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<mods:title id="6134BE03949F725BC160F2B2AE663284">Boreolimnus, a new leafhopper genus from northern North America, with a review of Cribrus Oman (Hemiptera, Cicadellidae, Deltocephalinae)</mods:title>
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<subSubSection id="BB8E6CFECC91BC793DEAF3DEAA89537E" type="nomenclature">
<paragraph id="4182633A1F2231183E0D8930B90519AB">
<taxonomicName id="5671A39EBC5117B31733B3A20639A06F" ID-CoL="8R4QD" authority="Oman, 1949" authorityName="Oman" authorityYear="1949" class="Insecta" family="Cicadellidae" genus="Cribrus" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="genus">
<emphasis id="251DBA018A8522C78A7733DC903CC6A4" italics="true">Cribrus</emphasis>
<bibRefCitation id="E136A79DEEAA1CA13A98D6C2F591D203" author="Oman PW" journalOrPublisher="Entomological Society of Washington, Washington, DC" refId="B24" refString="Oman PW (1949) The Nearctic Leafhoppers (Homoptera: Cicadellidae): a Generic Classification and Check List. Entomological Society of Washington, Washington, DC, 153 pp." title="The Nearctic Leafhoppers (Homoptera: Cicadellidae): a Generic Classification and Check List." year="1949">
<collectingCountry id="03BCA9DEF7766DE6C0A6523DFC0C2B38" name="Oman">Oman</collectingCountry>
, 1949
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="SECID0ED6AE" type="type_taxon">
<paragraph id="5653CC493E58CE2252CD1C785DE06727">
<heading id="A0049D1B0B0B3A62DAA55C0B87677235" reason="title">Type species.</heading>
</paragraph>
<paragraph id="D135EEAA024F4DEF12E2E928DED647A7">
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<taxonomicName id="ED331EE9BC8FA68349539523ED4D5061" ID-CoL="8RLMX" authority="Beamer &amp; Tuthill, 1934" authorityName="Beamer &amp; Tuthill" authorityYear="1934" class="Insecta" family="Cicadellidae" genus="Laevicephalus" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="shingwauki">
<emphasis id="329FA2113F5AFFFEBBB1EDA3942E23CA" italics="true">Laevicephalus shingwauki</emphasis>
<bibRefCitation id="41380337EF81A5DB948726A32F4E6370" author="Beamer RH &amp; Tuthill LD" journalOrPublisher="Journal of the Kansas Entomological Society" pagination="1-24" refId="B4" refString="Beamer RH, Tuthill LD (1934) Some new species and a new genus of deltocephaloid leafhoppers (Homoptera, Cicadellidae). Journal of the Kansas Entomological Society 7: 1 - 24. http: // www. jstor. org / stable / 25081372" title="Some new species and a new genus of deltocephaloid leafhoppers (Homoptera, Cicadellidae)." url="http://www.jstor.org/stable/25081372" volume="7" year="1934">Beamer &amp; Tuthill, 1934</bibRefCitation>
</taxonomicName>
, by original designation (
<treatmentCitation id="9EBB3A8891380DCEF01E0BC567E52AD7" author="Oman PW" page="166" year="1949">
<bibRefCitation id="D7F9E6869F87D920D6F9651A4F7426C9" author="Oman PW" firstAuthor="Oman" journalOrPublisher="Entomological Society of Washington, Washington, DC" page="166" refId="B24" refString="Oman PW (1949) The Nearctic Leafhoppers (Homoptera: Cicadellidae): a Generic Classification and Check List. Entomological Society of Washington, Washington, DC, 153 pp." title="The Nearctic Leafhoppers (Homoptera: Cicadellidae): a Generic Classification and Check List." year="1949">
<collectingCountry id="3F6CA32594D87E570F2E03F2C224E0A7" name="Oman">Oman</collectingCountry>
1949: 166
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).
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</paragraph>
</subSubSection>
<subSubSection id="SECID0EY6AE" type="diagnosis">
<paragraph id="3161FD46BBAC3C62479BA892B1156180">
<heading id="205A00981AFA36711F155D55A37C82A6" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="A0B04779ADE264B59275B722F10B8879">
Separated from other genera of
<taxonomicName id="69557FFCC6AD16D5055D57C7B7C68674" authorityName="Distant" authorityYear="1908" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="tribe" tribe="Paralimnini">Paralimnini</taxonomicName>
by the following combination of characters: male subgenital plates with uniseriate macrosetae, plates truncate and shorter than pygofer; pygofer without processes; pygofer with a prominent pair of dorsal spots; connective linear with posterior stem about as long as wide (connective loop-shaped sensu
<bibRefCitation id="5F6D66E8BDF88BB2D33C824FD7D228DB" author="Emeljanov AF" firstAuthor="Emeljanov" journalOrPublisher="Entomological Review" pagination="547-562" refId="B13" refString="Emeljanov AF (1999) A key to genera of the subfamily Deltocephalinae s. l. (Homoptera, Cicadellidae) from Kazakhstan, Middle Asia, and Mongolia with description of new genera and subgenera. Entomological Review 79: 547562." title="A key to genera of the subfamily Deltocephalinae s. l. (Homoptera, Cicadellidae) from Kazakhstan, Middle Asia, and Mongolia with description of new genera and subgenera." volume="79" year="1999">Emeljanov (1999)</bibRefCitation>
); aedeagus with swollen atrium, short shaft with apical gonopore and one pair of apical processes; frontoclypeus and pronotum with longitudinal stripes; wings usually brachypterous, fore wing with three closed anteapical cells.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIAAG" type="description">
<paragraph id="A506DEC268171369B0788D5CBFAD5076">
<heading id="B57BCBE8A2425BE18466D7B53901331C" reason="title">Description.</heading>
</paragraph>
<paragraph id="BDA5266604D6C2CE9CF50B7C74024743">
Small leafhoppers with typical
<taxonomicName id="3281AC650EA82A99E2159B46135757D3" authorityName="Distant" authorityYear="1908" class="Insecta" family="Cicadellidae" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="tribe" tribe="Paralimnini">Paralimnini</taxonomicName>
structure. Colour generally stramineous, head and pronotum with longitudinal stripes, wing with indistinct brown infuscation around cell borders (Figs
<figureCitation id="31334C9C4869C778E0272BAD7D0521A2" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">2631</figureCitation>
).
</paragraph>
<paragraph id="A0EAACA3EF35B57650C5C6796982D624">
Head with crown bluntly angled, medial length about equal to width between eyes (Figs
<figureCitation id="58852FE2ADE4F507F876E17AEAF47914" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">26</figureCitation>
,
<figureCitation id="9393AF58FD80E177727746FF97CA42E8" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">27</figureCitation>
). Crown glabrous at base, margin and face shagreen. Lateral frontal sutures terminating just ventral of ocelli, ocelli about their own diameter distant from eye (Fig.
<figureCitation id="1ACDBA6DA585D5490513D583BE167E34" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">30</figureCitation>
). Mesal margin of eye notched. Anteclypeus with margins slightly convex, distal third distinctly tapered. Lorum about half width of anteclypeus, well separated from genal margin. Antennae about as long as head width.
</paragraph>
<caption id="DFDEFD0F9A81137D7EB1C7DAFF2EBF73" ID-DOI="10.3897/zookeys.1217.126602.figures26-40" ID-arpha="A9660009-BDB8-5D6E-9FEB-CF93E59FFE8F" httpUri="https://binary.pensoft.net/fig/1169803" startId="F5">
<paragraph id="E6684259AFC13E921F15392CD989BD91">
<label id="17A2D7D340864301E7282291768A6B94">Figures 2640.</label>
</paragraph>
<paragraph id="1DD666C32A55DD142C808C4406938DEF">
<taxonomicName id="DB05E6643569E5F9D8E1258278C5D867" baseAuthorityName="Sanders &amp; DeLong" baseAuthorityYear="1917" class="Insecta" family="Cicadellidae" genus="Cribrus" kingdom="Animalia" order="Hemiptera" phylum="Arthropoda" rank="species" species="concinnus">
<emphasis id="5D85C7F05595F4083A57E751B02723EC" italics="true">Cribrus concinnus</emphasis>
</taxonomicName>
<emphasis id="27C821AAC8A394DC5DC10EFFAE1E27D9" bold="true">26</emphasis>
dorsal habitus, male
<emphasis id="B0A83A61D77BD495242C404C295FCEBB" bold="true">27</emphasis>
dorsal habitus, female
<emphasis id="5D6C90C42C42F577858999B15B41A135" bold="true">28</emphasis>
abdomen ventral, female
<emphasis id="6623D78F20263D0B018F87A36CFAF388" bold="true">29</emphasis>
lateral habitus, male
<emphasis id="8212D16A6E70F2803CBD2BA6639F4710" bold="true">30</emphasis>
face
<emphasis id="DAF15660B02B1C4D8BA76672253DA5B3" bold="true">31</emphasis>
lateral habitus, female
<emphasis id="9B08E4D4ED4B7F59B62B8594FBA33018" bold="true">32</emphasis>
male pygofer, dorsal
<emphasis id="D9A7E7D3CFDDFD7AB2DE0235168CAE4E" bold="true">33</emphasis>
male pygofer, lateral
<emphasis id="7B61DD7B7A8254935F08BB1069B8CBA3" bold="true">34</emphasis>
male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal
<emphasis id="56BA9689A624D2140C01EC123B24BC14" bold="true">36</emphasis>
aedeagus, lateral
<emphasis id="2EF445BD0D4CD8751B2FDE0A964BA2DA" bold="true">37</emphasis>
female genital capsule, ventral
<emphasis id="5C9B50003D2ABA3DB585AEDEA7736657" bold="true">38</emphasis>
first valvifer, lateral, with enlargement
<emphasis id="610F9B2A3A6AC6CF70A5BFFB61FC21FD" bold="true">39</emphasis>
second valvifers, lateral, with enlargement
<emphasis id="F03D193C8D7EF00DF3F6C0EFDA362846" bold="true">40</emphasis>
female sternite VII, ventral.
</paragraph>
</caption>
<paragraph id="44C77FC046CDC542BA2E125D185D9355">Pronotum slightly narrower than width of head across eyes, about as long as medial length of head. Fore femur with AM 1 near ventral margin, row IC with a few fine setae, row AV consisting of a few, widely spaced, very short setae. Fore tibia with 1 AD and 4 PD macrosetae. Hind femur with 2 + 2 + 1 macrosetae. First hind tarsomere with two rows of plantar setae, four apical platellae between a pair of longer normal setae. Fore wing with three closed anteapical cells, although venation may be distorted due to brachyptery.</paragraph>
<paragraph id="5303398DBB13FCE121A0344CFCBE0204">
Male abdomen with apodemes on sternite II poorly developed, shorter than width. Pygofer about twice as long as wide, with a patch of long macrosetae posterodorsally and a few small macrosetae scattered ventrally (Figs
<figureCitation id="C591B6BF32BEB491FDF1A52E211E9ABA" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">32</figureCitation>
,
<figureCitation id="00E364DB946871F87FED11E17B50C228" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">33</figureCitation>
). Pygofer dorsally with a pair of heavily sclerotized spots basal to segment X. Segment X about as long as wide, completely sclerotized dorsally and laterally. Valve parabolic. Subgenital plates truncate, shorter than pygofer, bearing a single row of macrosetae laterally (Fig.
<figureCitation id="D6258DE2C3B8547FD5A56FE5309FA7AC" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">34</figureCitation>
). Connective with arms fused anteriorly, tapered towards posterior end, stem abruptly broadened apically, wider than arms. Style apophysis with lateral lobe weakly developed, medial lobe with corrugated sculpture but no distinct teeth. Aedeagus with swollen atrium, shaft very short with apical gonopore, with one pair of apical processes (Figs
<figureCitation id="98A86FB3F8DDABA62B98C05615010FFE" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">35</figureCitation>
,
<figureCitation id="88D9921B617287D479127141679B5F5D" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">36</figureCitation>
).
</paragraph>
<paragraph id="BAEECEF1AA1247419A8CCF29221B26E6">
Female pygofer with moderate length macrosetae (Fig.
<figureCitation id="CD90513E40F1F7B5615EECC9E9793AF5" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">37</figureCitation>
). Ovipositor not projecting beyond pygofer. Gonoplac without macrosetae. First valvula slightly concave; sculpture imbricate dorsally and strigate ventrally (Fig.
<figureCitation id="06733DB888F55A844734551F2F642CCD" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">38</figureCitation>
). Second valvulae evenly tapered distally, with rounded teeth decreasing in size distally (Fig.
<figureCitation id="6F9709258D8033DD556CDD4FE53AE018" captionStart="Figures 2640" captionStartId="F5" captionText="Figures 2640. Cribrus concinnus 26 dorsal habitus, male 27 dorsal habitus, female 28 abdomen ventral, female 29 lateral habitus, male 30 face 31 lateral habitus, female 32 male pygofer, dorsal 33 male pygofer, lateral 34 male subgenital plate, styles, connective, dorsal 35 aedeagus, caudal 36 aedeagus, lateral 37 female genital capsule, ventral 38 first valvifer, lateral, with enlargement 39 second valvifers, lateral, with enlargement 40 female sternite VII, ventral." figureDoi="10.3897/zookeys.1217.126602.figures26-40" httpUri="https://binary.pensoft.net/fig/1169803">39</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>