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<document id="6E73DFDB2A0C6AE5CBFD5E99C7F46875" ID-CLB-Dataset="31635" ID-DOI="http://dx.doi.org/10.3897/zookeys.713.14732" ID-GBIF-Dataset="29242e82-d634-42a5-b250-ab2d71ad5894" ID-PMC="PMC5674209" ID-Pensoft-Pub="1313-2970-713-1" ID-PubMed="29134039" ID-ZooBank="D70E71F8669C4557B266D4FF8359A511" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-713-1" ModsDocOrigin="ZooKeys 713" ModsDocTitle="Monogenea of fishes from the lagoon flats of Palmyra Atoll in the Central Pacific" checkinTime="1509722113451" checkinUser="pensoft" docAuthor="Vidal-Martinez, Victor Manuel, Soler-Jimenez, Lilia Catherinne, Aguirre-Macedo, Ma. Leopoldina, Mclaughlin, John, Jaramillo, Alejandra G., Shaw 2, Jenny C., James, Anna, Hechinger, Ryan F., Kuris, Armand M. &amp; Lafferty, Kevin D." docDate="2017" docId="4991C91FE87CBD89F302D69BBE954A10" docLanguage="en" docName="ZooKeys 713: 1-23" docOrigin="ZooKeys 713" docSource="http://dx.doi.org/10.3897/zookeys.713.14732" docTitle="Acleotrema girellae Johnston &amp; Tiegs 1922" docType="treatment" docVersion="7" lastPageNumber="10" masterDocId="FF87FFACA374FFBBFF8D606A501CFFC0" masterDocTitle="Monogenea of fishes from the lagoon flats of Palmyra Atoll in the Central Pacific" masterLastPageNumber="23" masterPageNumber="1" pageNumber="9" updateTime="1701382954004" updateUser="ExternalLinkService">
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<mods:title id="5DC599BE42F87A7672F8DE5E449FE9CA">Monogenea of fishes from the lagoon flats of Palmyra Atoll in the Central Pacific</mods:title>
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<mods:namePart id="9215A2AEAEE94AA29F1F008FF03F8E97">Vidal-Martinez, Victor Manuel</mods:namePart>
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<mods:namePart id="DA10E39B4E24B5D9A0A620E4AD2AB519">Soler-Jimenez, Lilia Catherinne</mods:namePart>
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<mods:namePart id="FEC20B9CB518095C6EAA9AFE6613D57A">Aguirre-Macedo, Ma. Leopoldina</mods:namePart>
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<mods:namePart id="91853AAF5368764A55727CFB180DF1A9">Mclaughlin, John</mods:namePart>
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<mods:namePart id="EC3A03057C687CF987C3135FA1321C57">Jaramillo, Alejandra G.</mods:namePart>
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<mods:namePart id="3275D74CD8A30D56C12CB1FC884604E8">Shaw 2, Jenny C.</mods:namePart>
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<mods:namePart id="D3B465F8C05ADC3B796039E03D530878">James, Anna</mods:namePart>
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<mods:namePart id="0E8D618EFF096DC82882DC3A535047CE">Hechinger, Ryan F.</mods:namePart>
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<mods:namePart id="F4E435EC7FE654BF1C7A9A4A32C34257">Kuris, Armand M.</mods:namePart>
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<mods:namePart id="71182A95636526DF5F430245C5607622">Lafferty, Kevin D.</mods:namePart>
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<mods:date id="DC0E8482F1EDE31647BC5F8FE026E177">2017</mods:date>
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<mods:number id="353DA5CB3E599F96829EE22605A35DB7">713</mods:number>
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<mods:classification id="85076FBAA40446FCC6298BAA31555C94">journal article</mods:classification>
<mods:identifier id="574156E753059BA6160E6847F9E98019" type="DOI">http://dx.doi.org/10.3897/zookeys.713.14732</mods:identifier>
<mods:identifier id="9C3C2220C49B91E1F362FEA23A84CDAF" type="Pensoft-Pub">1313-2970-713-1</mods:identifier>
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<treatment id="4991C91FE87CBD89F302D69BBE954A10" ID-GBIF-Taxon="137233634" LSID="urn:lsid:plazi:treatment:4991C91FE87CBD89F302D69BBE954A10" httpUri="http://treatment.plazi.org/id/4991C91FE87CBD89F302D69BBE954A10" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<subSubSection id="3E4E655DADA500B7711194D42DB2B692" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="E0E7450763194190F1021B3ABB9648E1" pageId="8" pageNumber="9">
<taxonomicName id="DDCBAA53FDB79343ACCB8B93BB4638BE" ID-CoL="9CP5" authority="Johnston &amp; Tiegs, 1922" authorityName="Johnston &amp; Tiegs" authorityYear="1922" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema girellae" order="Dactylogyridea" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="girellae">Acleotrema girellae Johnston &amp; Tiegs, 1922</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="47D6417A4379F2E2CB9C85F4A3FA3562" pageId="8" pageNumber="9" type="type host">
<paragraph id="6A2D01CFC6D79B5E30DEE878C23A5478" pageId="8" pageNumber="9">Type host.</paragraph>
<paragraph id="2690B6E7F29D86BA9A71E17F9BC97DFE" pageId="8" pageNumber="9">
<taxonomicName id="540A09CBDE5E62B170C71D57269972BD" family="Kyphosidae" genus="Girella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Girella tricuspidata" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="tricuspidata">Girella tricuspidata</taxonomicName>
(Quoy and Gaimard) (
<taxonomicName id="67401F09A9ECD714F117FE6CD2950440" family="Kyphosidae" lsidName="" pageId="8" pageNumber="9" rank="family">Kyphosidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="0C27C8952A0098E7C4BBBC1E6B17B110" pageId="8" pageNumber="9" type="other host and localities">
<paragraph id="F2C3C04DE8DEEE87B2F5E1C869E61E84" pageId="8" pageNumber="9">Other host and localities.</paragraph>
<paragraph id="7BC6E6140AC6898330DA666C219549F3" pageId="8" pageNumber="9">
<taxonomicName id="205A3C176893EDDD2A6C337048A53256" family="Kyphosidae" genus="Girella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Girella tricuspidata" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="tricuspidata">Girella tricuspidata</taxonomicName>
from off Caloundra, southeast of Queensland, Australia (
<bibRefCitation id="376C06616588409A823A179A03C57D79" author="Johnston, TH" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="15" pageNumber="16" pagination="83 - 131" title="New Gyrodactyloid Trematodes from Australian fishes together with a reclassification of the Super-Family Gyrodactyloidea." volume="47" year="1922">Johnston and Tiegs 1922</bibRefCitation>
).
<taxonomicName id="62B8DA2BDAC2E540F5AA80DC3A1C0352" family="Kyphosidae" genus="Kyphosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kyphosus cinerascens" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cinerascens">Kyphosus cinerascens</taxonomicName>
collected off
<normalizedToken id="A974642F1EEA4BCBDDF68B03227C1ED4" originalValue="Hawai’i">Hawai'i</normalizedToken>
(as
<taxonomicName id="15EC120BFEF9E5C7994F633CD62C1DF3" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema kyphosi" order="Dactylogyridea" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="kyphosi">Acleotrema kyphosi</taxonomicName>
) (
<bibRefCitation id="0A8A62FA74B8158BD198DEF0C323DE31" author="Yamaguti, S" journalOrPublisher="University of Hawaii, Honolulu" pageId="19" pageNumber="20" title="Monogenetic trematodes of Hawaiian fishes." url="https://doi.org/10.2307/3277429" year="1968">Yamaguti 1968</bibRefCitation>
).
<taxonomicName id="00B69A35E986448EE30EFD099925BE87" family="Kyphosidae" genus="Kyphosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kyphosus elegans" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="elegans">Kyphosus elegans</taxonomicName>
(Peters) from Chamela Bay, Mexico (as
<taxonomicName id="099BDDB84CD076BAA6F13EED285F25DE" class="Monogenea" family="Diplectanidae" genus="Heteroplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplectanum kyphosi" order="Dactylogyridea" pageId="8" pageNumber="9" phylum="Platyhelminthes" rank="species" species="kyphosi">Heteroplectanum kyphosi</taxonomicName>
) (
<bibRefCitation id="97734DE568E1081047591E596D63C2A7" author="Leon-Regagnon, V" journalOrPublisher="Journal of Helminthology" pageId="16" pageNumber="17" pagination="9 - 16" title="Description of Heteroplectanumoliveri sp. n. (Monogenea: Diplectanidae) and Comments on the Helminth fauna of Kyphosuselegans (Perciformes: Kyphosidae) from Chamela Bay, Mexico." volume="64" year="1997">
<normalizedToken id="406ECA5781C497ACED17860FE0508CCE" originalValue="León-Régagnon">Leon-Regagnon</normalizedToken>
et al. 1997
</bibRefCitation>
).
<taxonomicName id="F9F4731CF748322A4CC1C13E8CB81303" family="Kyphosidae" genus="Kyphosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kyphosus" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="genus">Kyphosus</taxonomicName>
spp. (as
<taxonomicName id="11008622E41AD29EC30A46868BEE6968" lsidName="A. girellae" pageId="8" pageNumber="9" rank="species" species="girellae">A. girellae</taxonomicName>
) from Australia, the Mediterranean Sea and Mexican Pacific (
<bibRefCitation id="C4F032932728A97EBCE6A928F4D0908B" author="Domingues, MV" journalOrPublisher="Systematic Parasitology" pageId="14" pageNumber="15" pagination="35 - 41" title="The status of Acleotrema Johnston and Tiegs, 1922 and Heteroplectanum Rakotofiringa, Oliver and Lambert, 1987 (Monogenoidea: Diplectanidae), with the redescription of Acleotremagirellae Johnston and Tiegs, 1922." url="https://doi.org/10.1007/s11230-006-9051-9" volume="66" year="2007">Domingues and Boeger 2007</bibRefCitation>
) (all
<taxonomicName id="364556E240C3CBC60E727DB500E37A55" family="Kyphosidae" lsidName="" pageId="8" pageNumber="9" rank="family">Kyphosidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="F499AF56F45DDB4B6AD7808B3653C58C" pageId="8" pageNumber="9" type="current host">
<paragraph id="89016238D33BC5A6B3B746FAAB7432BD" pageId="8" pageNumber="9">Current host.</paragraph>
<paragraph id="8D04373E0EBF413E014E0BCDD69453E2" pageId="8" pageNumber="9">
<taxonomicName id="FBFFF72D1DA2074378879B9EA9B69761" family="Kyphosidae" genus="Kyphosus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Kyphosus cinerascens" order="Perciformes" pageId="8" pageNumber="9" phylum="Chordata" rank="species" species="cinerascens">Kyphosus cinerascens</taxonomicName>
(
<taxonomicName id="4E3BF0A67FEC442274F2E2F204BAD50A" family="Kyphosidae" lsidName="" pageId="8" pageNumber="9" rank="family">Kyphosidae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="E812503914E98247EAEBE7D092922EE3" pageId="8" pageNumber="9" type="site infection">
<paragraph id="3D2D68ADAB5CD7AEA1C945B216443F64" pageId="8" pageNumber="9">Site infection.</paragraph>
<paragraph id="B36BA2DE83A043F7186049064031A71B" pageId="8" pageNumber="9">Gills.</paragraph>
</subSubSection>
<subSubSection id="6EC77A10F0021CFC19C0F7652A47148E" pageId="9" pageNumber="10" type="specimens deposited">
<paragraph id="C388499ADEDBFDFB3BA37D8735140D21" pageId="9" pageNumber="10">
<pageBreakToken id="53831845030297227F54F8CF1367336F" pageId="9" pageNumber="10" start="start">Specimens</pageBreakToken>
deposited.
</paragraph>
<paragraph id="4BDA3B402FE5B9ED2D84F6E84C3110F3" pageId="9" pageNumber="10">CHCM No. 540 (paratypes) (1 slide, 2 specimen), USNM No. 1459852 (voucher).</paragraph>
</subSubSection>
<subSubSection id="C82ADDAB3E627F6092105876278CE005" pageId="9" pageNumber="10" type="prevalence and mean intensity">
<paragraph id="A02DB9AD0F4EA50C7CC383DE3C031331" pageId="9" pageNumber="10">Prevalence and mean intensity.</paragraph>
<paragraph id="24D4902880B47DEDAC01151C110696DD" pageId="9" pageNumber="10">
100 and 84
<normalizedToken id="03EAF74162DA105D5C7C617397FFC8D5" originalValue="±">+/-</normalizedToken>
90 (n=2).
</paragraph>
</subSubSection>
<subSubSection id="91D02BFD6ABE141B867477B48B930A42" pageId="9" pageNumber="10" type="remarks">
<paragraph id="3893FDDFF9635850C89F17B51A186CB9" pageId="9" pageNumber="10">Remarks.</paragraph>
<paragraph id="A9C29DC6F7B87C3C0B6F16C9D481FC63" pageId="9" pageNumber="10">
<taxonomicName id="621CBEEAE1FFD4B848B9B2650582431C" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema girellae" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="girellae">Acleotrema girellae</taxonomicName>
was originally described from the gills of
<taxonomicName id="6EEC62EBCF67361B68D78B58BD2C29B7" lsidName="G. tricuspidata" pageId="9" pageNumber="10" rank="species" species="tricuspidata">G. tricuspidata</taxonomicName>
collected off Caloundra, southeast Queensland, Australia (
<bibRefCitation id="B107ACE431FB5046943168B0EFF73517" author="Johnston, TH" journalOrPublisher="Proceedings of the Linnean Society of New South Wales" pageId="15" pageNumber="16" pagination="83 - 131" title="New Gyrodactyloid Trematodes from Australian fishes together with a reclassification of the Super-Family Gyrodactyloidea." volume="47" year="1922">Johnston and Tiegs 1922</bibRefCitation>
). In 1937, Price transferred this species to
<taxonomicName id="9674BD197CC03E3F5778EEF8F762D522" class="Monogenea" family="Diplectanidae" genus="Diplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplectanum" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Diplectanum</taxonomicName>
as
<taxonomicName id="3F1E56E280A36ED895C17C8B9F0EF05B" lsidName="D. girellae" pageId="9" pageNumber="10" rank="species" species="girellae">D. girellae</taxonomicName>
, considering
<taxonomicName id="2C24C25A32DA2A34701A8D8FA09DA225" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Acleotrema</taxonomicName>
a junior synonym of
<taxonomicName id="FCFDE3BC7E721F7E4B37FF4FA9AAE4F5" class="Monogenea" family="Diplectanidae" genus="Diplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplectanum" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Diplectanum</taxonomicName>
, based on the presence of squamodiscs. However,
<bibRefCitation id="FDEB8167CFEEE45647FF5C0FDF523AD4" author="Yamaguti, S" journalOrPublisher="London-New York, Interscience Publishers" pageId="18" pageNumber="19" title="Monogenea and Aspidocotylea. Systema Helmithum IV." year="1963">Yamaguti (1963)</bibRefCitation>
accepted
<taxonomicName id="0ACAAD30093C84EE86A03A91F1DF808A" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Acleotrema</taxonomicName>
as a valid genus.
<bibRefCitation id="276422524B5824045D2178401074F9C9" pageId="9" pageNumber="10">Rakotofiringa et al. (1987)</bibRefCitation>
proposed the genus
<taxonomicName id="1A9A21F555B2D1C115D7076FCAF33011" class="Monogenea" family="Diplectanidae" genus="Heteroplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplectanum" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Heteroplectanum</taxonomicName>
and several species have been transferred to this new genus, including
<taxonomicName id="B6831958D7E3CC272425815AF3E63DAF" class="Monogenea" family="Diplectanidae" genus="Diplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplectanum kyphosi" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="kyphosi">Diplectanum kyphosi</taxonomicName>
(considered a synonym of
<taxonomicName id="B82B8EE0224B165F9BFB100267239B4E" lsidName="A. girellae" pageId="9" pageNumber="10" rank="species" species="girellae">A. girellae</taxonomicName>
) as
<taxonomicName id="2A251C6251D6208EFF71C3B1003D8D0E" class="Monogenea" family="Diplectanidae" genus="Heteroplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplectanum kyphosi" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="kyphosi">Heteroplectanum kyphosi</taxonomicName>
(Yamaguti, 1968)
<bibRefCitation id="DD86C2D0425F097DD1CE520526F6985D" author="Oliver, G" journalOrPublisher="Universite des Sciences et Techniques du Languedoc, Montpellier, France" pageId="17" pageNumber="18" title="Les Diplectanidae Bychowsky 1957 (Monogenea, Monopisthocotylea, Dactylogyridea). Systematique. Biologie. Ontogenie. Ecologie. Essai de phylogenese. These d'etat." year="1987">Oliver 1987</bibRefCitation>
. However,
<bibRefCitation id="013B4771F0E1C17D1E9F6351C10E5A7F" author="Domingues, MV" journalOrPublisher="Systematic Parasitology" pageId="14" pageNumber="15" pagination="35 - 41" title="The status of Acleotrema Johnston and Tiegs, 1922 and Heteroplectanum Rakotofiringa, Oliver and Lambert, 1987 (Monogenoidea: Diplectanidae), with the redescription of Acleotremagirellae Johnston and Tiegs, 1922." url="https://doi.org/10.1007/s11230-006-9051-9" volume="66" year="2007">Domingues and Boeger (2007)</bibRefCitation>
considered that species of
<taxonomicName id="D8C89DE98C5A5117AB2CDA9B217115AE" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Acleotrema</taxonomicName>
share unique features and can be distinguished from other diplectanids (including species of
<taxonomicName id="0D9A3AD004926BA0E9A218348F9AC6A0" class="Monogenea" family="Diplectanidae" genus="Diplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplectanum" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Diplectanum</taxonomicName>
), presenting arguments for considering
<taxonomicName id="7C140150B1E740E483FFB00EF9C41801" class="Monogenea" family="Diplectanidae" genus="Heteroplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplectanum" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Heteroplectanum</taxonomicName>
as a junior synonym of
<taxonomicName id="B144483844DE44110B1D4BFE4DA34A7F" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="genus">Acleotrema</taxonomicName>
. Therefore,
<taxonomicName id="A259DD933B69059600442FEEF155A5CB" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema kyphosi" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="kyphosi">Acleotrema kyphosi</taxonomicName>
Yamaguti, 1968,
<taxonomicName id="DD4C47156D13504DB4B3B822A5291156" class="Monogenea" family="Diplectanidae" genus="Diplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplectanum girellae" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="girellae">Diplectanum girellae</taxonomicName>
(Johnston &amp; Tiegs, 1922),
<taxonomicName id="B114B6264530DE0F5A540912C2865F39" class="Monogenea" family="Diplectanidae" genus="Heteroplectanum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Heteroplectanum kyphosi" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="kyphosi">Heteroplectanum kyphosi</taxonomicName>
(Yamaguti, 1968) Oliver, 1987,
<taxonomicName id="CCA0F2A2422CF3895AC4E95877C739A6" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema gibsoni" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="gibsoni">Acleotrema gibsoni</taxonomicName>
Young, 1970 and
<taxonomicName id="27E06EFD4908E1660D45CAA6A2317486" class="Monogenea" family="Diplectanidae" genus="Acleotrema" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Acleotrema heronensis" order="Dactylogyridea" pageId="9" pageNumber="10" phylum="Platyhelminthes" rank="species" species="heronensis">Acleotrema heronensis</taxonomicName>
Young, 1970, are considered synonyms of
<taxonomicName id="08EBD5C9F1DB8DD20193B7AE4D08DF39" lsidName="A. girellae" pageId="9" pageNumber="10" rank="species" species="girellae">A. girellae</taxonomicName>
. This species differs from its congeners by having: a tubular MCO with the distal extremity recurved and bifid; and a sclerotised sac with radial musculature involving the proximal portion of the MCO. New geographical record for Palmyra Atoll.
</paragraph>
</subSubSection>
</treatment>
</document>
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