treatments-xml/data/E0/8A/66/E08A66F2F1B05A1E8BE328BE42094967.xml

<document ID-DOI="http://dx.doi.org/10.3897/asp.80.e73829" ID-Pensoft-Pub="1864-8312-80-309" ID-Pensoft-UUID="11907D1B6F665D96B5FD2F9751431366" ID-ZooBank="D5C0468B99A14EF39237D9BC51A8BDA3" ModsDocID="1864-8312-80-309" checkinTime="1660711556436" checkinUser="pensoft" docAuthor="Benedetti, Alipio Rezende &amp; Pinto-da-Rocha, Ricardo" docDate="2022" docId="E08A66F2F1B05A1E8BE328BE42094967" docLanguage="en" docName="ArthropodSystPhyl 80: 309-388" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 80" docPubDate="2022-08-16" docSource="http://dx.doi.org/10.3897/asp.80.e73829" docTitle="Ayacucho spiniger Benedetti &amp; Pinto-da-Rocha 2022, comb. nov." docType="treatment" docVersion="1" id="11907D1B6F665D96B5FD2F9751431366" lastPageNumber="309" masterDocId="11907D1B6F665D96B5FD2F9751431366" masterDocTitle="Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species" masterLastPageNumber="388" masterPageNumber="309" pageNumber="309" updateTime="1660711556436" updateUser="pensoft">
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<mods:title>Systematic revision and total evidence phylogenetic analysis of the Andean family Metasarcidae Kury, 1994 (Opiliones: Laniatores), with description of two new genera and twenty new species</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Benedetti, Alipio Rezende</mods:namePart>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil &amp; Instituto Federal de Minas Gerais, campus Sao Joao Evangelista, Av. Primeiro de Junho, 1043, Sao Joao Evangelista, MG, Brazil</mods:affiliation>
<mods:nameIdentifier type="email">alipiobenedetti@gmail.com</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Pinto-da-Rocha, Ricardo</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3959-2205</mods:nameIdentifier>
<mods:affiliation>Departamento de Zoologia, Instituto de Biociencias, Universidade de Sao Paulo, Caixa Postal: 11.461, 05422 - 970, Sao Paulo, SP, Brazil</mods:affiliation>
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<mods:title>Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:date>2022</mods:date>
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<mods:number>2022-08-16</mods:number>
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3.21. 
<taxonomicName LSID="E08A66F2-F1B0-5A1E-8BE3-28BE42094967" authority="(Roewer, 1957)" authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho spiniger" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="spiniger" status="comb. nov.">Ayacucho spiniger (Roewer, 1957)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="309">comb. nov.</taxonomicNameLabel>
</paragraph>
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<subSubSection pageId="0" pageNumber="309" type="description">
<paragraph pageId="0" pageNumber="309">
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">Figs 3B</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">, 7Q, R</figureCitation>
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Penis of Ayacucho, dorsal, lateral and ventral views, respectively. A - C A. spiniger (Roewer, 1957) comb. nov.; D - F A. glauberrochai sp. nov.; G - I A. tapacocha nom. nov.; Legend bars = 0.1 mm" figureDoi="10.3897/asp.80.e73829.figure16" httpUri="https://binary.pensoft.net/fig/729890" pageId="0" pageNumber="309">, 16 A-C</figureCitation>
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">, 23A, B</figureCitation>
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">, 28</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="reference_group">
<paragraph pageId="0" pageNumber="309">
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti &amp; Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Palcares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Palcares spiniger" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="spiniger" status="comb. nov.">Palcares spiniger</taxonomicName>
Roewer, 1957: 72 (desc.), fig. 4 (dorsal habitus, femur IV), 5 (details of femur IV armature), 6 (pedipalpus); 
<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
: 145 (cat.)
</paragraph>
<paragraph pageId="0" pageNumber="309">
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Benedetti &amp; Pinto-da-Rocha" baseAuthorityYear="2022" class="Arachnida" family="Metasarcidae" genus="Palcares" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Palcares serrifemur" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="serrifemur" status="comb. nov.">Palcares serrifemur</taxonomicName>
Roewer, 1959: 70 (desc.) fig. 2A (dorsal habitus, femur IV), 2b (details of femur IV armature); 
<bibRefCitation author="Kury, AB" journalOrPublisher="Tropical Zoology" pageId="0" pageNumber="309" refId="B51" refString="Kury, AB, 2003. Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol.7: 1-337." title="Annotated catalogue of the Laniatores of the New World (Arachnida, Opiliones). Revista Iberica de Aracnologia, vol. 7: 1 - 337." year="2003">Kury 2003</bibRefCitation>
: 145 (cat.), 
<emphasis bold="true" pageId="0" pageNumber="309">syn. n.</emphasis>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="redescription">
<paragraph pageId="0" pageNumber="309">Redescription.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">
MALE: 
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=4) DSW: 3.4-3.8 (3.7); DSL: 4.2-4.6 (4.6); CL: 1.4-1.5 (1.4). FIVL: 4.9-5.9 (5.9). ChL: 1.3-2.7 (2.7). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Coloration</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">23A</figureCitation>
) Dorsal scutum and legs yellow-orange. Anterior and posterior margins of DS and free tergites darker. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Dorsum</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3B</figureCitation>
) Alpha-type DSS, with shallower constriction II (almost faint). Anterior margin of DS with median elevation completely covered by granules. Ocularium covered with tiny granules and two long spines. Carapace densely covered by tiny granules. Areas I-IV densely covered with small granules; I with a pair of small spiniform tubercles; II and IV each one with a pair of tubercles twice the size of those on area I; III with pair of spines. Lateral margins of dorsal scutum covered with minute granules. Posterior margin of dorsal scutum with a small to slightly larger granules, armed with 2-3 spines; a single tubercle located to the right of the pair of spines, when present. Free tergites covered with granules of different sizes and armed with spines. Free tergite I with a pair of spines; II with 2-3 spines, two large and one three times lower; III with three spines. Anal operculum with three tubercles. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Chelicerae</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3B</figureCitation>
) Swollen in large males (swollen in holotype); similar to females in small males. Segment I densely covered with granules. Segment II predominantly smooth, with setae at certain points of the frontal face, with two teeth on finger. Segment III with one tooth. 
<taxonomicName genus="Pedipalpus" lsidName="Pedipalpus" pageId="0" pageNumber="309" rank="genus">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Pedipalpus</emphasis>
</taxonomicName>
: With small granules distributed mainly on the dorsal surface of the femur, tibia and patella. Trochanter with a ventrodistal setiferous tubercle. Femur with a ventrobasal setiferous tubercle and a row of six ventral setiferous tubercles. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Venter</emphasis>
: Coxae I-IV with granules sparsely distributed. Genital area almost smooth. Free sternites I-III with rows of small tubercles. Genital operculum granulate. 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Legs</emphasis>
: (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Habitus, dorsal of males of Ayacucho. A A. spielbergi sp. nov.; B A. spiniger (Roewer, 1957) comb. nov.; C A. tapacocha nom. nov.; D A. titschacki Roewer, 1949; E Ayacucho triarmatus nom. nov.; F A. weyrauchi (Roewer, 1952) comb. nov.; G Ayacucho vargasllosai sp. nov.; H A. uniseriatus (Roewer, 1959) comb. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure3" httpUri="https://binary.pensoft.net/fig/729877" pageId="0" pageNumber="309">3B</figureCitation>
, 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7Q, R</figureCitation>
) Coxae I-II each one with a prolateral and a retrolateral apophyses; III with a retrolateral apophysis; IV with granules distributed more densely on the prolateral face and an apical prodorsal apophysis. Trochanters I-IV with sparsely distributed granules. Femora I-III granular. Femur IV slightly sinuous, granular; with a retrolateral row of 15 tubercles along the apical ⅔ and a prolateral row of 12-13 tubercles along the apical 
<normalizedToken originalValue="½">1/2</normalizedToken>
; retro and prolateral tubercles size growing apically; the smallest tubercles are short and blunt, while the largest are long and with an approximately straight or lanceolate apex. Patella IV with granules distributed throughout its length and with three dorsoapical tubercles. Tibiae I-IV with sparse granules. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=8) 6-8 (7), 11-14 (11), 7-8 (7), 8-9 (8). 
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Penis</emphasis>
: (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Penis of Ayacucho, dorsal, lateral and ventral views, respectively. A - C A. spiniger (Roewer, 1957) comb. nov.; D - F A. glauberrochai sp. nov.; G - I A. tapacocha nom. nov.; Legend bars = 0.1 mm" figureDoi="10.3897/asp.80.e73829.figure16" httpUri="https://binary.pensoft.net/fig/729890" pageId="0" pageNumber="309">16A-C</figureCitation>
) VP rectangular; distal margin slightly concave; large lateral projections. MS C1 -C3 subapical long and apically curved; MS A1 sub basal long and straight; MS B1 sub basal long and straight (MS A1 and MS B1 shorter than MS C1-C3); MS D1 short, dorsally placed, next to MS A. Lateral sacs long and with blunt apex; with long T3-like microsetae. Stylus dorsally curved, with subapical ventral projection. Dorsal process long and tapered. Promontory convex. - 
<emphasis bold="true" pageId="0" pageNumber="309">
FEMALE: 
<emphasis italics="true" pageId="0" pageNumber="309">Measurements</emphasis>
</emphasis>
(
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=4) DSW: 3.4-3.8; DSL: 4.2-4.6; CL: 1.4-1.5. FIVL: 4.9-5.5. ChL: 1.3-1.6. (Fig. 
<figureCitation captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Live specimens of Ayacucho. A Ayacucho spiniger (Roewer, 1957) comb. nov., male; B female; C A. tapacocha nom. nov., male; D female; E A. titschacki Roewer, 1949, male; F A. vargasllosai sp. nov., male; G A. uniseriatus (Roewer, 1959) comb. nov., male; H: female." figureDoi="10.3897/asp.80.e73829.figure23" httpUri="https://binary.pensoft.net/fig/729897" pageId="0" pageNumber="309">23B</figureCitation>
) DSS with constriction II well marked. Granules of ocularium smaller than in male. Pedipalpal femur with a proapical spine, absent in males. Femora IV unarmed. Dorsodistal apophysis of coxa IV slightly lower than in male. Tarsal segmentation: (
<emphasis italics="true" pageId="0" pageNumber="309">n</emphasis>
=4): 7-8, 10-13, 7-9, 8.
</paragraph>
<caption doi="10.3897/asp.80.e73829.figure15" httpUri="https://binary.pensoft.net/fig/729889" pageId="0" pageNumber="309" start="Figure 15" startId="F15">
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" pageId="0" pageNumber="309">Figure 15.</emphasis>
Penis of 
<taxonomicName authorityName="Roewer" authorityYear="1949" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho</emphasis>
</taxonomicName>
, dorsal, lateral and ventral views, respectively. 
<emphasis bold="true" pageId="0" pageNumber="309">A</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="309">C</emphasis>
<taxonomicName genus="A." lsidName="A. pomacocha" pageId="0" pageNumber="309" rank="species" species="pomacocha">
<emphasis italics="true" pageId="0" pageNumber="309">A. pomacocha</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="309">D</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="309">F</emphasis>
<taxonomicName genus="A." lsidName="A. querococha" pageId="0" pageNumber="309" rank="species" species="querococha">
<emphasis italics="true" pageId="0" pageNumber="309">A. querococha</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
; 
<emphasis bold="true" pageId="0" pageNumber="309">G</emphasis>
- 
<emphasis bold="true" pageId="0" pageNumber="309">I</emphasis>
<taxonomicName genus="A." lsidName="A. silvae" pageId="0" pageNumber="309" rank="species" species="silvae">
<emphasis italics="true" pageId="0" pageNumber="309">A. silvae</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">sp. nov.</emphasis>
; Legend bars = 0.1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="diagnosis">
<paragraph pageId="0" pageNumber="309">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="309">
Similar to 
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho inermis" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="inermis">
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho inermis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
because of two rows of tubercles on the femur IV (pro and retrolateral; Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7Q, R</figureCitation>
). Differs from 
<taxonomicName authorityName="Benedetti &amp; Pinto-da-Rocha" authorityYear="2022" baseAuthorityName="Roewer" baseAuthorityYear="1957" class="Arachnida" family="Metasarcidae" genus="Ayacucho" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ayacucho inermis" order="Opiliones" pageId="0" pageNumber="309" phylum="Arthropoda" rank="species" species="inermis">
<emphasis italics="true" pageId="0" pageNumber="309">Ayacucho inermis</emphasis>
</taxonomicName>
<emphasis bold="true" pageId="0" pageNumber="309">comb. nov.</emphasis>
because femur IV with some lanceolate tubercles (Fig. 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Male trochanter-tibia IV of Ayacucho, dorsal and ventral view, respectively. A - B A. bambamarca (Roewer, 1957) comb. nov.; C - D A. glauberrochai sp. nov.; E - F A. inermis (Roewer, 1957) comb. nov.; G - H A. pasolinii sp. nov.; I - J A. pomacocha sp. nov.; K - L A. querococha sp. nov.; M - N A. silvae sp. nov.; O - P A. spielbergi sp. nov.; Q - R A. spiniger (Roewer, 1957) comb. nov.; S - T: A. tapacocha nom. nov.; U - V A. titschacki Roewer, 1949; W - X: A. triarmatus nom. nov.; Legend bars = 1 mm." figureDoi="10.3897/asp.80.e73829.figure7" httpUri="https://binary.pensoft.net/fig/729881" pageId="0" pageNumber="309">7Q, R</figureCitation>
); ventral plate of penis with conspicuous lateral projections; long MS C on VP; lateral sacs with long T3-like microsetae, dorsal process long in relation to stylus (Fig. 
<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Penis of Ayacucho, dorsal, lateral and ventral views, respectively. A - C A. spiniger (Roewer, 1957) comb. nov.; D - F A. glauberrochai sp. nov.; G - I A. tapacocha nom. nov.; Legend bars = 0.1 mm" figureDoi="10.3897/asp.80.e73829.figure16" httpUri="https://binary.pensoft.net/fig/729890" pageId="0" pageNumber="309">16A-C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="distribution">
<paragraph pageId="0" pageNumber="309">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="309">
(Fig. 
<figureCitation captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Distribution of Ayacucho spp. in Peru" figureDoi="10.3897/asp.80.e73829.figure28" httpUri="https://binary.pensoft.net/fig/729902" pageId="0" pageNumber="309">28</figureCitation>
) PERU. 
<normalizedToken originalValue="Junín">Junin</normalizedToken>
. 
<normalizedToken originalValue="Campañillaya">Campanillaya</normalizedToken>
. Hacienda Maraynioc. Palca
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="309" type="materials_examined">
<paragraph pageId="0" pageNumber="309">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="309">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">Type material</emphasis>
: Of 
<taxonomicName genus="P." lsidName="P. spiniger" pageId="0" pageNumber="309" rank="species" species="spiniger">
<emphasis italics="true" pageId="0" pageNumber="309">P. spiniger</emphasis>
</taxonomicName>
: 
<materialsCitation accessionNumber="RII 11418" collectingDate="1955-01-06" collectorName="Weyrauch, Of" country="PERU" elevation="2600" location="Palca in Rio Tarma" municipality="Campanillaya" specimenCount="♂" stateProvince="Junin" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Holotype</typeStatus>
</emphasis>
<specimenCount>♂</specimenCount>
, ' 
<collectingCountry name="Peru">PERU</collectingCountry>
, 
<normalizedToken originalValue="Junín">
<collectingRegion country="Peru" name="Junin">Junin</collectingRegion>
</normalizedToken>
, 
<normalizedToken originalValue="Campañillaya">
<collectingMunicipality>Campanillaya</collectingMunicipality>
</normalizedToken>
, near to 
<location LSID="urn:lsid:plazi:treatment:E08A66F2F1B05A1E8BE328BE42094967:75D3CBDB2B45219DC18FDFE9FAA273D0" country="PERU" municipality="Campanillaya" name="Palca in Rio Tarma" stateProvince="Junin">Palca in Rio Tarma</location>
, 
<elevation metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="2.6" unit="m" value="2600.0">2,600 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="1955-01-06">06/I/1955</collectingDate>
, 
<collectorName>Weyrauch</collectorName>
leg. (SMF 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII11418">RII 11418</accessionNumber>
/24). 
<collectorName>Of</collectorName>
<taxonomicName genus="P." lsidName="P. serrifemur" pageId="0" pageNumber="309" rank="species" species="serrifemur">
<emphasis italics="true" pageId="0" pageNumber="309">P. serrifemur</emphasis>
</taxonomicName>
</materialsCitation>
: 
<materialsCitation accessionNumber="RII 12764" collectingDate="1956-10-10" collectorName="Weyrauch" country="PERU" county="Hacienda Maraynioc" elevation="3500" location="Chanchamayo bay" municipality="east Tarma" specimenCount="♂" stateProvince="Junin" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Holotype</typeStatus>
</emphasis>
<specimenCount>♂</specimenCount>
, ' 
<collectingCountry name="Peru">PERU</collectingCountry>
, 
<normalizedToken originalValue="Junín">
<collectingRegion country="Peru" name="Junin">Junin</collectingRegion>
</normalizedToken>
, 
<collectingCounty>Hacienda Maraynioc</collectingCounty>
, 
<collectingMunicipality>east Tarma</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:E08A66F2F1B05A1E8BE328BE42094967:21233B53019186E234270829A137D92D" country="PERU" county="Hacienda Maraynioc" municipality="east Tarma" name="Chanchamayo bay" stateProvince="Junin">Chanchamayo bay</location>
, 
<elevation metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="m" value="3500.0">
<quantity metricMagnitude="3" metricUnit="m" metricValue="3.5" unit="m" value="3500.0">3,500 m</quantity>
a.s.l.
</elevation>
, 
<collectingDate value="1956-10-10">10/X/1956</collectingDate>
, 
<collectorName>Weyrauch</collectorName>
leg. (SMF 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII12764">RII 12764</accessionNumber>
/35)
</materialsCitation>
- 
<materialsCitation accessionNumber="RII 12764" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" country="Peru" specimenCount="1" specimenCount-female="1" stateProvince="Junin" typeStatus="Paratype">
<emphasis bold="true" pageId="0" pageNumber="309">
<typeStatus>Paratype</typeStatus>
</emphasis>
<specimenCount type="female">1 ♀</specimenCount>
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(SMF 
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/protein/RII12764">RII 12764</accessionNumber>
/35)
</materialsCitation>
. 
<materialsCitation collectingDate="2011-04-19" collectorName="R. Pinto-da-Rocha, A. Benedetti, J. Ochoa, D. Silva" country="PERU" latitude="-11.358028" location="Palca" longLatPrecision="1" longitude="-75.55664" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Junin" typeStatus="Additional material">
<emphasis bold="true" italics="true" pageId="0" pageNumber="309">
<typeStatus isOtherTypeStatus="true">Additional material</typeStatus>
</emphasis>
: 
<specimenCount type="male">3 ♂</specimenCount>
, 
<specimenCount type="female">3 ♀</specimenCount>
' 
<collectingCountry name="Peru">PERU</collectingCountry>
, 
<normalizedToken originalValue="Junín">
<collectingRegion country="Peru" name="Junin">Junin</collectingRegion>
</normalizedToken>
, 
<location LSID="urn:lsid:plazi:treatment:E08A66F2F1B05A1E8BE328BE42094967:AD888DDC826E147D7B1E5F68D36FC5ED" country="PERU" latitude="-11.358028" longLatPrecision="1" longitude="-75.55664" name="Palca" stateProvince="Junin">Palca</location>
, 
<geoCoordinate degrees="11" direction="south" minutes="21" orientation="latitude" precision="1" seconds="28.9" value="-11.358028">11°21′28.9″S</geoCoordinate>
<geoCoordinate degrees="75" direction="west" minutes="33" orientation="longitude" precision="1" seconds="23.9" value="-75.55664">75°33′23.9″W</geoCoordinate>
, 
<collectingDate value="2011-04-19">19/IV/2011</collectingDate>
, 
<collectorName>R. Pinto-da-Rocha</collectorName>
, 
<collectorName>A. Benedetti</collectorName>
, 
<collectorName>J. Ochoa</collectorName>
&amp; 
<collectorName>D. Silva</collectorName>
leg. (MUBI)
</materialsCitation>
; 
<specimenCount type="male">3 ♂</specimenCount>
, 
<specimenCount type="female">2 ♀</specimenCount>
, 
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MUSM); 
<specimenCount type="male">7 ♂</specimenCount>
, 
<specimenCount type="female">5 ♀</specimenCount>
<normalizedToken originalValue="‘ditto’">'ditto'</normalizedToken>
(MZSP 36980).
</paragraph>
</subSubSection>
</treatment>
</document>