based on gladius outlines; B 3-reconstructions of the same gladiuses systematically sorted (after Fuchs, 2020). Where known arm configurations are indicated. Note the gladius outline of Vampyrofugies (based on Rowe et al., 2023) is approximated" figureDoi="http://doi.org/10.5281/zenodo.12006496" httpUri="https://zenodo.org/record/12006496/files/figure.png" pageId="5" pageNumber="6">Fig. 4B</figureCitation>
|
||
). The latter is a contemporary taxon and belongs to the prototeuthid clade owing to the absence of a hyperbolar zone. Tithonian
|
||
<taxonomicName id="4C137892EC30EC3AFC4B63DF7E49808E" box="[929,1139,683,707]" class="Cephalopoda" genus="Leptotheuthis" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="gigas">
|
||
<emphasis id="B967DF03EC30EC3AFC4B63DF7E49808E" box="[929,1139,683,707]" italics="true" pageId="5" pageNumber="6">Leptotheuthis gigas</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C137892EC30EC3AFAED63DF79AC80AE" authority="Fuchs & Weis & Thuy, 2024" authorityName="Fuchs & Weis & Thuy" authorityYear="2024" class="Cephalopoda" genus="Simoniteuthis" higherTaxonomySource="CoL" kingdom="Animalia" order="Vampyromorpha" pageId="5" pageNumber="6" phylum="Mollusca" rank="species" species="michaelyi" status="n. gen.">
|
||
<emphasis id="B967DF03EC30EC3AFAED63DF79AC80AE" italics="true" pageId="5" pageNumber="6">Simoniteuthis michaelyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2546278EC30EC3AFC7763B879DC80A9" box="[925,998,716,740]" pageId="5" pageNumber="6" rank="genus">n. gen.</taxonomicNameLabel>
|
||
n. sp. through a wider median field and a constriction of the lateral fields in the posterior third of the gladius.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |