657 lines
82 KiB
XML
657 lines
82 KiB
XML
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<mods:title id="B67F999AC9ED3E2AF88BC59C9BFE403E">Integrative revision of the Lygodactylus gutturalis (Bocage, 1873) complex unveils extensive cryptic diversity and traces its evolutionary history</mods:title>
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<mods:namePart id="CBE136033D3FBB32C312BA07B26419AD">Lobón-Rovira, Javier</mods:namePart>
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<mods:namePart id="D2A4C900900DA64F665572B8E588ACE5">Kusamba, Chifundera</mods:namePart>
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<mods:namePart id="D1D19CA77050B30FBBA91437825D4A19">Júlio, Timóteo</mods:namePart>
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<mods:namePart id="A727EC8486CF73C79D3D9729A6B527EA">Pauwels, Olivier S. G.</mods:namePart>
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<mods:date id="7611B099C456D36C12E602813522AE84">2024</mods:date>
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<treatment id="03B30955FFCAFFDEFF1F46B0FD56FE68" ID-DOI="http://doi.org/10.5281/zenodo.13302871" ID-Zenodo-Dep="13302871" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68" httpUri="http://treatment.plazi.org/id/03B30955FFCAFFDEFF1F46B0FD56FE68" lastPageId="33" lastPageNumber="480" pageId="29" pageNumber="476">
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<heading id="D0ED0F2FFFCAFFE2FF1F46B0FD1EF950" allCaps="true" bold="true" box="[221,658,1738,1764]" centered="true" fontSize="11" level="1" pageId="29" pageNumber="476" reason="1">
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<taxonomicName id="4C1AC3C0FFCAFFE2FF1F46B0FDA7F950" ID-CoL="5HP8" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[221,555,1738,1764]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="hramoja" status="sp. nov.">
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<emphasis id="B96E6451FFCAFFE2FF1F46B0FDA7F950" bold="true" box="[221,555,1738,1764]" italics="true" pageId="29" pageNumber="476">LYGODACTYLUS HRAMOJA</emphasis>
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<taxonomicNameLabel id="A25DD92AFFCAFFE2FDED46B0FD1EF950" box="[559,658,1738,1764]" pageId="29" pageNumber="476" rank="species">SP.NOV.</taxonomicNameLabel>
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</emphasis>
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<subSubSection id="C300EBC8FFCAFFE2FFB3468BFE16F8D5" pageId="29" pageNumber="476" type="description">
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<paragraph id="8BA5B843FFCAFFE2FFB3468BFD76F8BD" blockId="29.[113,762,1777,1801]" box="[113,762,1777,1801]" pageId="29" pageNumber="476">
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(
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<figureCitation id="1321A4C6FFCAFFE2FFBE468BFF67F8BD" box="[124,235,1777,1801]" captionStart-0="Figure 19" captionStart-1="Figure 20" captionStartId-0="31.[115,180,1898,1922]" captionStartId-1="32.[128,193,1898,1922]" captionTargetBox-0="[114,1458,920,1870]" captionTargetBox-1="[129,1473,449,1870]" captionTargetId-0="figure-423@31.[114,1458,920,1870]" captionTargetId-1="figure-170@32.[129,1473,449,1870]" captionTargetPageId-0="31" captionTargetPageId-1="32" captionText-0="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." captionText-1="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi-0="http://doi.org/10.5281/zenodo.13302905" figureDoi-1="http://doi.org/10.5281/zenodo.13302907" httpUri-0="https://zenodo.org/record/13302905/files/figure.png" httpUri-1="https://zenodo.org/record/13302907/files/figure.png" pageId="29" pageNumber="476">Figs 19–20</figureCitation>
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,
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<tableCitation id="C6988DF8FFCAFFE2FF37468BFEB3F8BD" box="[245,319,1777,1801]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="29" pageNumber="476" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
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; Supporting Information,
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<figureCitation id="1321A4C6FFCAFFE2FD8B468BFD01F8BD" box="[585,653,1777,1801]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="29" pageNumber="476">Fig. S7</figureCitation>
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; Table S8)
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</paragraph>
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<emphasis id="B96E6451FFCAFFE2FFB34750FEC1F8F6" box="[113,333,1834,1858]" italics="true" pageId="29" pageNumber="476">Zoobank registration:</emphasis>
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https://zoobank.org/
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<subSubSection id="C300EBC8FFCAFFE2FF5247FBFA85FAFA" pageId="29" pageNumber="476" type="reference_group">
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<paragraph id="8BA5B843FFCAFFE2FF5247FBFD10F80F" blockId="29.[144,732,1921,1979]" pageId="29" pageNumber="476">
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<emphasis id="B96E6451FFCAFFE2FF5247FBFD10F80F" italics="true" pageId="29" pageNumber="476">
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<taxonomicName id="4C1AC3C0FFCAFFE2FF5247FBFE62F82F" ID-CoL="3WQ3T" authorityName="' (Schmidt" authorityYear="1919" box="[144,494,1921,1947]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="subSpecies" species="picturatus" subSpecies="gutturalis">Lygodactylus picturatus gutturalis</taxonomicName>
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:
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<bibRefCitation id="EF8BC5B2FFCAFFE2FE3B47FBFD19F82F" author="Schmidt KP" box="[505,661,1921,1947]" pageId="29" pageNumber="476" pagination="385 - 624" refId="ref37222" refString="Schmidt KP. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909 - 1915. Part I: turtles, crocodiles, lizards, and chamaeleons. Bulletin of the American Museum of Natural History 1919; 39: 385 - 624." type="journal article" year="1919">Schmidt (1919)</bibRefCitation>
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[part];
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<bibRefCitation id="EF8BC5B2FFCAFFE2FF1347DBFEE8F80E" author="Pasteur G" box="[209,356,1952,1978]" pageId="29" pageNumber="476" pagination="1 - 132" refId="ref36328" refString="Pasteur G. Recherches sur l'evolution des lygodactyles, lezards Afro- Malgaches actuels. Travaux de l'Institut Scientifique Cherifien, Zoologie 1965 (1964); 29: 1 - 132." type="journal article" year="1964">Pasteur (1964)</bibRefCitation>
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[part];
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||
<bibRefCitation id="EF8BC5B2FFCAFFE2FE6D47DAFDDAF80E" author="Wermuth H" box="[431,598,1952,1978]" pageId="29" pageNumber="476" pagination="1 - 246" refId="ref37733" refString="Wermuth H. Liste der rezenten Amphibien und Reptilien. Gekkonidae, Pygopodidae, Xantusiidae. Das Tierreich 1965; 80: I - XXIII, 1 - 246." type="journal article" year="1965">Wermuth (1965)</bibRefCitation>
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[part].
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</emphasis>
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<paragraph id="8BA5B843FFCAFFE2FCEC456EFA85FAFA" blockId="29.[810,1459,1300,1645]" pageId="29" pageNumber="476">
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|
||
<taxonomicName id="4C1AC3C0FFCAFFE2FCEC456EFB92FA9A" ID-CoL="3WQ3T" authority="(Bocage, 1873)" box="[814,1054,1300,1326]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="gutturalis">Lygodactylus gutturalis</taxonomicName>
|
||
:
|
||
<bibRefCitation id="EF8BC5B2FFCAFFE2FBEB456EFB12FA9B" author="Roll B" box="[1065,1182,1300,1327]" pageId="29" pageNumber="476" pagination="645 - 8" refId="ref36800" refString="Roll B. Lygodactylus gutturalis (Bocage). Sauria (Suppl.) 2005; 27: 645 - 8." type="journal article" year="2005">Röll (2005)</bibRefCitation>
|
||
[part];
|
||
<bibRefCitation id="EF8BC5B2FFCAFFE2FB28456EFA23FA9B" author="Spawls S & Howell K & Hinkel H & Menegon M" box="[1258,1455,1300,1327]" pageId="29" pageNumber="476" refId="ref37334" refString="Spawls S, Howell K, Hinkel H, Menegon M. 2018. Field Guide to East African Reptiles. London: Bloomsbury." type="book" year="2018">Spawls et al. (2018)</bibRefCitation>
|
||
[part]; Behangana et al. (2020)
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300EBC8FFCAFFE2FCE84520FAE1F9D9" pageId="29" pageNumber="476" type="discussion">
|
||
<paragraph id="8BA5B843FFCAFFE2FCE84520FAE1F9D9" blockId="29.[810,1459,1300,1645]" pageId="29" pageNumber="476">
|
||
<emphasis id="B96E6451FFCAFFE2FCE84520FB0DFAC6" bold="true" box="[810,1153,1370,1394]" pageId="29" pageNumber="476">
|
||
<taxonomicName id="4C1AC3C0FFCAFFE2FCE84520FBA8FAC6" ID-CoL="C7JCW" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[810,1060,1370,1394]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFCAFFE2FCE84520FBA8FAC6" bold="true" box="[810,1060,1370,1394]" italics="true" pageId="29" pageNumber="476">Lygodactylus karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFCAFFE2FBEE4520FB0DFAC6" box="[1068,1153,1370,1394]" pageId="29" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from the northern Albertine Rift in the Lendu Plateau of
|
||
<collectingCountry id="F30DF8D3FFCAFFE2FB014500FB75FA26" box="[1219,1273,1402,1426]" name="Democratic Republic of the Congo" pageId="29" pageNumber="476">DRC</collectingCountry>
|
||
and several highlands of
|
||
<collectingCountry id="F30DF8D3FFCAFFE2FC4145E3FC59FA05" box="[899,981,1433,1457]" name="Uganda" pageId="29" pageNumber="476">Uganda</collectingCountry>
|
||
, and it is the sister taxon to
|
||
<emphasis id="B96E6451FFCAFFE2FB3845E3FA3FFA05" bold="true" box="[1274,1459,1432,1457]" pageId="29" pageNumber="476">
|
||
<taxonomicName id="4C1AC3C0FFCAFFE2FB3845E3FAD6FA04" ID-CoL="C7JCX" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1274,1370,1432,1456]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFCAFFE2FB3845E3FAD6FA04" bold="true" box="[1274,1370,1432,1456]" italics="true" pageId="29" pageNumber="476">L. kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFCAFFE2FAA245E3FA3FFA05" box="[1376,1459,1433,1457]" pageId="29" pageNumber="476" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in clade (B1), from which it differs by
|
||
<emphasis id="B96E6451FFCAFFE2FB6F45C3FB3AFA64" box="[1197,1206,1465,1488]" italics="true" pageId="29" pageNumber="476">c</emphasis>
|
||
. 6.07% for the
|
||
<emphasis id="B96E6451FFCAFFE2FA8D45C2FAFAFA64" box="[1359,1398,1464,1488]" italics="true" pageId="29" pageNumber="476">16S</emphasis>
|
||
mitochondrial gene (Table 2) and a lack of nuclear haplotype sharing in
|
||
<emphasis id="B96E6451FFCAFFE2FC86458DFC0DF9BB" box="[836,897,1527,1551]" italics="true" pageId="29" pageNumber="476">RAG1</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFCAFFE2FC50458DFC77F9BB" box="[914,1019,1527,1551]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
|
||
). The new species differs from
|
||
<taxonomicName id="4C1AC3C0FFCAFFE2FAFF4582FA3EF9BB" ID-CoL="3WQ3T" box="[1341,1458,1527,1551]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFCAFFE2FAFF4582FA3EF9BB" box="[1341,1458,1527,1551]" italics="true" pageId="29" pageNumber="476">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C1AC3C0FFCAFFE2FC9A466DFC42F99A" ID-CoL="3WQ3G" authorityName=", Schmidt" authorityYear="1919" box="[856,974,1558,1582]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="476" phylum="Chordata" rank="species" species="depressus">
|
||
<emphasis id="B96E6451FFCAFFE2FC9A466DFC42F99A" box="[856,974,1558,1582]" italics="true" pageId="29" pageNumber="476">L. depressus</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<emphasis id="B96E6451FFCAFFE2FC34466DFC73F99A" box="[1014,1023,1559,1582]" italics="true" pageId="29" pageNumber="476">c</emphasis>
|
||
. 12.49% and 10.45%, respectively, for the
|
||
<emphasis id="B96E6451FFCAFFE2FCE8464FFCDDF9F9" box="[810,849,1589,1613]" italics="true" pageId="29" pageNumber="476">16S</emphasis>
|
||
(uncorrected p-distance) mitochondrial gene (Table 2) and a lack of nuclear haplotype sharing in
|
||
<emphasis id="B96E6451FFCAFFE2FB6A462FFB69F9D9" box="[1192,1253,1621,1645]" italics="true" pageId="29" pageNumber="476">RAG1</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFCAFFE2FB37462FFAD0F9D9" box="[1269,1372,1621,1645]" captionStart="Figure 2" captionStartId="11.[113,178,1269,1293]" captionTargetBox="[117,1455,147,1239]" captionTargetId="figure-288@11.[114,1458,144,1242]" captionTargetPageId="11" captionText="Figure 2. A, Multilocus species tree and Bayesian chronogram of divergences among Afro-American Lygodactylus clade (Clade C of Gippner et al. 2021), based on an alignment of 13 concatenated markers (including eight nuclear-encoded genes, four protein-coding mitochondrial genes, and the 16S mitochondrial rRNA gene, see Gippner et al.2021). Branches with posterior probability> 0.9 are denoted by asterisks (*) at relevant nodes. Blue bars depict 95% High Probability Density (HPD) intervals of estimated divergence dates.B, Median-joining nuclear allele network showing the relationships between the L. picturatus-fischeri groups, inferred from the RAG1 nuclear gene (262 bp including all available RAG1 sequences of the L. picturatus-fischeri groups).C, Median-joining nuclear allele network of the RAG1 nuclear gene (980 bp) showing the relationships within the L. gutturalis subgroup, excluding L. gutturalis s.s. In all networks, circle frequency of alleles and small lines represent mutational steps." figureDoi="http://doi.org/10.5281/zenodo.13302840" httpUri="https://zenodo.org/record/13302840/files/figure.png" pageId="29" pageNumber="476">Fig. 2B–C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300EBC8FFCAFFE1FCE846F4FD35FA0C" lastPageId="30" lastPageNumber="477" pageId="29" pageNumber="476" type="materials_examined">
|
||
<paragraph id="8BA5B843FFCAFFE2FCE846F4FB5BF897" blockId="29.[810,1459,1678,1827]" pageId="29" pageNumber="476">
|
||
<materialsCitation id="3B72B21EFFCAFFE2FCE846F4FB5BF897" collectingDate="2015-07-05" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes & Wilber Lukwago & Mathias Behangana." country="Uganda" county="Northern Region" elevation="1193" latitude="3.83156" location="Agoro Town" longLatPrecision="1" longitude="33.01974" municipality="Imatong Foothills" pageId="29" pageNumber="476" specimenCode="UTEP 22590, DFH 593" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B96E6451FFCAFFE2FCE846F4FC06F912" box="[810,906,1678,1702]" italics="true" pageId="29" pageNumber="476">
|
||
<typeStatus id="54A106E1FFCAFFE2FCE846F4FC09F912" box="[810,901,1678,1702]" pageId="29" pageNumber="476" type="holotype">Holotype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCode id="DBBC1038FFCAFFE2FC5C46F5FBA0F912" box="[926,1068,1678,1702]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22590</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFCAFFE2FBFF46F4FB20F912" box="[1085,1196,1678,1702]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 593</specimenCode>
|
||
), male with original tail, collected on a large tree at the edge of a village at
|
||
<location id="8EC5EE98FFCAFFE2FAE946D4FA23F971" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68:8EC5EE98FFCAFFE2FAE946D4FA23F971" box="[1323,1455,1710,1734]" country="Uganda" county="Northern Region" latitude="3.83156" longLatPrecision="1" longitude="33.01974" municipality="Imatong Foothills" name="Agoro Town" pageId="29" pageNumber="476">Agoro Town</location>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFCAFFE2FCE846B7FC61F951" box="[810,1005,1741,1765]" pageId="29" pageNumber="476">Imatong Foothills</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C4C0CFFFCAFFE2FBC346B7FB33F951" box="[1025,1215,1741,1765]" pageId="29" pageNumber="476">Northern Region</collectingCounty>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFCAFFE2FB1146B7FAA9F951" box="[1235,1317,1741,1765]" name="Uganda" pageId="29" pageNumber="476">Uganda</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFCAFFE2FAFB46B7FA23F951" box="[1337,1455,1741,1765]" degrees="03.83156" direction="north" orientation="latitude" pageId="29" pageNumber="476" precision="1" value="3.83156">N03.83156</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFCAFFE2FCE84696FC10F8B0" box="[810,924,1772,1796]" degrees="33.01974" direction="east" orientation="longitude" pageId="29" pageNumber="476" precision="1" value="33.01974">E33.01974</geoCoordinate>
|
||
,
|
||
<elevation id="00375F70FFCAFFE2FC6A4696FBA0F8B0" box="[936,1068,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="29" pageNumber="476" unit="m" value="1193.0">
|
||
<quantity id="4CE215A6FFCAFFE2FC6A4696FC75F8B0" box="[936,1017,1772,1796]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="29" pageNumber="476" unit="m" value="1193.0">1193 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFCAFFE2FB954696FB43F8B0" box="[1111,1231,1772,1796]" pageId="29" pageNumber="476" value="2015-07-05">
|
||
<collectingDate id="EFE0676BFFCAFFE2FB954696FB43F8B0" box="[1111,1231,1772,1796]" pageId="29" pageNumber="476" value="2015-07-05">5 July 2015</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFCAFFE2FB3A4696FA23F8B0" box="[1272,1455,1772,1796]" pageId="29" pageNumber="476">Daniel F. Hughes</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFCAFFE2FCE84771FC58F897" box="[810,980,1803,1827]" pageId="29" pageNumber="476">Wilber Lukwago</collectorName>
|
||
, and
|
||
<collectorName id="26EFDD95FFCAFFE2FBCA4771FB5BF897" box="[1032,1239,1803,1827]" pageId="29" pageNumber="476">Mathias Behangana.</collectorName>
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFCAFFE1FCE8473CFD35FA0C" blockId="29.[810,1459,1861,1979]" lastBlockId="30.[129,778,1284,1465]" lastPageId="30" lastPageNumber="477" pageId="29" pageNumber="476">
|
||
<emphasis id="B96E6451FFCAFFE2FCE8473CFBB8F8E8" box="[810,1076,1861,1885]" italics="true" pageId="29" pageNumber="476">
|
||
<typeStatus id="54A106E1FFCAFFE2FCE8473CFC05F8E9" box="[810,905,1862,1885]" pageId="29" pageNumber="476" type="paratype">Paratypes</typeStatus>
|
||
(
|
||
<specimenCount id="9D1C73CAFFCAFFE2FC5E473FFBA4F8E8" box="[924,1064,1861,1884]" count="6" pageId="29" pageNumber="476" type="generic">six specimens</specimenCount>
|
||
):
|
||
</emphasis>
|
||
<materialsCitation id="3B72B21EFFCAFFE2FB89473FFCDCF8C8" collectionCode="UTEP, DFH" pageId="29" pageNumber="476" specimenCode="UTEP 22588-89, DFH 591-92" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFCAFFE2FB89473FFA8EF8E9" box="[1099,1282,1861,1885]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22588–89</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFCAFFE2FAD7473FFA20F8E9" box="[1301,1452,1861,1885]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 591–92</specimenCode>
|
||
) and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B72B21EFFCAFFE2FC94471FFC5FF82F" collectingDate="2015-07-05" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes & Wilber Lukwago & Mathias Behangana." country="Uganda" county="Northern Region" elevation="1193" latitude="3.83156" location="Agoro Town" longLatPrecision="1" longitude="33.01974" municipality="Imatong Foothills" pageId="29" pageNumber="476" specimenCode="UTEP 22591, DFH 594" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFCAFFE2FC94471FFC6CF8C8" box="[854,992,1892,1916]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22591</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFCAFFE2FC2F471EFBD4F8C8" box="[1005,1112,1892,1916]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 594</specimenCode>
|
||
), males, with same collection data as the holotype
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFCAFFE1FC2247FEFD34FA8F" collectingDate="2015-07-05" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes & Wilber Lukwago & Mathias Behangana" country="Uganda" county="Northern Region" elevation="1153" lastPageId="30" lastPageNumber="477" latitude="3.80548" location="Agoro Town" longLatPrecision="1" longitude="32.98836" municipality="Imatong Foothills" pageId="29" pageNumber="476" specimenCode="UTEP 22592, DFH 641" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFCAFFE2FC2247FEFBE3F82F" box="[992,1135,1923,1947]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="29" pageNumber="476" type="Museum">UTEP 22592</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFCAFFE2FB4247FEFB62F82F" box="[1152,1262,1923,1948]" collectionCode="DFH" pageId="29" pageNumber="476">DFH 641</specimenCode>
|
||
), female, collected at
|
||
<location id="8EC5EE98FFCAFFE2FC8747D9FC46F80E" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68:8EC5EE98FFCAFFE2FC8747D9FC46F80E" box="[837,970,1955,1979]" country="Uganda" county="Northern Region" latitude="3.80548" longLatPrecision="1" longitude="32.98836" municipality="Imatong Foothills" name="Agoro Town" pageId="29" pageNumber="476">Agoro Town</location>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFCAFFE2FC1447D9FB1CF80F" box="[982,1168,1955,1979]" pageId="29" pageNumber="476">Imatong Foothills</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62C4C0CFFFCAFFE2FB5E47D9FADEF80F" box="[1180,1362,1955,1979]" pageId="29" pageNumber="476">Northern Region</collectingCounty>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFCAFFE2FA9C47D9FA3CF80F" box="[1374,1456,1955,1979]" name="Uganda" pageId="29" pageNumber="476">Uganda</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFC9FFE1FF43457EFF7BFAA8" box="[129,247,1284,1308]" degrees="03.80548" direction="north" orientation="latitude" pageId="30" pageNumber="477" precision="1" value="3.80548">N03.80548</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFC9FFE1FEC0457EFEFFFAA8" box="[258,371,1284,1308]" degrees="32.98836" direction="east" orientation="longitude" pageId="30" pageNumber="477" precision="1" value="32.98836">E32.98836</geoCoordinate>
|
||
,
|
||
<elevation id="00375F70FFC9FFE1FEBC457EFD8CFAA8" box="[382,512,1284,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.153" pageId="30" pageNumber="477" unit="m" value="1153.0">
|
||
<quantity id="4CE215A6FFC9FFE1FEBC457EFE42FAAF" box="[382,462,1284,1308]" metricMagnitude="3" metricUnit="m" metricValue="1.153" pageId="30" pageNumber="477" unit="m" value="1153.0">1153 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFC9FFE1FDEB457EFD13FAA8" box="[553,671,1284,1308]" pageId="30" pageNumber="477" value="2015-07-05">
|
||
<collectingDate id="EFE0676BFFC9FFE1FDEB457EFD13FAA8" box="[553,671,1284,1308]" pageId="30" pageNumber="477" value="2015-07-05">5 July 2015</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFC9FFE1FD07457EFF62FA8F" pageId="30" pageNumber="477">Daniel F. Hughes</collectorName>
|
||
,
|
||
<collectorName id="26EFDD95FFC9FFE1FF394559FE26FA8F" box="[251,426,1315,1339]" pageId="30" pageNumber="477">Wilber Lukwago</collectorName>
|
||
, and
|
||
<collectorName id="26EFDD95FFC9FFE1FE2A4559FD34FA8F" box="[488,696,1315,1339]" pageId="30" pageNumber="477">Mathias Behangana</collectorName>
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B72B21EFFC9FFE1FD04455EFE6DFAEE" collectionCode="UTEP, DFH" country="Uganda" pageId="30" pageNumber="477" specimenCode="UTEP 22593, DFH 130" specimenCount="1" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFC9FFE1FD04455EFF4EFAEE" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="30" pageNumber="477" type="Museum">UTEP 22593</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFC9FFE1FF164538FECFFAEE" box="[212,323,1346,1370]" collectionCode="DFH" pageId="30" pageNumber="477">DFH 130</specimenCode>
|
||
), juvenile, and
|
||
</materialsCitation>
|
||
<materialsCitation id="3B72B21EFFC9FFE1FE2E4539FD39FA0C" collectingDate="2015-05-31" collectionCode="UTEP, DFH" collectorName="Daniel F. Hughes & Mathias Behangana" country="Uganda" elevation="1730" latitude="1.8227" location="Mount Kadam" longLatPrecision="1" longitude="34.74603" municipality="Northern Region" pageId="30" pageNumber="477" specimenCode="UTEP 22594, DFH 131" specimenCount="1" stateProvince="Nakapiripirit" typeStatus="paratype">
|
||
<specimenCode id="DBBC1038FFC9FFE1FE2E4539FDF7FAEE" box="[492,635,1346,1370]" collectionCode="UTEP" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="30" pageNumber="477" type="Museum">UTEP 22594</specimenCode>
|
||
(
|
||
<specimenCode id="DBBC1038FFC9FFE1FD4F4538FD71FAEE" box="[653,765,1346,1370]" collectionCode="DFH" pageId="30" pageNumber="477">DFH 131</specimenCode>
|
||
), female, collected at
|
||
<collectingRegion id="49DE76A1FFC9FFE1FE9D4518FE6AFACE" box="[351,486,1378,1402]" country="Uganda" name="Nakapiripirit" pageId="30" pageNumber="477">Nakapiripirit</collectingRegion>
|
||
,
|
||
<location id="8EC5EE98FFC9FFE1FE354518FD1BFACE" LSID="urn:lsid:plazi:treatment:03B30955FFCAFFDEFF1F46B0FD56FE68:8EC5EE98FFC9FFE1FE354518FD1BFACE" box="[503,663,1378,1402]" country="Uganda" latitude="1.8227" longLatPrecision="1" longitude="34.74603" municipality="Northern Region" name="Mount Kadam" pageId="30" pageNumber="477" stateProvince="Nakapiripirit">Mount Kadam</location>
|
||
,
|
||
<collectingMunicipality id="6BC12239FFC9FFE1FD6A4518FF41FA2D" pageId="30" pageNumber="477">Northern Region</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F30DF8D3FFC9FFE1FF1845FBFEA0FA2D" box="[218,300,1409,1433]" name="Uganda" pageId="30" pageNumber="477">Uganda</collectingCountry>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFC9FFE1FEFA45FBFE23FA2D" box="[312,431,1409,1433]" degrees="01.82270" direction="north" orientation="latitude" pageId="30" pageNumber="477" precision="1" value="1.8227">N01.82270</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE2EDE84FFC9FFE1FE7E45FBFDA1FA2D" box="[444,557,1409,1433]" degrees="34.74603" direction="east" orientation="longitude" pageId="30" pageNumber="477" precision="1" value="34.74603">E34.74603</geoCoordinate>
|
||
,
|
||
<elevation id="00375F70FFC9FFE1FDF845FBFD4EFA2D" box="[570,706,1409,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="30" pageNumber="477" unit="m" value="1730.0">
|
||
<quantity id="4CE215A6FFC9FFE1FDF845FBFD01FA2D" box="[570,653,1409,1433]" metricMagnitude="3" metricUnit="m" metricValue="1.73" pageId="30" pageNumber="477" unit="m" value="1730.0">1730 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
on
|
||
<date id="FFA49E83FFC9FFE1FD2D45FBFF6BFA0C" pageId="30" pageNumber="477" value="2015-05-31">
|
||
<collectingDate id="EFE0676BFFC9FFE1FD2D45FBFF6BFA0C" pageId="30" pageNumber="477" value="2015-05-31">31 May 2015</collectingDate>
|
||
</date>
|
||
by
|
||
<collectorName id="26EFDD95FFC9FFE1FEC845DAFE36FA0C" box="[266,442,1440,1464]" pageId="30" pageNumber="477">Daniel F. Hughes</collectorName>
|
||
and
|
||
<collectorName id="26EFDD95FFC9FFE1FE2945DAFD39FA0C" box="[491,693,1440,1464]" pageId="30" pageNumber="477">Mathias Behangana</collectorName>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF65E8CBFFC9FFE1FF434412FEC7FB60" ID-DOI="http://doi.org/10.5281/zenodo.13302899" ID-Zenodo-Dep="13302899" httpUri="https://zenodo.org/record/13302899/files/figure.png" pageId="30" pageNumber="477" startId="30.[129,194,1128,1152]" targetBox="[129,1473,144,1100]" targetPageId="30" targetType="figure">
|
||
<paragraph id="8BA5B843FFC9FFE1FF434412FEC7FB60" blockId="30.[129,1430,1127,1236]" pageId="30" pageNumber="477">
|
||
<emphasis id="B96E6451FFC9FFE1FF434412FF69FB34" bold="true" box="[129,229,1128,1152]" pageId="30" pageNumber="477">Figure 18.</emphasis>
|
||
Specimens of
|
||
<emphasis id="B96E6451FFC9FFE1FEAF441DFD0CFB34" bold="true" box="[365,640,1127,1152]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FEAF441DFDBCFBCB" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[365,560,1127,1151]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FEAF441DFDBCFBCB" bold="true" box="[365,560,1127,1151]" italics="true" pageId="30" pageNumber="477">Lygodactylus kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FDF74412FD0CFB34" box="[565,640,1128,1152]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life or freshly euthanized from (A) Mpishi (holotype: UTEP 22566), near Kibira National Park, Bubanza Province, Burundi, (B) N'Komo River (UTEP 22586), South Kivu Province, DRC, and (C) Bujumbura city, Bujumbura Mairie Province (UTEP 22576), Burundi. Photo of habitat at (D) Mpishi, Bubanza Province, Burundi. Photographs by E.G. (A–C) and C.K. (D).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C300EBC8FFC9FFE0FF43459AFE1CFF7C" lastPageId="31" lastPageNumber="478" pageId="30" pageNumber="477" type="diagnosis">
|
||
<paragraph id="8BA5B843FFC9FFE1FF43459AFDF5F8E4" blockId="30.[129,778,1503,1967]" pageId="30" pageNumber="477">
|
||
<emphasis id="B96E6451FFC9FFE1FF43459AFE63FA43" box="[129,495,1503,1527]" italics="true" pageId="30" pageNumber="477">
|
||
Diagnosis:
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FF3445A5FE63FA43" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[246,495,1503,1527]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FF3445A5FE63FA43" bold="true" box="[246,495,1503,1527]" pageId="30" pageNumber="477">Lygodactylus karamoja</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FE35459AFDC6FA4C" box="[503,586,1504,1528]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is the sister taxon of
|
||
<emphasis id="B96E6451FFC9FFE1FF594585FEDCF9A3" bold="true" box="[155,336,1535,1559]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FF594585FF75F9A3" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[155,249,1535,1559]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FF594585FF75F9A3" bold="true" box="[155,249,1535,1559]" italics="true" pageId="30" pageNumber="477">L. kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FF3C4585FEDCF9A3" box="[254,336,1535,1559]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
that is also a large-sized
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FD8B4585FD44F9A3" box="[585,712,1535,1559]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96E6451FFC9FFE1FD8B4585FD44F9A3" box="[585,712,1535,1559]" italics="true" pageId="30" pageNumber="477">Lygodactylus</emphasis>
|
||
</taxonomicName>
|
||
[maximum
|
||
<collectionCode id="ED0B2086FFC9FFE1FF054664FF78F982" box="[199,244,1566,1590]" pageId="30" pageNumber="477">SVL</collectionCode>
|
||
<quantity id="4CE215A6FFC9FFE1FF3C4664FED1F982" box="[254,349,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="30" pageNumber="477" unit="mm" value="37.7">37.7 mm</quantity>
|
||
(mean 34.4 ±
|
||
<quantity id="4CE215A6FFC9FFE1FE3D4664FDD4F982" box="[511,600,1566,1590]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="30" pageNumber="477" unit="mm" value="2.9">2.9 mm</quantity>
|
||
)] and shares the gular patterning of the
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FEBE4644FE79F9E2" box="[380,501,1598,1622]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFC9FFE1FEBE4644FE79F9E2" box="[380,501,1598,1622]" italics="true" pageId="30" pageNumber="477">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
subgroup. Seven to eight supralabials and 5–7 infralabials. Dorsal pholidosis with granular scales that become flattened, larger, and imbricate on original tails. Large triangular mental followed by usually two (or occasionally three) symmetric postmental scales (Supporting Information,
|
||
<figureCitation id="1321A4C6FFC9FFE1FECE46A0FED9F946" box="[268,341,1754,1778]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="30" pageNumber="477">Fig. S7</figureCitation>
|
||
). Ventral pholidosis with large, flattened, and imbricate scales. Ventral scales usually denticulated posteriorly. Six terminal scansors on the tail tip. Digits elongated with five terminal scansors on the fourth toe (
|
||
<tableCitation id="C6988DF8FFC9FFE1FDDC4742FDE4F8E4" box="[542,616,1848,1872]" captionStart="Table 3" captionStartId="12.[107,162,413,438]" captionTargetPageId="12" captionText="Table 3. Summary of external and osteological morphological data of all members within the L. gutturalis complex.Measurements are shown in mm (average and SD). For abbreviations see Material and methods section." httpUri="http://table.plazi.org/id/DF65E8CBFFDBFFF3FFA941E7FE07FE66" pageId="30" pageNumber="477" tableUuid="DF65E8CBFFDBFFF3FFA941E7FE07FE66">Table 3</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFC9FFE1FF5E4722FA4FFACE" blockId="30.[129,778,1503,1967]" lastBlockId="30.[825,1475,1283,1965]" pageId="30" pageNumber="477">
|
||
Like other
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FEDF4722FE10F8C4" box="[285,412,1880,1904]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="genus">
|
||
<emphasis id="B96E6451FFC9FFE1FEDF4722FE10F8C4" box="[285,412,1880,1904]" italics="true" pageId="30" pageNumber="477">Lygodactylus</emphasis>
|
||
</taxonomicName>
|
||
within the
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FDE84722FD09F8C4" box="[554,645,1880,1904]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFC9FFE1FDE84722FD09F8C4" box="[554,645,1880,1904]" italics="true" pageId="30" pageNumber="477">gutturalis</emphasis>
|
||
</taxonomicName>
|
||
group, this species can be easily differentiated from members of the
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FD374702FF57F81A" authorityName="Gunther" authorityYear="1893" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="angularis">
|
||
<emphasis id="B96E6451FFC9FFE1FD374702FF57F81A" italics="true" pageId="30" pageNumber="477">L. angularis</emphasis>
|
||
</taxonomicName>
|
||
group by its characteristic
|
||
<emphasis id="B96E6451FFC9FFE1FE2447ECFE7BF81A" bold="true" box="[486,503,1942,1966]" pageId="30" pageNumber="477">∩</emphasis>
|
||
<emphasis id="B96E6451FFC9FFE1FE3547EDFE72F81A" box="[503,510,1943,1966]" italics="true" pageId="30" pageNumber="477">-</emphasis>
|
||
shaped gular chevrons (vs. V-shaped gular chevrons in
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FB9A457EFB40FAAF" authorityName="Gunther" authorityYear="1893" box="[1112,1228,1283,1307]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="angularis">
|
||
<emphasis id="B96E6451FFC9FFE1FB9A457EFB40FAAF" box="[1112,1228,1283,1307]" italics="true" pageId="30" pageNumber="477">L. angularis</emphasis>
|
||
</taxonomicName>
|
||
group), and from other members of the
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FC27455EFBEEFA8F" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[997,1122,1316,1339]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="picturatus">
|
||
<emphasis id="B96E6451FFC9FFE1FC27455EFBEEFA8F" box="[997,1122,1316,1339]" italics="true" pageId="30" pageNumber="477">L. picturatus</emphasis>
|
||
</taxonomicName>
|
||
group based on dorsal coloration (light brown with five or six laterodorsal cream ocelli vs. usually blueish dorsum with yellow to white head in
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FB3A4518FAFEFACD" baseAuthorityName="Lobon-Rovira and Bauer" baseAuthorityYear="2021" box="[1272,1394,1378,1401]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="picturatus">
|
||
<emphasis id="B96E6451FFC9FFE1FB3A4518FAFEFACD" box="[1272,1394,1378,1401]" italics="true" pageId="30" pageNumber="477">L. picturatus</emphasis>
|
||
</taxonomicName>
|
||
group).
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFC9FFE0FC9745FAFE1CFF7C" blockId="30.[825,1475,1283,1965]" lastBlockId="31.[113,762,145,200]" lastPageId="31" lastPageNumber="478" pageId="30" pageNumber="477">
|
||
<emphasis id="B96E6451FFC9FFE1FC9745FAFB26FA2D" bold="true" box="[853,1194,1408,1433]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FC9745FAFBC2FA2C" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[853,1102,1408,1432]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FC9745FAFBC2FA2C" bold="true" box="[853,1102,1408,1432]" italics="true" pageId="30" pageNumber="477">Lygodactylus karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FB9445FBFB26FA2D" box="[1110,1194,1409,1433]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated from other species within the
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FB8D45DAFB26FA0C" box="[1103,1194,1440,1464]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFC9FFE1FB8D45DAFB26FA0C" box="[1103,1194,1440,1464]" italics="true" pageId="30" pageNumber="477">gutturalis</emphasis>
|
||
</taxonomicName>
|
||
subgroup by only minor morphometric and meristic data, reflecting its cryptic nature. However, we provide some characters that are putatively diagnostic.
|
||
<emphasis id="B96E6451FFC9FFE1FC404584FB5CF9A2" bold="true" box="[898,1232,1534,1558]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FC404584FBF5F9A2" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[898,1145,1534,1558]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FC404584FBF5F9A2" bold="true" box="[898,1145,1534,1558]" italics="true" pageId="30" pageNumber="477">Lygodactylus karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FBBC4584FB5CF9A2" box="[1150,1232,1534,1558]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FA904585FA4EF9A2" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1362,1474,1534,1558]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="dysmicus">
|
||
<emphasis id="B96E6451FFC9FFE1FA904585FA4EF9A2" box="[1362,1474,1534,1558]" italics="true" pageId="30" pageNumber="477">L. dysmicus</emphasis>
|
||
</taxonomicName>
|
||
by its larger size (maximum
|
||
<collectionCode id="ED0B2086FFC9FFE1FB9C4664FB07F982" box="[1118,1163,1566,1590]" pageId="30" pageNumber="477">SVL</collectionCode>
|
||
<quantity id="4CE215A6FFC9FFE1FB504664FB62F981" box="[1170,1262,1566,1590]" metricMagnitude="-2" metricUnit="m" metricValue="3.77" pageId="30" pageNumber="477" unit="mm" value="37.7">37.7 mm</quantity>
|
||
vs.
|
||
<quantity id="4CE215A6FFC9FFE1FAD44667FAFBF982" box="[1302,1399,1565,1590]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="30" pageNumber="477" unit="mm" value="27.6">27.6 mm</quantity>
|
||
); fewer precloacal pores (8 vs. 9); nostril not in contact with rostral scale (vs. nostril contacting rostral); and lower number of ventral scales across the body (16–18 vs.
|
||
<quantity id="4CE215A6FFC9FFE1FB004606FB7BF927" box="[1218,1271,1659,1684]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" pageId="30" pageNumber="477" unit="in" value="21.0">21 in</quantity>
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FB3C4606FAFCF927" baseAuthorityName="Perret" baseAuthorityYear="1963" box="[1278,1392,1659,1683]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="dysmicus">
|
||
<emphasis id="B96E6451FFC9FFE1FB3C4606FAFCF927" box="[1278,1392,1659,1683]" italics="true" pageId="30" pageNumber="477">L. dysmicus</emphasis>
|
||
</taxonomicName>
|
||
). It can be easily distinguished from
|
||
<emphasis id="B96E6451FFC9FFE1FB9D46E1FA99F907" bold="true" box="[1119,1301,1690,1715]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FB9D46E1FB32F906" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1119,1214,1690,1714]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FB9D46E1FB32F906" bold="true" box="[1119,1214,1690,1714]" italics="true" pageId="30" pageNumber="477">L. kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FB0146E1FA99F907" box="[1219,1301,1691,1715]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the gular pattern that comprises two or three thinner
|
||
<emphasis id="B96E6451FFC9FFE1FB3846C0FA87F966" bold="true" box="[1274,1291,1722,1746]" pageId="30" pageNumber="477">∩</emphasis>
|
||
-shaped chevrons that never extend beyond the posterior part of the lower jaw vs. three thick
|
||
<emphasis id="B96E6451FFC9FFE1FC6E4683FC31F8A5" bold="true" box="[940,957,1785,1809]" pageId="30" pageNumber="477">∩</emphasis>
|
||
-shaped chevrons reaching the chest; there are usually two symmetrical postmental scales vs. usually three smaller postmental scales; and a proportionally smaller orbital diameter (
|
||
<collectionCode id="ED0B2086FFC9FFE1FC83472DFCE7F8DB" box="[833,875,1879,1903]" pageId="30" pageNumber="477">OD</collectionCode>
|
||
/HL 0.19–0.23 vs.
|
||
<quantity id="4CE215A6FFC9FFE1FBF1472DFB39F8DB" box="[1075,1205,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="6.096" metricValueMax="6.604" metricValueMin="5.587999999999999" pageId="30" pageNumber="477" unit="in" value="0.24" valueMax="0.26" valueMin="0.22">0.22–0.26 in</quantity>
|
||
<emphasis id="B96E6451FFC9FFE1FB7F472DFAF4F8DB" bold="true" box="[1213,1400,1878,1903]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FB7F472DFA91F8DA" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1213,1309,1878,1902]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="kibera" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FB7F472DFA91F8DA" bold="true" box="[1213,1309,1878,1902]" italics="true" pageId="30" pageNumber="477">L. kibera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FAE6472DFAF4F8DB" box="[1316,1400,1879,1903]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). Also,
|
||
<emphasis id="B96E6451FFC9FFE1FCFB470CFB95F83A" bold="true" box="[825,1049,1910,1934]" pageId="30" pageNumber="477">
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FCFB470CFC31F83A" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[825,957,1910,1934]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC9FFE1FCFB470CFC31F83A" bold="true" box="[825,957,1910,1934]" italics="true" pageId="30" pageNumber="477">L. karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC9FFE1FC07470CFB95F83A" box="[965,1049,1910,1934]" pageId="30" pageNumber="477" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be differentiated in the same way as
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FCFB47ECFC22F819" box="[825,942,1941,1965]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="gutturalis">
|
||
<emphasis id="B96E6451FFC9FFE1FCFB47ECFC22F819" box="[825,942,1941,1965]" italics="true" pageId="30" pageNumber="477">L. gutturalis</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C1AC3C0FFC9FFE1FC2F47ECFBEDF819" authorityName=", Schmidt" authorityYear="1919" box="[1005,1121,1941,1965]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="477" phylum="Chordata" rank="species" species="depressus">
|
||
<emphasis id="B96E6451FFC9FFE1FC2F47ECFBEDF819" box="[1005,1121,1941,1965]" italics="true" pageId="30" pageNumber="477">L. depressus</emphasis>
|
||
</taxonomicName>
|
||
, based on gular patterning (always two or three
|
||
<emphasis id="B96E6451FFC8FFE0FF3940EBFE80FF1D" bold="true" box="[251,268,145,169]" pageId="31" pageNumber="478">∩</emphasis>
|
||
-shaped chevrons vs. one
|
||
<emphasis id="B96E6451FFC8FFE0FDDA40EBFDA5FF1D" bold="true" box="[536,553,145,169]" pageId="31" pageNumber="478">∩</emphasis>
|
||
-shaped followed by one posterior central mark).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300EBC8FFC8FFE0FFB34093FEECFE2A" pageId="31" pageNumber="478" type="etymology">
|
||
<paragraph id="8BA5B843FFC8FFE0FFB34093FEECFE2A" blockId="31.[113,762,233,414]" pageId="31" pageNumber="478">
|
||
<emphasis id="B96E6451FFC8FFE0FFB34093FF6DFEB5" box="[113,225,233,257]" italics="true" pageId="31" pageNumber="478">Etymology:</emphasis>
|
||
The name
|
||
<taxonomicName id="4C1AC3C0FFC8FFE0FEA64093FE4BFEB5" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[356,455,233,257]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="478" phylum="Chordata" rank="species" species="karamoja">
|
||
‘
|
||
<emphasis id="B96E6451FFC8FFE0FEAB4093FE4BFEB5" box="[361,455,233,257]" italics="true" pageId="31" pageNumber="478">karamoja</emphasis>
|
||
</taxonomicName>
|
||
’ is a noun in apposition and refers to the Karamoja region in north-eastern
|
||
<collectingCountry id="F30DF8D3FFC8FFE0FDA14173FD3EFE95" box="[611,690,265,289]" name="Uganda" pageId="31" pageNumber="478">Uganda</collectingCountry>
|
||
where many individuals of this species have been found. The species is named in honour of this arid region, which is occupied by the Karamojong people who are mostly nomadic pastoralists related to the Maasai in
|
||
<collectingCountry id="F30DF8D3FFC8FFE0FEDA41FCFED0FE2A" box="[280,348,390,414]" name="Kenya" pageId="31" pageNumber="478">Kenya</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300EBC8FFC8FFDEFFB341BAFE7BFE91" lastPageId="33" lastPageNumber="480" pageId="31" pageNumber="478" type="description">
|
||
<paragraph id="8BA5B843FFC8FFE0FFB341BAFBCCFD08" blockId="31.[113,763,447,847]" lastBlockId="31.[810,1459,144,700]" pageId="31" pageNumber="478">
|
||
<emphasis id="B96E6451FFC8FFE0FFB341BAFE1EFE63" box="[113,402,447,471]" italics="true" pageId="31" pageNumber="478">
|
||
Description of the
|
||
<typeStatus id="54A106E1FFC8FFE0FE8241C5FE1EFE63" box="[320,402,447,471]" pageId="31" pageNumber="478" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFC8FFE0FE7E41C5FD80FE63" box="[444,524,447,471]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302905" httpUri="https://zenodo.org/record/13302905/files/figure.png" pageId="31" pageNumber="478">Fig. 19</figureCitation>
|
||
): Measurements and meristic characters of the
|
||
<typeStatus id="54A106E1FFC8FFE0FEB841A4FE5AFE42" box="[378,470,478,502]" pageId="31" pageNumber="478" type="holotype">holotype</typeStatus>
|
||
are presented in Supporting Information, Table S8. Adult male, with a snout–vent length (
|
||
<collectionCode id="ED0B2086FFC8FFE0FFBB4267FF20FD81" box="[121,172,541,565]" pageId="31" pageNumber="478">SVL</collectionCode>
|
||
) of
|
||
<quantity id="4CE215A6FFC8FFE0FF1F4267FEB2FD81" box="[221,318,541,565]" metricMagnitude="-2" metricUnit="m" metricValue="3.4299999999999997" pageId="31" pageNumber="478" unit="mm" value="34.3">34.3 mm</quantity>
|
||
and an original tail length slightly larger (TL =
|
||
<quantity id="4CE215A6FFC8FFE0FF7B4246FEAFFDE1" box="[185,291,572,597]" metricMagnitude="-2" metricUnit="m" metricValue="3.61" pageId="31" pageNumber="478" unit="mm" value="36.1">36.1 mm</quantity>
|
||
). Body slender, nape moderately distinct. Head as broad as body and moderate head length (
|
||
<collectionCode id="ED0B2086FFC8FFE0FD584226FD46FDC0" box="[666,714,604,628]" pageId="31" pageNumber="478">HW</collectionCode>
|
||
/HL 0.72). Canthus rostralis not prominent. Eye diameter
|
||
<quantity id="4CE215A6FFC8FFE0FD6B4201FD76FD27" box="[681,762,635,659]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="31" pageNumber="478" unit="mm" value="1.9">1.9 mm</quantity>
|
||
with circular pupil. Ear to eye distance twice the orbital diameter (
|
||
<quantity id="4CE215A6FFC8FFE0FF6B42C0FE8FFD66" box="[169,259,698,722]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="31" pageNumber="478" unit="mm" value="3.2">3.2 mm</quantity>
|
||
). Snout rounded and slightly pointed. Granular scales on frontal area larger than occipital scales, with 22 small granular scales between the eyes. Dorsal scales granular from rostral to tail. Rostral undivided, in contact with 1st supralabial, prenasals, and one large internasal scale. Eight supralabials and 5–6 infralabials. Prenasal scale present and in contact with 1st supralabial. Nostril circular, bordered by 1st supralabial, prenasal, and one supranasal. Postnasal not contacting nostril, separated by 1st supralabial, which it contacts posteriorly. Four rows of scales between supralabials and orbit. Mental large, triangular, and rounded, bordered posteriorly by two large rounded symmetric postmental scales. Six asymmetrical post-postmental scales. Gular scales granular, rounded, and slightly smaller than ventral scales. Ventral scales imbricate, rounded, and larger than dorsal scales, with 16 scales rows across the venter. Body relatively robust and slightly elongated (
|
||
<collectionCode id="ED0B2086FFC8FFE0FB5C41B3FB5EFE55" box="[1182,1234,457,481]" pageId="31" pageNumber="478">TRL</collectionCode>
|
||
/
|
||
<collectionCode id="ED0B2086FFC8FFE0FB1941B3FA82FE55" box="[1243,1294,457,481]" pageId="31" pageNumber="478">SVL</collectionCode>
|
||
0.42). Tail with 34 enlarged transverse scales and six pairs of terminal scansors at the tip. Eight precloacal pores. Fore- and hind limbs moderately short, stout; forearm medium-sized (
|
||
<collectionCode id="ED0B2086FFC8FFE0FB1F425DFB71FD8B" box="[1245,1277,551,575]" pageId="31" pageNumber="478">FL</collectionCode>
|
||
/
|
||
<collectionCode id="ED0B2086FFC8FFE0FAC4425DFAB4FD8B" box="[1286,1336,551,575]" pageId="31" pageNumber="478">SVL</collectionCode>
|
||
0.16); tibia short (CL/
|
||
<collectionCode id="ED0B2086FFC8FFE0FC55423DFC46FDEB" box="[919,970,583,607]" pageId="31" pageNumber="478">SVL</collectionCode>
|
||
0.17). Digits elongated and unwebbed with 5–6 terminal scansors. Thumb rudimentary with a small terminal claw. Relative length of digits: I <II =
|
||
<collectionCode id="ED0B2086FFC8FFE0FB0942FCFB50FD29" box="[1227,1244,646,669]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="478" type="Museum">V</collectionCode>
|
||
<III <IV (manus); I <II <
|
||
<collectionCode id="ED0B2086FFC8FFE0FCBC42DFFC03FD08" box="[894,911,677,700]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="31" pageNumber="478" type="Museum">V</collectionCode>
|
||
<III <IV (pes).
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFC8FFDFFCE842A4FDE8FEF1" blockId="31.[810,1459,734,852]" lastBlockId="32.[129,778,144,325]" lastPageId="32" lastPageNumber="479" pageId="31" pageNumber="478">
|
||
<emphasis id="B96E6451FFC8FFE0FCE842A4FC60FD42" box="[810,1004,734,758]" italics="true" pageId="31" pageNumber="478">Coloration: In life</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFC8FFE0FBC142A4FBC2FD42" box="[1027,1102,734,758]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="31" pageNumber="478">Fig. 20</figureCitation>
|
||
), dorsal coloration from light to dark brown to greyish or almost black (
|
||
<figureCitation id="1321A4C6FFC8FFE0FB0B4287FAAFFCA1" box="[1225,1315,765,789]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="31" pageNumber="478">Fig. 20D</figureCitation>
|
||
), with lighter cream-beige lines with five or six lighter cream dots surrounded by black flanks. Some specimens have lighter cream-beige dots along the vertebral line, giving them a granitic appearance (
|
||
<figureCitation id="1321A4C6FFF7FFDFFD2140EAFF27FF73" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="32" pageNumber="479">Fig. 20A</figureCitation>
|
||
). Black line from nostril to the anterior insertion of the forelimb. Gular region white and two or three black,
|
||
<emphasis id="B96E6451FFF7FFDFFDAE40B5FDF1FF53" bold="true" box="[620,637,207,231]" pageId="32" pageNumber="479">∩</emphasis>
|
||
-shaped chevrons. Venter uniformly orange or white. Tip of tail and digits dark brown.
|
||
<emphasis id="B96E6451FFF7FFDFFF124174FED2FE91" box="[208,350,270,293]" italics="true" pageId="32" pageNumber="479">In preservative</emphasis>
|
||
(
|
||
<typeStatus id="54A106E1FFF7FFDFFEAE4177FE5CFE91" box="[364,464,269,293]" pageId="32" pageNumber="479" type="holotype">holotype</typeStatus>
|
||
;
|
||
<figureCitation id="1321A4C6FFF7FFDFFE194177FDAEFE91" box="[475,546,269,293]" captionStart="Figure 19" captionStartId="31.[115,180,1898,1922]" captionTargetBox="[114,1458,920,1870]" captionTargetId="figure-423@31.[114,1458,920,1870]" captionTargetPageId="31" captionText="Figure 19. Holotype of L. karamojasp. nov. (UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda.A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302905" httpUri="https://zenodo.org/record/13302905/files/figure.png" pageId="32" pageNumber="479">Fig. 19</figureCitation>
|
||
): dorsum dark brown and venter with uniform light cream coloration.
|
||
</paragraph>
|
||
<caption id="DF65E8CBFFC8FFE0FFB14710FDDAF80E" ID-DOI="http://doi.org/10.5281/zenodo.13302905" ID-Zenodo-Dep="13302905" httpUri="https://zenodo.org/record/13302905/files/figure.png" pageId="31" pageNumber="478" startId="31.[115,180,1898,1922]" targetBox="[114,1458,920,1870]" targetPageId="31" targetType="figure">
|
||
<paragraph id="8BA5B843FFC8FFE0FFB14710FDDAF80E" blockId="31.[113,1443,1898,1979]" pageId="31" pageNumber="478">
|
||
<emphasis id="B96E6451FFC8FFE0FFB14710FF54F837" bold="true" box="[115,216,1898,1923]" pageId="31" pageNumber="478">Figure 19.</emphasis>
|
||
Holotype of
|
||
<emphasis id="B96E6451FFC8FFE0FE964711FD96F837" bold="true" box="[340,538,1898,1923]" pageId="31" pageNumber="478">
|
||
<taxonomicName id="4C1AC3C0FFC8FFE0FE964711FE40F836" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[340,460,1898,1922]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="478" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFC8FFE0FE964711FE40F836" bold="true" box="[340,460,1898,1922]" italics="true" pageId="31" pageNumber="478">L. karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFC8FFE0FE0D4711FD96F837" box="[463,538,1899,1923]" pageId="31" pageNumber="478" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(UTEP 22590) from Agoro Town, Imatong Foothills, Northern Region, Uganda. A, Dorsal and ventral view of body, (B) details of cloacal section, (C) regenerated tail tip, and (D) left pes. E, Detailed view of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BA5B843FFF7FFDFFF43411DFAEAFF53" blockId="32.[129,778,358,413]" lastBlockId="32.[825,1474,144,231]" pageId="32" pageNumber="479">
|
||
<emphasis id="B96E6451FFF7FFDFFF43411DFF68FECA" box="[129,228,359,382]" italics="true" pageId="32" pageNumber="479">Variation:</emphasis>
|
||
The meristic and morphometric data are summarized in Supporting Information, Table S8. Coloration of this species seems to be consistent, with gular coloration slightly lighter in females. Number and shape of postmentals and internasals vary between specimens (Supporting Information,
|
||
<figureCitation id="1321A4C6FFF7FFDFFAD240B5FADAFF53" box="[1296,1366,207,231]" captionStart="Figure 7" captionStartId="18.[129,194,1068,1092]" captionTargetBox="[129,1473,144,1040]" captionTargetId="figure-493@18.[129,1473,144,1040]" captionTargetPageId="18" captionText="Figure 7. Holotype of L. heeneni [RMCA (= MRAC) R.8477] from Kapiri, Lualaba Province, DRC. A, Ventral and dorsal view of body. B, Details of head in dorsal, ventral, and lateral views (from top to bottom). Photos by J.L.R." figureDoi="http://doi.org/10.5281/zenodo.13302858" httpUri="https://zenodo.org/record/13302858/files/figure.png" pageId="32" pageNumber="479">Fig. S7</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BA5B843FFF7FFDEFCFB4172FE7BFE91" blockId="32.[825,1474,263,414]" lastBlockId="33.[113,762,144,293]" lastPageId="33" lastPageNumber="480" pageId="32" pageNumber="479">
|
||
<emphasis id="B96E6451FFF7FFDFFCFB4172FBA1FE94" box="[825,1069,264,288]" italics="true" pageId="32" pageNumber="479">Habitat and distribution</emphasis>
|
||
(
|
||
<figureCitation id="1321A4C6FFF7FFDFFB8A4172FB0BFE94" box="[1096,1159,264,288]" captionStart="Figure 6" captionStartId="17.[113,178,866,890]" captionTargetBox="[117,1454,147,835]" captionTargetId="figure-489@17.[114,1458,144,838]" captionTargetPageId="17" captionText="Figure 6. Geographical distribution of the L.gutturalis complex in Africa, on a hybrid map of the major vegetation divisions (Sayre et al. 2013) and 1 arc-second elevation map across tropical Africa (NASA 2000). Different colours depict records of different species within the L. gutturalis subgroup; see inset for explanations of symbols.White circles denote records of L. cf. gutturalis that cannot be assigned to any of the nominate taxa. Black stars represent the type localities of each nominate taxon within the L. picturatus subgroup (1– L. picturatus, 1S– L. picturatus sudanensis, 1U– L. picturatus ukerewensis, 2– L. inexpectatus, 3– L. keniensis, 4– L. kimhowelli, 5– L. manni, 6– L. mombasicus, 7– L. scorteccii, 8– L. tsavoensis, 9– L. wojnowskii, 10– L. williamsi)." figureDoi="http://doi.org/10.5281/zenodo.13302854" httpUri="https://zenodo.org/record/13302854/files/figure.png" pageId="32" pageNumber="479">Figs 6</figureCitation>
|
||
,
|
||
<figureCitation id="1321A4C6FFF7FFDFFB564172FB22FE94" box="[1172,1198,264,288]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="32" pageNumber="479">20</figureCitation>
|
||
<emphasis id="B96E6451FFF7FFDFFB6C4172FA4EFEAB" box="[1198,1474,263,287]" italics="true" pageId="32" pageNumber="479">
|
||
):
|
||
<taxonomicName id="4C1AC3C0FFF7FFDFFB0B417DFA4EFEAB" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[1225,1474,263,287]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="479" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF7FFDFFB0B417DFA4EFEAB" bold="true" box="[1225,1474,263,287]" pageId="32" pageNumber="479">Lygodactylus karamoja</emphasis>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
<taxonomicNameLabel id="A25DD92AFFF7FFDFFCFB415DFC1CFE8B" box="[825,912,295,319]" pageId="32" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
|
||
has been recorded from mid-elevation savannahs of the northern Albertine Rift to the Karamoja Region, eastern
|
||
<collectingCountry id="F30DF8D3FFF7FFDFFCFB411CFC07FECA" box="[825,907,358,382]" name="Uganda" pageId="32" pageNumber="479">Uganda</collectingCountry>
|
||
, an arid area characterized by mostly woodland savannah habitat with several isolated mountain ranges. This suggests that specimens from western
|
||
<collectingCountry id="F30DF8D3FFF6FFDEFEB340EAFE3EFF1C" box="[369,434,144,168]" name="Kenya" pageId="33" pageNumber="480">Kenya</collectingCountry>
|
||
and southern
|
||
<collectingCountry id="F30DF8D3FFF6FFDEFD8540EAFD47FF1C" box="[583,715,144,168]" name="South Sudan" pageId="33" pageNumber="480">South Sudan</collectingCountry>
|
||
also belong to this species. If confirmed, this species would have a distribution that is similar to the lacertid
|
||
<taxonomicName id="4C1AC3C0FFF6FFDEFD9740B5FE0CFEB2" authority="(Greenbaum et al. 2018)" baseAuthorityName="Greenbaum" baseAuthorityYear="2018" class="Reptilia" family="Lacertidae" genus="Adolfus" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="480" phylum="Chordata" rank="species" species="jacksoni">
|
||
<emphasis id="B96E6451FFF6FFDEFD9740B5FD76FF53" box="[597,762,207,231]" italics="true" pageId="33" pageNumber="480">Adolfus jacksoni</emphasis>
|
||
(Greenbaum
|
||
<emphasis id="B96E6451FFF6FFDEFEC34095FEBAFEB2" box="[257,310,238,262]" italics="true" pageId="33" pageNumber="480">et al.</emphasis>
|
||
2018)
|
||
</taxonomicName>
|
||
. However, we recommend caution until molecular data become available.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF65E8CBFFF7FFDFFF424710FDD7F80E" ID-DOI="http://doi.org/10.5281/zenodo.13302907" ID-Zenodo-Dep="13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="32" pageNumber="479" startId="32.[128,193,1898,1922]" targetBox="[129,1473,449,1870]" targetPageId="32" targetType="figure">
|
||
<paragraph id="8BA5B843FFF7FFDFFF424710FDD7F80E" blockId="32.[128,1444,1898,1979]" pageId="32" pageNumber="479">
|
||
<emphasis id="B96E6451FFF7FFDFFF424710FF69F837" bold="true" box="[128,229,1898,1923]" pageId="32" pageNumber="479">Figure 20.</emphasis>
|
||
Specimens of
|
||
<emphasis id="B96E6451FFF7FFDFFEAE4711FDBFF837" bold="true" box="[364,563,1898,1923]" pageId="32" pageNumber="479">
|
||
<taxonomicName id="4C1AC3C0FFF7FFDFFEAE4711FE68F836" authorityName="Lobón-Rovira & Bauer & Pinto & Trape & Conradie & Kusamba & Júlio & Cael & Stanley & Hughes & Behangana & Masudi & Pauwels & Greenbaum" authorityYear="2024" box="[364,484,1898,1922]" class="Reptilia" family="Gekkonidae" genus="Lygodactylus" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="479" phylum="Chordata" rank="species" species="karamoja" status="sp. nov.">
|
||
<emphasis id="B96E6451FFF7FFDFFEAE4711FE68F836" bold="true" box="[364,484,1898,1922]" italics="true" pageId="32" pageNumber="479">L. karamoja</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25DD92AFFF7FFDFFE2A4711FDBFF837" box="[488,563,1899,1923]" pageId="32" pageNumber="479" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda. Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C300EBC8FFF6FFDEFFB3413DFD56FE68" pageId="33" pageNumber="480" type="biology_ecology">
|
||
<paragraph id="8BA5B843FFF6FFDEFFB3413DFD56FE68" blockId="33.[113,762,327,476]" pageId="33" pageNumber="480">
|
||
<emphasis id="B96E6451FFF6FFDEFFB3413DFE9DFEEA" box="[113,273,327,351]" italics="true" pageId="33" pageNumber="480">Natural history:</emphasis>
|
||
A diurnal and arboreal species that is always found on trees.
|
||
<collectionCode id="ED0B2086FFF6FFDEFED3411CFED8FEC9" box="[273,340,358,381]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="33" pageNumber="480" type="Museum">UTEP</collectionCode>
|
||
22592, a gravid female, was found in July on a tree at night.
|
||
<collectionCode id="ED0B2086FFF6FFDEFEE341FCFEE8FE29" box="[289,356,390,413]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15719" lsid="urn:lsid:biocol.org:col:15719" name="University of Texas at El Paso Biodiversity Collections" pageId="33" pageNumber="480" type="Museum">UTEP</collectionCode>
|
||
22593–94 were found during the day on trees trunks
|
||
<emphasis id="B96E6451FFF6FFDEFF2C41DFFF7BFE08" box="[238,247,421,444]" italics="true" pageId="33" pageNumber="480">c</emphasis>
|
||
.
|
||
<quantity id="4CE215A6FFF6FFDEFEC041DFFEA5FE08" box="[258,297,421,445]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="480" unit="m" value="1.0">1 m</quantity>
|
||
above ground in an open clearing of grass and rocky outcrops that were surrounded by dry forest (
|
||
<figureCitation id="1321A4C6FFF6FFDEFDB541BEFD45FE68" box="[631,713,452,476]" captionStart="Figure 20" captionStartId="32.[128,193,1898,1922]" captionTargetBox="[129,1473,449,1870]" captionTargetId="figure-170@32.[129,1473,449,1870]" captionTargetPageId="32" captionText="Figure 20. Specimens of L. karamoja sp. nov. in life from (A–B) Agoro Town (holotype: UTEP 22590), Imatong Foothills and (D–E) Nakapiripirit (UTEP 22594), Mount Kadam, Northern Region, Uganda.Photo of habitat at (C) Imatong Foothills, and (F) Mount Kadam, Northern Region, Uganda. Photographs by D.F.H." figureDoi="http://doi.org/10.5281/zenodo.13302907" httpUri="https://zenodo.org/record/13302907/files/figure.png" pageId="33" pageNumber="480">Fig. 20F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |