366 lines
31 KiB
XML
366 lines
31 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1058.68628" ID-GBIF-Dataset="4477866e-bbd0-4938-86ed-bd708b00559c" ID-PMC="PMC8421324" ID-Pensoft-Pub="1313-2970-1058-1" ID-Pensoft-UUID="61FB9E86BEE45968AAC868B21AF7ED86" ID-PubMed="34557053" ID-ZooBank="654932353DDB4E1B8848EAB69F2C20FD" ModsDocID="1313-2970-1058-1" checkinTime="1630349293316" checkinUser="pensoft" docAuthor="Vagalinski, Boyan & Golovatch, Sergei I." docDate="2021" docId="C891E4EE1E0E5AD4BDDA2959F415A530" docLanguage="en" docName="ZooKeys 1058: 1-127" docOrigin="ZooKeys 1058" docPubDate="2021-08-30" docSource="http://dx.doi.org/10.3897/zookeys.1058.68628" docTitle="Svaniulus waltheri Vagalinski, gen. nov., sp. nov." docType="treatment" docVersion="5" id="61FB9E86BEE45968AAC868B21AF7ED86" lastPageNumber="1" masterDocId="61FB9E86BEE45968AAC868B21AF7ED86" masterDocTitle="The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae)" masterLastPageNumber="127" masterPageNumber="1" pageNumber="1" updateTime="1668150880892" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The millipede tribe Brachyiulini in the Caucasus (Diplopoda, Julida, Julidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Vagalinski, Boyan</mods:namePart>
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<mods:affiliation>Institute of Biodiversity and Ecosystem Research at the Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria</mods:affiliation>
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<mods:nameIdentifier type="email">boyan_vagalinski@excite.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation>Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-08-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1058</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>127</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1058.68628</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1058.68628</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1058-1</mods:identifier>
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<mods:identifier type="ZooBank">654932353DDB4E1B8848EAB69F2C20FD</mods:identifier>
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||
<mods:identifier type="Pensoft-UUID">61FB9E86BEE45968AAC868B21AF7ED86</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7019436" ID-GBIF-Taxon="185371656" ID-Zenodo-Dep="7019436" LSID="urn:lsid:zoobank.org:act:15B28D68-1A07-49AA-9F81-717705C8F0AD" httpUri="http://treatment.plazi.org/id/C891E4EE1E0E5AD4BDDA2959F415A530" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/15B28D68-1A07-49AA-9F81-717705C8F0AD" authority="Vagalinski, gen. nov., sp. nov." authorityName="Vagalinski, gen. nov., sp. nov." class="Diplopoda" family="Julidae" genus="Svaniulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Svaniulus waltheri" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="waltheri">Svaniulus waltheri Vagalinski, gen. nov., sp. nov.</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Svaniulus waltheri gen. nov., sp. nov., ♂ paratype A head and body rings 1 - 3 B telson, lateral views. Not to scale." figureDoi="10.3897/zookeys.1058.68628.figure50" httpUri="https://binary.pensoft.net/fig/581985" pageId="0" pageNumber="1">Figs 50</figureCitation>
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<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">, 51</figureCitation>
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<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Svaniulus waltheri gen. nov., sp. nov., gonopods of ♂ paratype A right promere, caudal view B apical part of flagellum C right opisthomere, oral view D distal part of the same, latero-oral view E distal part of left opisthomere, caudo-mesal view F apical part of the same, meso-caudal view. Scale bars: 0.1 mm (A, C-E), 0.05 mm (F), 0.01 mm (B). Abbreviations: bpp basoposterior process, dr distal ridge, f flagellum, g (supposed) gonocoxal gland, ml mesomeroidal lobe, mp mesal process, mr median ridge, msl mesal lobe of basoposterior process, s solenomere, sf spiniform filaments, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure52" httpUri="https://binary.pensoft.net/fig/581987" pageId="0" pageNumber="1">, 52</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Material examined</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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(SMNG):
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<materialsCitation collectingDate="2011-10-10" collectorName="F. Walther" country="Georgia" county="Lebarde" elevation="1260" latitude="42.7258" location="Taleri" longLatPrecision="6" longitude="42.4703" municipality="Tekhuri Valley" specimenCount="♂" stateProvince="Samegrelo-Zemo Svaneti" typeStatus="Holotype">
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<emphasis italics="true" pageId="0" pageNumber="1">
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<typeStatus>Holotype</typeStatus>
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</emphasis>
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:
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<specimenCount>♂</specimenCount>
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(in head to pleurotergum 7 and rest of body, gonopods intact),
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<collectingCountry name="Georgia">Georgia</collectingCountry>
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:
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<collectingRegion country="Georgia" name="Samegrelo and Zemo Svaneti">Samegrelo-Zemo Svaneti</collectingRegion>
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,
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<collectingCounty>Lebarde</collectingCounty>
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,
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<collectingMunicipality>Tekhuri Valley</collectingMunicipality>
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, 30 rkm from
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<location LSID="urn:lsid:plazi:treatment:C891E4EE1E0E5AD4BDDA2959F415A530:3C98EFFA08AE2FCEA56F6C10D3A69B81" country="Georgia" county="Lebarde" latitude="42.7258" longLatPrecision="6" longitude="42.4703" municipality="Tekhuri Valley" name="Taleri" stateProvince="Samegrelo-Zemo Svaneti">Taleri</location>
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towards
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<location LSID="urn:lsid:plazi:treatment:C891E4EE1E0E5AD4BDDA2959F415A530:C13644A59D4C8423108F6B13F993F900" country="Georgia" county="Lebarde" latitude="42.7258" longLatPrecision="6" longitude="42.4703" municipality="Tekhuri Valley" name="Lebarde" stateProvince="Samegrelo-Zemo Svaneti">Lebarde</location>
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[at the influx of Lebarde and Tekhuri rivers],
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<geoCoordinate degrees="42.7258" direction="north" orientation="latitude" precision="5" value="42.7258">42.7258°N</geoCoordinate>
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,
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<geoCoordinate degrees="42.4703" direction="east" orientation="longitude" precision="5" value="42.4703">42.4703°E</geoCoordinate>
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,
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<elevation metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">
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<quantity metricMagnitude="3" metricUnit="m" metricValue="1.26" unit="m" value="1260.0">1260 m</quantity>
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a.s.l.
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</elevation>
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,
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<collectingDate value="2011-10-10">10.X.2011</collectingDate>
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,
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<collectorName>F. Walther</collectorName>
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leg.
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</materialsCitation>
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</emphasis>
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">
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<typeStatus>Paratypes</typeStatus>
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</emphasis>
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:
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<specimenCount type="male">1 ♂</specimenCount>
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(in 3 pieces, leg 7, mid-body and end-body leg, hypoproct, left flange of pleurotergum 7, and penis dissected, gonopods prepared for SEM),
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<specimenCount type="female">1 ♀</specimenCount>
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(in head to body ring 3 and rest of body, vulvae dissected, right one prepared for SEM), same collecting data as of the
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<typeStatus>holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Differs from its only known congener,
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., mostly by being on average slightly larger, by the relatively more slender male antennae, by the position and shape of the ventral protrusions of male pleurotergum 7, and by the following details of gonopod structure: promere more slender, distally significantly narrowing, with the mesal process barely exceeding the apex, vs. the same being somewhat stouter, only apically abruptly narrowing, and with the mesal process significantly exceeding the apex in
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov.; opisthomere lacking a distomesal process, vs. this being present in
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., having the basoposterior process with a distinct apical corner, vs. the same ending in a broad and flat margin in
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov. sp. nov., and with the solenomere being somewhat enlarged apically, vs. the same narrowing all the way to the top in
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="name">
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<paragraph pageId="0" pageNumber="1">Name.</paragraph>
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<paragraph pageId="0" pageNumber="1">Honours Frank Walther, a malacologist and an active collector of various invertebrate groups, including the material used for the present description.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Measurements</emphasis>
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:
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</emphasis>
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holotype in S X, 43+1+T, L = 26 mm, H = 1.8 mm; paratype ♂ in S IX, 43+1+T, L = 27.5 mm, H = 1.75 mm; paratype ♀ in S IX, 44+1+T, L = 37.5 mm, H = 2.65 mm.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Colouration</emphasis>
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(apparently somewhat faded) (Fig.
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<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Svaniulus waltheri gen. nov., sp. nov., ♂ paratype A head and body rings 1 - 3 B telson, lateral views. Not to scale." figureDoi="10.3897/zookeys.1058.68628.figure50" httpUri="https://binary.pensoft.net/fig/581985" pageId="0" pageNumber="1">50</figureCitation>
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): Similar to
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., but generally lighter, predominantly grey rather than brown, without significant difference between dorsal and ventral sides, and without transverse dark stripes along pro-metazonal sutures; prozonae dark grey, with numerous, densely set, light brown-beige spots arranged transversely above and below ozopore level; metazonae mostly grey, with numerous, minute, dark brown/blackish dots throughout, dorsally with a very narrow, transverse, blurred, dark brown band in posterior section; hind margins ochre-brown.
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</paragraph>
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<caption doi="10.3897/zookeys.1058.68628.figure50" httpUri="https://binary.pensoft.net/fig/581985" pageId="0" pageNumber="1" start="Figure 50" startId="F50">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 50.</emphasis>
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<taxonomicName class="Diplopoda" family="Julidae" genus="Svaniulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Svaniulus waltheri" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="waltheri">
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<emphasis italics="true" pageId="0" pageNumber="1">Svaniulus waltheri</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., ♂ paratype
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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head and body rings 1-3
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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telson, lateral views. Not to scale.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">External structures</emphasis>
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:
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</emphasis>
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Eye patches in adults vertically rather elongate, due to partial reduction of the higher developmental (vertical) rows, consisting of 21-35 markedly convex ommatidia. Vertigial, supralabral, and labral setae: two, four, and 16-18, respectively. Antennae (Fig.
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<figureCitation captionStart="Figure 50" captionStartId="F50" captionText="Figure 50. Svaniulus waltheri gen. nov., sp. nov., ♂ paratype A head and body rings 1 - 3 B telson, lateral views. Not to scale." figureDoi="10.3897/zookeys.1058.68628.figure50" httpUri="https://binary.pensoft.net/fig/581985" pageId="0" pageNumber="1">50A</figureCitation>
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) 1.6-1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as head in the males, and 1.25
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<normalizedToken originalValue="×">x</normalizedToken>
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in the female; antennomere 2> 5 ≥ 3> 4> 6. Gnathochilarium as in
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., but lamellae linguales with three rather than four setae in a longitudinal row, and male stipites without setose hump.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Body rings slightly to moderately vaulted. Male hypoproct (Fig.
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<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">51A</figureCitation>
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) very similar to that in
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<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
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<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., but almost perfectly semi-circular and with a densely and bluntly serrate margin, rather than apically somewhat flattened and with only several faint undulations. Tarsus of mid-body legs subequal to, to slightly longer than, tibia, and ca. 3
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as apical claw.
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</paragraph>
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<caption doi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1" start="Figure 51" startId="F51">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 51.</emphasis>
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<taxonomicName class="Diplopoda" family="Julidae" genus="Svaniulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Svaniulus waltheri" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="waltheri">
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<emphasis italics="true" pageId="0" pageNumber="1">Svaniulus waltheri</emphasis>
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</taxonomicName>
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gen. nov., sp. nov., ♂ (
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<emphasis bold="true" pageId="0" pageNumber="1">A-D</emphasis>
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) and ♀ (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) paratypes
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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hypoproct, ventral view
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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right flange of pleurotergum 7, ventro-lateral view
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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penis, caudal view
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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right opisthomere, oral view
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations:
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ct</emphasis>
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central tube,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">g</emphasis>
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(supposed) gonocoxal gland,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ml</emphasis>
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mesomeroidal lobe,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">msl</emphasis>
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mesal lobe of basoposterior process,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">pt</emphasis>
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posterior tube,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">s</emphasis>
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solenomere,
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">sp</emphasis>
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apical process of solenomere.
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</paragraph>
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</caption>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Male sexual characters</emphasis>
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:
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</emphasis>
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Walking legs with adhesive pads still present in caudalmost pairs. Pleurotergum 7 ventrally forming rather narrow and oar-like lobes (Fig.
|
||
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">51B</figureCitation>
|
||
) with a slightly concave apical margin, originating from border zone between pro- and metazona, protruding mostly ventrad behind gonopods. Penis (Fig.
|
||
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">51C</figureCitation>
|
||
) similar to that in
|
||
<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., but trapezoidal (widening distally) rather than barrel-shaped.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
All other external somatic characters as in
|
||
<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">51D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Svaniulus waltheri gen. nov., sp. nov., gonopods of ♂ paratype A right promere, caudal view B apical part of flagellum C right opisthomere, oral view D distal part of the same, latero-oral view E distal part of left opisthomere, caudo-mesal view F apical part of the same, meso-caudal view. Scale bars: 0.1 mm (A, C-E), 0.05 mm (F), 0.01 mm (B). Abbreviations: bpp basoposterior process, dr distal ridge, f flagellum, g (supposed) gonocoxal gland, ml mesomeroidal lobe, mp mesal process, mr median ridge, msl mesal lobe of basoposterior process, s solenomere, sf spiniform filaments, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure52" httpUri="https://binary.pensoft.net/fig/581987" pageId="0" pageNumber="1">52</figureCitation>
|
||
): Promere (Fig.
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Svaniulus waltheri gen. nov., sp. nov., gonopods of ♂ paratype A right promere, caudal view B apical part of flagellum C right opisthomere, oral view D distal part of the same, latero-oral view E distal part of left opisthomere, caudo-mesal view F apical part of the same, meso-caudal view. Scale bars: 0.1 mm (A, C-E), 0.05 mm (F), 0.01 mm (B). Abbreviations: bpp basoposterior process, dr distal ridge, f flagellum, g (supposed) gonocoxal gland, ml mesomeroidal lobe, mp mesal process, mr median ridge, msl mesal lobe of basoposterior process, s solenomere, sf spiniform filaments, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure52" httpUri="https://binary.pensoft.net/fig/581987" pageId="0" pageNumber="1">52A</figureCitation>
|
||
) similar to that in
|
||
<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., but more slender, distally mostly symmetrical rather than with a markedly convex lateral margin; mesal process barely, rather than considerably, surpassing the apex of promere, and ending with a finer, less strongly bent tip; median and distal ridges less strongly pronounced compared to the type species. Opisthomere (Figs
|
||
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">51D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 52" captionStartId="F52" captionText="Figure 52. Svaniulus waltheri gen. nov., sp. nov., gonopods of ♂ paratype A right promere, caudal view B apical part of flagellum C right opisthomere, oral view D distal part of the same, latero-oral view E distal part of left opisthomere, caudo-mesal view F apical part of the same, meso-caudal view. Scale bars: 0.1 mm (A, C-E), 0.05 mm (F), 0.01 mm (B). Abbreviations: bpp basoposterior process, dr distal ridge, f flagellum, g (supposed) gonocoxal gland, ml mesomeroidal lobe, mp mesal process, mr median ridge, msl mesal lobe of basoposterior process, s solenomere, sf spiniform filaments, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure52" httpUri="https://binary.pensoft.net/fig/581987" pageId="0" pageNumber="1">52C-F</figureCitation>
|
||
): lacking a distomesal process; basoposterior process narrower in comparison to
|
||
<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., with a distinct apical corner rather than with a flat margin, and with a much smaller mesal lobe; mesal side with a slender lobe (presumably gonocoxal gland) and a rather deep and narrow anteromesal sinus; solenomere somewhat enlarged distally rather than narrowing; otherwise of a very similar shape to that of the type species.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.1058.68628.figure52" httpUri="https://binary.pensoft.net/fig/581987" pageId="0" pageNumber="1" start="Figure 52" startId="F52">
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">Figure 52.</emphasis>
|
||
<taxonomicName class="Diplopoda" family="Julidae" genus="Svaniulus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Svaniulus waltheri" order="Julida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="waltheri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Svaniulus waltheri</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov., gonopods of ♂ paratype
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
|
||
right promere, caudal view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
apical part of flagellum
|
||
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
|
||
right opisthomere, oral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
|
||
distal part of the same, latero-oral view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
|
||
distal part of left opisthomere, caudo-mesal view
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
apical part of the same, meso-caudal view. Scale bars: 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">A, C-E</emphasis>
|
||
), 0.05 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
|
||
), 0.01 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
|
||
). Abbreviations:
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">bpp</emphasis>
|
||
basoposterior process,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">dr</emphasis>
|
||
distal ridge,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">f</emphasis>
|
||
flagellum,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">g</emphasis>
|
||
(supposed) gonocoxal gland,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">ml</emphasis>
|
||
mesomeroidal lobe,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">mp</emphasis>
|
||
mesal process,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">mr</emphasis>
|
||
median ridge,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">msl</emphasis>
|
||
mesal lobe of basoposterior process,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">s</emphasis>
|
||
solenomere,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">sf</emphasis>
|
||
spiniform filaments,
|
||
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">sp</emphasis>
|
||
apical process of solenomere.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
<emphasis bold="true" pageId="0" pageNumber="1">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">Female sexual characters</emphasis>
|
||
:
|
||
</emphasis>
|
||
First two leg pairs of same condition as in
|
||
<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov. Vulva (Fig.
|
||
<figureCitation captionStart="Figure 51" captionStartId="F51" captionText="Figure 51. Svaniulus waltheri gen. nov., sp. nov., ♂ (A-D) and ♀ (E) paratypes A hypoproct, ventral view B right flange of pleurotergum 7, ventro-lateral view C penis, caudal view D right opisthomere, oral view E left vulva, latero-caudal view. Scale bars: 0.1 mm. Abbreviations: ct central tube, g (supposed) gonocoxal gland, ml mesomeroidal lobe, msl mesal lobe of basoposterior process, pt posterior tube, s solenomere, sp apical process of solenomere." figureDoi="10.3897/zookeys.1058.68628.figure51" httpUri="https://binary.pensoft.net/fig/581986" pageId="0" pageNumber="1">51E</figureCitation>
|
||
) externally also very similar to the type species, mainly differing in being more elongated and with the operculum being equal to, rather than slightly shorter than, bursa. Receptaculum seminis with a somewhat wider and shorter central tube, not forming a distinct ampulla at bottom; posterior tube gradually widening towards bottom, rather than constricting to form an ampulla.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="remarks">
|
||
<paragraph pageId="0" pageNumber="1">Remark.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">
|
||
Despite the distance between the type localities of
|
||
<taxonomicName lsidName="S. waltheri" pageId="0" pageNumber="1" rank="species" species="waltheri">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. waltheri</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov. and
|
||
<taxonomicName lsidName="S. ryvkini" pageId="0" pageNumber="1" rank="species" species="ryvkini">
|
||
<emphasis italics="true" pageId="0" pageNumber="1">S. ryvkini</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., sp. nov. being only ca. 35 air-km, the distribution areas of the two species are probably divided by the Egrisi Mountain Range. With summit parts above 3000 m a.s.l., it could have split the population of the common ancestor of these two closely related species during colder periods in the geological past.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="1" type="distribution">
|
||
<paragraph pageId="0" pageNumber="1">General distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="1">SWGC.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |