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<document id="FEFBF704D1F765FF2265EB6C64A1BBA8" ID-CLB-Dataset="39200" ID-DOI="10.11646/zootaxa.4060.1.11" ID-GBIF-Dataset="1fcc6102-d3b3-4fda-8a8e-a4ecff813b9f" ID-ISSN="1175-5326" ID-Zenodo-Dep="233206" ID-ZooBank="4ECBC9B2-D644-4FB2-BC1A-F67D0B97B474" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461365081050" checkinUser="plazi" docAuthor="Ho, Hsuan-Ching" docDate="2015" docId="B72287E4FF9BFFFBFF21DDC2FD5428E0" docLanguage="en" docName="zt04060p085.pdf" docOrigin="Zootaxa 4060 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ophichthus bicolor McCosker &amp; Ho, sp. nov." docType="treatment" docUuid="2CF4B7F6-4DBA-4BC7-B9E2-5283066E96DE" docUuidSource="ZooBank" docVersion="9" lastPageNumber="80" masterDocId="4B1BFF9CFF9EFFF2FFB6D83EFFCC2B0C" masterDocTitle="New species of the snake eels Echelus and Ophichthus (Anguilliformes: Ophichthidae) from Taiwan" masterLastPageNumber="85" masterPageNumber="71" pageNumber="76" updateTime="1698661944915" updateUser="ExternalLinkService">
<mods:mods id="CA703EA241048AC7314A151622C543AF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="1BDED7B816E8D221664DA7726A90A35D">
<mods:title id="D2D0F50C7E81E82536287BC2BDB0561A">New species of the snake eels Echelus and Ophichthus (Anguilliformes: Ophichthidae) from Taiwan</mods:title>
</mods:titleInfo>
<mods:name id="D8AB3C6ED983E28E852397CC2BEE8CE1" type="personal">
<mods:role id="31E5F211B364A49ED4336F3217EA5F30">
<mods:roleTerm id="D56307F8F0D4A770D39C0AA0910AF5DB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="107FC89BC13A97B5DE1DDC1F659EFDE5">Ho, Hsuan-Ching</mods:namePart>
</mods:name>
<mods:typeOfResource id="E147D6234CF9A60E76E7EE059998A199">text</mods:typeOfResource>
<mods:relatedItem id="7BF841E9F86C1B7B23FB88535F83FE8C" type="host">
<mods:titleInfo id="23CB920958DBBF7AE5A95E710A768F4C">
<mods:title id="9A89A539A5E3E1EB1B3ED595B13547D0">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AD878745870266C6DC2FFA5863CF3405">
<mods:date id="D20672C99A4B0639AC504A32E6583FF6">2015</mods:date>
<mods:detail id="56096F13F6780FBDF8CB8BE6DC8D7179" type="volume">
<mods:number id="661C334D37D96334FAFD902533974360">4060</mods:number>
</mods:detail>
<mods:detail id="0BB03A4D5CC8FF25756ED9DB601F0288" type="issue">
<mods:number id="52F83CB8D3B877A33E7ADBF2B8770558">1</mods:number>
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<mods:extent id="FA6D994CA205E57AB36FB094CB935683" unit="page">
<mods:start id="69688D7977FD61A7FE2DEC1616FAF419">71</mods:start>
<mods:end id="2E7B82B2DBF2ED3A8E8E7431E7081441">85</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="F16E1F0C5331BA28482712237E17F926">journal article</mods:classification>
<mods:identifier id="98113A14953525ABAC22CCE1D86EBDEF" type="CLB-Dataset">39200</mods:identifier>
<mods:identifier id="584850FD62745938B64BF67D84082ED6" type="DOI">10.11646/zootaxa.4060.1.11</mods:identifier>
<mods:identifier id="94FE97888D9F73263B813E530C9C2EA4" type="GBIF-Dataset">1fcc6102-d3b3-4fda-8a8e-a4ecff813b9f</mods:identifier>
<mods:identifier id="AEE84D46232E4C8B573407BD6D6B6433" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="6541911A608F6DF1DC9BBB49D3C0EABE" type="Zenodo-Dep">233206</mods:identifier>
<mods:identifier id="FDEA8717595B88E6CFC7C6B9BE358815" type="ZooBank">4ECBC9B2-D644-4FB2-BC1A-F67D0B97B474</mods:identifier>
</mods:mods>
<treatment id="B72287E4FF9BFFFBFF21DDC2FD5428E0" ID-DOI="http://doi.org/10.5281/zenodo.5618734" ID-GBIF-Taxon="119649132" ID-Zenodo-Dep="5618734" LSID="urn:lsid:zoobank.org:act:2CF4B7F6-4DBA-4BC7-B9E2-5283066E96DE" httpUri="http://treatment.plazi.org/id/B72287E4FF9BFFFBFF21DDC2FD5428E0" lastPageId="9" lastPageNumber="80" pageId="5" pageNumber="76">
<subSubSection id="77916579FF9BFFF7FF21DDC2FEB82D52" pageId="5" pageNumber="76" type="nomenclature">
<paragraph id="3F3436F2FF9BFFF7FF21DDC2FD662D1A" blockId="5.[151,682,1532,1630]" box="[151,682,1532,1558]" pageId="5" pageNumber="76">
<heading id="647C819EFF9BFFF7FF21DDC2FD662D1A" bold="true" box="[151,682,1532,1558]" fontSize="10" level="2" pageId="5" pageNumber="76" reason="2">
<emphasis id="0DFFEAE0FF9BFFF7FF21DDC2FD662D1A" bold="true" box="[151,682,1532,1558]" pageId="5" pageNumber="76">
<taxonomicName id="F88B4D71FF9BFFF7FF21DDC2FD882D1A" ID-CoL="74P5J" authority="McCosker &amp; Ho" authorityName="McCosker &amp; Ho" box="[151,580,1532,1558]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="5" pageNumber="76" phylum="Chordata" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="0DFFEAE0FF9BFFF7FF21DDC2FEBA2D19" bold="true" box="[151,374,1532,1557]" italics="true" pageId="5" pageNumber="76">Ophichthus bicolor</emphasis>
McCosker &amp; Ho
</taxonomicName>
,
<taxonomicNameLabel id="16CC579BFF9BFFF7FDE7DDC3FD662D1A" box="[593,682,1533,1558]" pageId="5" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="3F3436F2FF9BFFF7FF21DE1FFEB12D36" blockId="5.[151,682,1532,1630]" box="[151,381,1569,1594]" pageId="5" pageNumber="76">
<emphasis id="0DFFEAE0FF9BFFF7FF21DE1FFEB12D36" bold="true" box="[151,381,1569,1594]" pageId="5" pageNumber="76">Bicolored snake eel</emphasis>
</paragraph>
<paragraph id="3F3436F2FF9BFFF7FF21DE7BFEB82D52" blockId="5.[151,682,1532,1630]" box="[151,372,1605,1630]" pageId="5" pageNumber="76">
(
<figureCitation id="A7B02A77FF9BFFF7FF29DE7BFEC72D52" box="[159,267,1605,1630]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="6.[151,244,1933,1954]" captionStartId-1="8.[151,244,955,976]" captionStartId-2="8.[151,244,1773,1794]" captionTargetBox-0="[388,1198,201,1884]" captionTargetBox-1="[261,1342,598,938]" captionTargetBox-2="[259,1336,1064,1751]" captionTargetId-0="figure@6.[388,1198,190,1904]" captionTargetId-1="figure@8.[250,1342,587,943]" captionTargetId-2="figure@8.[253,1345,1057,1752]" captionTargetPageId-0="6" captionTargetPageId-1="8" captionTargetPageId-2="8" captionText-0="FIGURE 4. Ophichthus bicolor sp. nov. A. NMMB-P 18103, holotype, 559 mm TL, preserved. B. NMMB-P 22212, paratype, 696 mm TL, fresh. C. NMMB-P 17544, paratype, 701 mm TL, fresh. Bars indicate the origins of the dorsal fin (left) and anal fin (right)." captionText-1="FIGURE 5. Semi-diagrammatic illustration of lateral head of Ophichthus bicolor McCosker &amp; Ho, sp. nov., from the holotype. Arrows point to the frontal pore (left) and median pore of supratemporal series (right)." captionText-2="FIGURE 6. Semi-diagrammatic illustration of dentition of Ophichthus bicolor McCosker &amp; Ho, sp. nov., from the holotype. A. Upper Jaw. B. Lower Jaw" httpUri-0="https://zenodo.org/record/233210/files/figure.png" httpUri-1="https://zenodo.org/record/233211/files/figure.png" httpUri-2="https://zenodo.org/record/233212/files/figure.png" pageId="5" pageNumber="76">Figs. 4–6</figureCitation>
;
<tableCitation id="72090349FF9BFFF7FEA1DE7BFEA72D52" box="[279,363,1605,1630]" captionStart="TABLE 2" captionStartId="7.[151,231,439,460]" captionTargetBox="[159,1184,620,1257]" captionTargetPageId="7" captionText="TABLE 2. Counts and proportions (in thousandths) of the holotype and selected paratypes of Ophichthus bicolor sp. nov." httpUri="http://table.plazi.org/id/6BF4667AFF99FFF5FF21D989FAEB2AC0" pageId="5" pageNumber="76" tableUuid="6BF4667AFF99FFF5FF21D989FAEB2AC0">Table 2</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="77916579FF9BFFF7FF21DEB3FD0A2CE6" pageId="5" pageNumber="76" type="materials_examined">
<paragraph id="3F3436F2FF9BFFF7FF21DEB3FF1A2DC6" blockId="5.[151,1437,1677,2026]" pageId="5" pageNumber="76">
<emphasis id="0DFFEAE0FF9BFFF7FF21DEB3FEC72DAA" bold="true" box="[151,267,1677,1702]" pageId="5" pageNumber="76">
<typeStatus id="E0308850FF9BFFF7FF21DEB3FECA2DAA" box="[151,262,1677,1702]" pageId="5" pageNumber="76" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="599AAE37FF9BFFF7FEA5DEB0FE4B2DAA" box="[275,391,1678,1702]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
18103 (male,
<quantity id="F8739B17FF9BFFF7FD97DEB3FD482DAA" box="[545,644,1677,1702]" metricMagnitude="-1" metricUnit="m" metricValue="5.59" pageId="5" pageNumber="76" unit="mm" value="559.0">559 mm</quantity>
), SW
<collectingCountry id="479C7662FF9BFFF7FD7CDEB3FCD32DAA" box="[714,799,1677,1702]" name="Taiwan" pageId="5" pageNumber="76">Taiwan</collectingCountry>
, Dong-gang, Pingtung, bottom trawl,
<emphasis id="0DFFEAE0FF9BFFF7FB7ADEB0FB212DA9" box="[1228,1261,1678,1701]" italics="true" pageId="5" pageNumber="76">ca.</emphasis>
<quantity id="F8739B17FF9BFFF7FB40DEB0FA8C2DA9" box="[1270,1344,1678,1702]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="76" unit="m" value="200.0">200 m</quantity>
,
<date id="4B351032FF9BFFF7FAFBDEB0FF1F2DC6" pageId="5" pageNumber="76" value="2012-09-07">7 Sept. 2012</date>
.
</paragraph>
<paragraph id="3F3436F2FF9BFFF7FF71DEEBFD0A2CE6" blockId="5.[151,1437,1677,2026]" pageId="5" pageNumber="76">
<emphasis id="0DFFEAE0FF9BFFF7FF71DEEBFE8A2DE2" bold="true" box="[199,326,1749,1774]" pageId="5" pageNumber="76">
<typeStatus id="E0308850FF9BFFF7FF71DEEBFE8D2DE2" box="[199,321,1749,1774]" pageId="5" pageNumber="76" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="298DFD7BFF9BFFF7FEFADEE8FE2A2DE2" box="[332,486,1749,1774]" pageId="5" pageNumber="76" type="generic">21 specimens</specimenCount>
,
<quantity id="F8739B17FF9BFFF7FE46DEE8FD482DE1" box="[496,644,1750,1774]" metricMagnitude="-1" metricUnit="m" metricValue="7.04" metricValueMax="9.19" metricValueMin="4.89" pageId="5" pageNumber="76" unit="mm" value="704.0" valueMax="919.0" valueMin="489.0">489–919 mm</quantity>
TL. Collected from nearby the
<typeStatus id="E0308850FF9BFFF7FC55DEEBFB892DE2" box="[995,1093,1749,1774]" pageId="5" pageNumber="76" type="holotype">holotype</typeStatus>
locality by bottom trawl:
<collectionCode id="599AAE37FF9BFFF7FAD0DEE8FA502DE2" LSID="urn:lsid:biocol.org:col:34700" box="[1382,1436,1750,1774]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="5" pageNumber="76">CAS</collectionCode>
234696 (2 spec.,
<quantity id="F8739B17FF9BFFF7FEE8DEC7FE352C1E" box="[350,505,1785,1810]" metricMagnitude="-1" metricUnit="m" metricValue="6.29" metricValueMax="6.409999999999999" metricValueMin="6.17" pageId="5" pageNumber="76" unit="mm" value="629.0" valueMax="641.0" valueMin="617.0">617–641 mm</quantity>
),
<date id="4B351032FF9BFFF7FDB8DEC4FD602C1E" box="[526,684,1786,1810]" pageId="5" pageNumber="76" value="2012-05-10">10 May 2012</date>
;
<collectionCode id="599AAE37FF9BFFF7FD0CDEC4FCE22C1E" box="[698,814,1786,1810]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
14077 (
<quantity id="F8739B17FF9BFFF7FC34DEC4FC242C1E" box="[898,1000,1786,1810]" metricMagnitude="-1" metricUnit="m" metricValue="4.9" pageId="5" pageNumber="76" unit="mm" value="490.0">490 mm</quantity>
),
<collectionCode id="599AAE37FF9BFFF7FC4BDEC4FBBC2C1E" box="[1021,1136,1786,1810]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
14082 (
<quantity id="F8739B17FF9BFFF7FB73DEC7FAE62C1E" box="[1221,1322,1785,1810]" metricMagnitude="-1" metricUnit="m" metricValue="6.13" pageId="5" pageNumber="76" unit="mm" value="613.0">613 mm</quantity>
),
<date id="4B351032FF9BFFF7FAF6DEC4FF1D2C3A" pageId="5" pageNumber="76" value="2011-08-10">10 Aug. 2011</date>
;
<collectionCode id="599AAE37FF9BFFF7FF68DF23FE9D2C3A" box="[222,337,1821,1846]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
16640 (
<quantity id="F8739B17FF9BFFF7FE10DF23FDC62C3A" box="[422,522,1821,1846]" metricMagnitude="-1" metricUnit="m" metricValue="5.06" pageId="5" pageNumber="76" unit="mm" value="506.0">506 mm</quantity>
),
<date id="4B351032FF9BFFF7FDA8DF20FD792C3A" box="[542,693,1822,1846]" pageId="5" pageNumber="76" value="2011-10-20">20 Oct. 2011</date>
;
<collectionCode id="599AAE37FF9BFFF7FD77DF23FCF82C3A" box="[705,820,1821,1846]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
17544 (
<quantity id="F8739B17FF9BFFF7FC3FDF20FC212C3A" box="[905,1005,1822,1846]" metricMagnitude="-1" metricUnit="m" metricValue="7.01" pageId="5" pageNumber="76" unit="mm" value="701.0">701 mm</quantity>
),
<date id="4B351032FF9BFFF7FBB7DF20FB5F2C3A" box="[1025,1171,1822,1846]" pageId="5" pageNumber="76" value="2012-08-08">8 Aug. 2012</date>
;
<collectionCode id="599AAE37FF9BFFF7FB16DF23FAD92C3A" box="[1184,1301,1821,1846]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
21657-61 (5 spec., 521–603), collected with the
<typeStatus id="E0308850FF9BFFF7FD9FDF7FFD422C56" box="[553,654,1857,1882]" pageId="5" pageNumber="76" type="holotype">holotype</typeStatus>
;
<collectionCode id="599AAE37FF9BFFF7FD2DDF7FFCC22C56" box="[667,782,1857,1882]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
21662-64 (4 spec.,
<quantity id="F8739B17FF9BFFF7FC50DF7FFB4C2C56" box="[998,1152,1857,1882]" metricMagnitude="-1" metricUnit="m" metricValue="5.83" metricValueMax="6.12" metricValueMin="5.54" pageId="5" pageNumber="76" unit="mm" value="583.0" valueMax="612.0" valueMin="554.0">554–612 mm</quantity>
), collected with NMMB- P14082;
<collectionCode id="599AAE37FF9BFFF7FF4ADF5BFEA32C72" box="[252,367,1893,1918]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
5791 (
<quantity id="F8739B17FF9BFFF7FE03DF5BFDD42C72" box="[437,536,1893,1918]" metricMagnitude="-1" metricUnit="m" metricValue="6.45" pageId="5" pageNumber="76" unit="mm" value="645.0">645 mm</quantity>
),
<date id="4B351032FF9BFFF7FD9ADF58FD092C72" box="[556,709,1893,1918]" pageId="5" pageNumber="76" value="1966-09-12">12 Sep. 1966</date>
;
<collectionCode id="599AAE37FF9BFFF7FD67DF58FCF52C71" box="[721,825,1894,1917]" httpUri="http://grbio.org/cool/kt9t-t1e8" name="National Museum of Marine Biology and Aquarium" pageId="5" pageNumber="76">NMMBP</collectionCode>
17544 (
<quantity id="F8739B17FF9BFFF7FC21DF5BFC362C72" box="[919,1018,1893,1918]" metricMagnitude="-1" metricUnit="m" metricValue="6.959999999999999" pageId="5" pageNumber="76" unit="mm" value="696.0">696 mm</quantity>
),
<date id="4B351032FF9BFFF7FBB8DF58FB522C72" box="[1038,1182,1894,1918]" pageId="5" pageNumber="76" value="2012-08-08">8 Aug. 2012</date>
;
<collectionCode id="599AAE37FF9BFFF7FB1DDF58FADF2C71" box="[1195,1299,1894,1917]" httpUri="http://grbio.org/cool/kt9t-t1e8" name="National Museum of Marine Biology and Aquarium" pageId="5" pageNumber="76">NMMBP</collectionCode>
14077 (
<quantity id="F8739B17FF9BFFF7FAC7DF58FF0B2CAE" metricMagnitude="-1" metricUnit="m" metricValue="4.89" pageId="5" pageNumber="76" unit="mm" value="489.0">489 mm</quantity>
),
<date id="4B351032FF9BFFF7FF6FDFB4FEB92CAE" box="[217,373,1930,1954]" pageId="5" pageNumber="76" value="2011-08-10">10 Aug. 2011</date>
. Collected from Chang-bin, Taitung, E.
<collectingCountry id="479C7662FF9BFFF7FC84DFB7FC482CAE" box="[818,900,1929,1954]" name="Taiwan" pageId="5" pageNumber="76">Taiwan</collectingCountry>
by hook and line:
<collectionCode id="599AAE37FF9BFFF7FBE2DFB7FB0B2CAE" box="[1108,1223,1929,1954]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
16624 (
<quantity id="F8739B17FF9BFFF7FAACDFB7FAB02CAE" box="[1306,1404,1929,1954]" metricMagnitude="-1" metricUnit="m" metricValue="7.859999999999999" pageId="5" pageNumber="76" unit="mm" value="786.0">786 mm</quantity>
),
<date id="4B351032FF9BFFF7FA38DFB4FECA2CCA" pageId="5" pageNumber="76" value="2012-01-08">8 Jan. 2012</date>
;
<collectionCode id="599AAE37FF9BFFF7FEA5DF90FEB72CC9" box="[275,379,1966,1989]" httpUri="http://grbio.org/cool/kt9t-t1e8" name="National Museum of Marine Biology and Aquarium" pageId="5" pageNumber="76">NMMBP</collectionCode>
16891 (
<quantity id="F8739B17FF9BFFF7FE6FDF90FDF02CCA" box="[473,572,1966,1990]" metricMagnitude="-1" metricUnit="m" metricValue="9.19" pageId="5" pageNumber="76" unit="mm" value="919.0">919 mm</quantity>
),
<date id="4B351032FF9BFFF7FDE6DF90FD272CCA" box="[592,747,1966,1990]" pageId="5" pageNumber="76" value="2012-05-10">10 May 2012</date>
. Collected from Ke-tzu-liao, Kaohsiung by bottom trawl, ca.
<quantity id="F8739B17FF9BFFF7FF21DFECFF2E2CE5" box="[151,226,2002,2026]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="76" unit="m" value="200.0">200 m</quantity>
:
<collectionCode id="599AAE37FF9BFFF7FF5BDFECFE932CE6" box="[237,351,2002,2026]" pageId="5" pageNumber="76">NMMB-P</collectionCode>
22212 (
<quantity id="F8739B17FF9BFFF7FE05DFEFFDD92CE6" box="[435,533,2001,2026]" metricMagnitude="-1" metricUnit="m" metricValue="6.959999999999999" pageId="5" pageNumber="76" unit="mm" value="696.0">696 mm</quantity>
),
<date id="4B351032FF9BFFF7FD9EDFECFD0E2CE6" box="[552,706,2001,2026]" pageId="5" pageNumber="76" value="2015-03-28">28 Mar. 2015</date>
.
</paragraph>
</subSubSection>
<caption id="6BF4667AFF98FFF4FF21DFB3FEE62CD1" httpUri="https://zenodo.org/record/233210/files/figure.png" pageId="6" pageNumber="77" targetBox="[388,1198,201,1884]" targetPageId="6">
<paragraph id="3F3436F2FF98FFF4FF21DFB3FEE62CD1" blockId="6.[151,1435,1933,2013]" pageId="6" pageNumber="77">
<emphasis id="0DFFEAE0FF98FFF4FF21DFB3FEC02CAE" bold="true" box="[151,268,1933,1954]" pageId="6" pageNumber="77">FIGURE 4.</emphasis>
<taxonomicName id="F88B4D71FF98FFF4FEA4DFB3FE072CAE" box="[274,459,1933,1954]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="6" pageNumber="77" phylum="Chordata" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="0DFFEAE0FF98FFF4FEA4DFB3FE072CAE" box="[274,459,1933,1954]" italics="true" pageId="6" pageNumber="77">Ophichthus bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16CC579BFF98FFF4FE64DFB3FDD22CAE" box="[466,542,1933,1954]" pageId="6" pageNumber="77" rank="species">
<emphasis id="0DFFEAE0FF98FFF4FE64DFB3FDD22CAE" bold="true" box="[466,542,1933,1954]" pageId="6" pageNumber="77">sp. nov.</emphasis>
</taxonomicNameLabel>
A. NMMB-P18103, holotype, 559 mm TL, preserved. B. NMMB-P 22212, paratype, 696 mm TL, fresh. C. NMMB-P 17544, paratype, 701 mm TL, fresh. Bars indicate the origins of the dorsal fin (left) and anal fin (right).
</paragraph>
</caption>
<subSubSection id="77916579FF99FFF5FF71D8A9FEBF2D42" pageId="7" pageNumber="78" type="diagnosis">
<paragraph id="3F3436F2FF99FFF5FF71D8A9FEBF2A84" blockId="7.[151,1437,151,392]" pageId="7" pageNumber="78">
<emphasis id="0DFFEAE0FF99FFF5FF71D8A9FE8D2BBC" bold="true" box="[199,321,151,176]" pageId="7" pageNumber="78">Diagnosis.</emphasis>
A moderately elongate species of
<taxonomicName id="F88B4D71FF99FFF5FD7ED8A9FC852BBC" box="[712,841,151,176]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="78" phylum="Chordata" rank="genus">
<emphasis id="0DFFEAE0FF99FFF5FD7ED8A9FC852BBC" box="[712,841,151,176]" italics="true" pageId="7" pageNumber="78">Ophichthus</emphasis>
</taxonomicName>
with: tail 53–58%, head 8.6–10.8%, and body depth at gill opening 2.5–3.4% of total length; dorsal-fin origin behind gill opening by 2.0–2.5 pectoral-fin lengths; pectoral fin pointed, not elongate; posterior nostril a hole above the upper lip, covered by a flap that extends slightly below the edge of the mouth; upper lip lacks barbels; pores small but conspicuous, SO 1+4, IO 4+2, POM 6+2; teeth large and conical, biserial anteriorly on vomer and maxillary, uniserial on mandible; coloration yellowish-tan, strongly contrasting with white throat and belly, fins pale; total vertebrae 155–163, mean vertebral formula 18-64-158.
</paragraph>
<caption id="6BF4667AFF99FFF5FF21D989FAEB2AC0" ID-Table-UUID="6BF4667AFF99FFF5FF21D989FAEB2AC0" box="[151,1319,439,460]" httpUri="http://table.plazi.org/id/6BF4667AFF99FFF5FF21D989FAEB2AC0" pageId="7" pageNumber="78" targetBox="[159,1184,620,1257]" targetIsTable="true" targetPageId="7">
<paragraph id="3F3436F2FF99FFF5FF21D989FAEB2AC0" blockId="7.[151,1319,439,460]" box="[151,1319,439,460]" pageId="7" pageNumber="78">
<emphasis id="0DFFEAE0FF99FFF5FF21D989FF322AC0" bold="true" box="[151,254,439,460]" pageId="7" pageNumber="78">TABLE 2.</emphasis>
Counts and proportions (in thousandths) of the holotype and selected paratypes of
<taxonomicName id="F88B4D71FF99FFF5FBADD989FB182AC0" box="[1051,1236,439,460]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="7" pageNumber="78" phylum="Chordata" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="0DFFEAE0FF99FFF5FBADD989FB182AC0" box="[1051,1236,439,460]" italics="true" pageId="7" pageNumber="78">Ophichthus bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="16CC579BFF99FFF5FB6DD989FAEB2AC0" box="[1243,1319,439,460]" pageId="7" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="3F3436F2FF99FFF5FD45D9D9FBDC2AF0" blockId="7.[159,1152,487,596]" box="[755,1040,487,508]" pageId="7" pageNumber="78">
<typeStatus id="E0308850FF99FFF5FD45D9D9FC802AF0" box="[755,844,487,508]" pageId="7" pageNumber="78" type="holotype">Holotype</typeStatus>
<typeStatus id="E0308850FF99FFF5FC05D9D9FBDC2AF0" box="[947,1040,487,508]" pageId="7" pageNumber="78" type="paratype">Paratypes</typeStatus>
</paragraph>
<paragraph id="3F3436F2FF99FFF5FF29DA2DFB822924" blockId="7.[159,1152,487,596]" box="[159,1102,531,552]" pageId="7" pageNumber="78">TL (mm) 559 489–919 (n=20)</paragraph>
<paragraph id="3F3436F2FF99FFF5FC02DA01FB532E18" blockId="7.[159,1152,487,596]" lastBlockId="7.[159,1183,1279,1300]" pageId="7" pageNumber="78">mean range Total vertebrae 160 158.3 155–163</paragraph>
<paragraph id="3F3436F2FF99FFF5FF29DA52FB4B2FE5" pageId="7" pageNumber="78">
<table id="4D8BC452FF99000DFF29DA52FB6C2FE5" box="[159,1184,620,1257]" gridcols="4" gridrows="15" pageId="7" pageNumber="78">
<tr id="81BB34B0FF99000DFF29DA52FB6C298D" box="[159,1184,620,641]" gridrow="0" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DA52FEB5298D" box="[159,377,620,641]" gridcol="0" gridrow="0" pageId="7" pageNumber="78">HL/TL</th>
<th id="C26A5DCCFF99000DFD45DA52FCDB298D" box="[755,791,620,641]" gridcol="1" gridrow="0" pageId="7" pageNumber="78">92</th>
<th id="C26A5DCCFF99000DFC05DA52FC11298D" box="[947,989,620,641]" gridcol="2" gridrow="0" pageId="7" pageNumber="78">95</th>
<th id="C26A5DCCFF99000DFBFDDA52FB6C298D" box="[1099,1184,620,641]" gridcol="3" gridrow="0" pageId="7" pageNumber="78">86–108</th>
</tr>
<tr id="81BB34B0FF99000DFF29DAA9FB6C29A0" box="[159,1184,663,684]" gridrow="1" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DAA9FEB529A0" box="[159,377,663,684]" gridcol="0" gridrow="1" pageId="7" pageNumber="78">Head and trunk/TL</th>
<td id="C26A5DCCFF99000DFD45DAA9FCDB29A0" box="[755,791,663,684]" gridcol="1" gridrow="1" pageId="7" pageNumber="78">445</td>
<td id="C26A5DCCFF99000DFC05DAA9FC1129A0" box="[947,989,663,684]" gridcol="2" gridrow="1" pageId="7" pageNumber="78">445</td>
<td id="C26A5DCCFF99000DFBFDDAA9FB6C29A0" box="[1099,1184,663,684]" gridcol="3" gridrow="1" pageId="7" pageNumber="78">424–462</td>
</tr>
<tr id="81BB34B0FF99000DFF29DAFAFB6C29D5" box="[159,1184,708,729]" gridrow="2" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DAFAFEB529D5" box="[159,377,708,729]" gridcol="0" gridrow="2" pageId="7" pageNumber="78">Trunk/TL</th>
<td id="C26A5DCCFF99000DFD45DAFAFCDB29D5" box="[755,791,708,729]" gridcol="1" gridrow="2" pageId="7" pageNumber="78">353</td>
<td id="C26A5DCCFF99000DFC05DAFAFC1129D5" box="[947,989,708,729]" gridcol="2" gridrow="2" pageId="7" pageNumber="78">350</td>
<td id="C26A5DCCFF99000DFBFDDAFAFB6C29D5" box="[1099,1184,708,729]" gridcol="3" gridrow="2" pageId="7" pageNumber="78">336–371</td>
</tr>
<tr id="81BB34B0FF99000DFF29DAD1FB6C2808" box="[159,1184,751,772]" gridrow="3" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DAD1FEB52808" box="[159,377,751,772]" gridcol="0" gridrow="3" pageId="7" pageNumber="78">Tail/TL</th>
<td id="C26A5DCCFF99000DFD45DAD1FCDB2808" box="[755,791,751,772]" gridcol="1" gridrow="3" pageId="7" pageNumber="78">555</td>
<td id="C26A5DCCFF99000DFC05DAD1FC112808" box="[947,989,751,772]" gridcol="2" gridrow="3" pageId="7" pageNumber="78">555</td>
<td id="C26A5DCCFF99000DFBFDDAD1FB6C2808" box="[1099,1184,751,772]" gridcol="3" gridrow="3" pageId="7" pageNumber="78">531–572</td>
</tr>
<tr id="81BB34B0FF99000DFF29DB22FB6C283D" box="[159,1184,796,817]" gridrow="4" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DB22FEB5283D" box="[159,377,796,817]" gridcol="0" gridrow="4" pageId="7" pageNumber="78">Dorsal fin-origin/TL</th>
<td id="C26A5DCCFF99000DFD45DB22FCDB283D" box="[755,791,796,817]" gridcol="1" gridrow="4" pageId="7" pageNumber="78">156</td>
<td id="C26A5DCCFF99000DFC05DB22FC11283D" box="[947,989,796,817]" gridcol="2" gridrow="4" pageId="7" pageNumber="78">153</td>
<td id="C26A5DCCFF99000DFBFDDB22FB6C283D" box="[1099,1184,796,817]" gridcol="3" gridrow="4" pageId="7" pageNumber="78">136–183</td>
</tr>
<tr id="81BB34B0FF99000DFF29DB79FB6C2850" box="[159,1184,839,860]" gridrow="5" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DB79FEB52850" box="[159,377,839,860]" gridcol="0" gridrow="5" pageId="7" pageNumber="78">Pectoral fin-length/HL</th>
<td id="C26A5DCCFF99000DFD45DB79FCDB2850" box="[755,791,839,860]" gridcol="1" gridrow="5" pageId="7" pageNumber="78">292</td>
<td id="C26A5DCCFF99000DFC05DB79FC112850" box="[947,989,839,860]" gridcol="2" gridrow="5" pageId="7" pageNumber="78">280</td>
<td id="C26A5DCCFF99000DFBFDDB79FB6C2850" box="[1099,1184,839,860]" gridcol="3" gridrow="5" pageId="7" pageNumber="78">216–340</td>
</tr>
<tr id="81BB34B0FF99000DFF29DB4AFB6C2885" box="[159,1184,884,905]" gridrow="6" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DB4AFEB52885" box="[159,377,884,905]" gridcol="0" gridrow="6" pageId="7" pageNumber="78">Pectoral fin-base/HL</th>
<td id="C26A5DCCFF99000DFD45DB4AFCDB2885" box="[755,791,884,905]" gridcol="1" gridrow="6" pageId="7" pageNumber="78">86</td>
<td id="C26A5DCCFF99000DFC05DB4AFC112885" box="[947,989,884,905]" gridcol="2" gridrow="6" pageId="7" pageNumber="78">103</td>
<td id="C26A5DCCFF99000DFBFDDB4AFB6C2885" box="[1099,1184,884,905]" gridcol="3" gridrow="6" pageId="7" pageNumber="78">77–128</td>
</tr>
<tr id="81BB34B0FF99000DFF29DBA1FB6C28B8" box="[159,1184,927,948]" gridrow="7" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DBA1FEB528B8" box="[159,377,927,948]" gridcol="0" gridrow="7" pageId="7" pageNumber="78">Upper jaw/HL</th>
<td id="C26A5DCCFF99000DFD45DBA1FCDB28B8" box="[755,791,927,948]" gridcol="1" gridrow="7" pageId="7" pageNumber="78">377</td>
<td id="C26A5DCCFF99000DFC05DBA1FC1128B8" box="[947,989,927,948]" gridcol="2" gridrow="7" pageId="7" pageNumber="78">435</td>
<td id="C26A5DCCFF99000DFBFDDBA1FB6C28B8" box="[1099,1184,927,948]" gridcol="3" gridrow="7" pageId="7" pageNumber="78">377–484</td>
</tr>
<tr id="81BB34B0FF99000DFF29DBF2FB6C28ED" box="[159,1184,972,993]" gridrow="8" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DBF2FEB528ED" box="[159,377,972,993]" gridcol="0" gridrow="8" pageId="7" pageNumber="78">Snout/HL</th>
<td id="C26A5DCCFF99000DFD45DBF2FCDB28ED" box="[755,791,972,993]" gridcol="1" gridrow="8" pageId="7" pageNumber="78">206</td>
<td id="C26A5DCCFF99000DFC05DBF2FC1128ED" box="[947,989,972,993]" gridcol="2" gridrow="8" pageId="7" pageNumber="78">188</td>
<td id="C26A5DCCFF99000DFBFDDBF2FB6C28ED" box="[1099,1184,972,993]" gridcol="3" gridrow="8" pageId="7" pageNumber="78">172–215</td>
</tr>
<tr id="81BB34B0FF99000DFF29DBC9FB6C2F00" box="[159,1184,1015,1036]" gridrow="9" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DBC9FEB52F00" box="[159,377,1015,1036]" gridcol="0" gridrow="9" pageId="7" pageNumber="78">Eye/HL</th>
<td id="C26A5DCCFF99000DFD45DBC9FCDB2F00" box="[755,791,1015,1036]" gridcol="1" gridrow="9" pageId="7" pageNumber="78">123</td>
<td id="C26A5DCCFF99000DFC05DBC9FC112F00" box="[947,989,1015,1036]" gridcol="2" gridrow="9" pageId="7" pageNumber="78">115</td>
<td id="C26A5DCCFF99000DFBFDDBC9FB6C2F00" box="[1099,1184,1015,1036]" gridcol="3" gridrow="9" pageId="7" pageNumber="78">103–135</td>
</tr>
<tr id="81BB34B0FF99000DFF29DC1AFB6C2F35" box="[159,1184,1060,1081]" gridrow="10" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DC1AFEB52F35" box="[159,377,1060,1081]" gridcol="0" gridrow="10" pageId="7" pageNumber="78">Depth at GO/HL</th>
<td id="C26A5DCCFF99000DFD45DC1AFCDB2F35" box="[755,791,1060,1081]" gridcol="1" gridrow="10" pageId="7" pageNumber="78">31</td>
<td id="C26A5DCCFF99000DFC05DC1AFC112F35" box="[947,989,1060,1081]" gridcol="2" gridrow="10" pageId="7" pageNumber="78">31</td>
<td id="C26A5DCCFF99000DFBFDDC1AFB6C2F35" box="[1099,1184,1060,1081]" gridcol="3" gridrow="10" pageId="7" pageNumber="78">26–41</td>
</tr>
<tr id="81BB34B0FF99000DFF29DC71FB6C2F68" box="[159,1184,1103,1124]" gridrow="11" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DC71FEB52F68" box="[159,377,1103,1124]" gridcol="0" gridrow="11" pageId="7" pageNumber="78">Width/depth at GO</th>
<td id="C26A5DCCFF99000DFD45DC71FCDB2F68" box="[755,791,1103,1124]" gridcol="1" gridrow="11" pageId="7" pageNumber="78">790</td>
<td id="C26A5DCCFF99000DFC05DC71FC112F68" box="[947,989,1103,1124]" gridcol="2" gridrow="11" pageId="7" pageNumber="78">913</td>
<td id="C26A5DCCFF99000DFBFDDC71FB6C2F68" box="[1099,1184,1103,1124]" gridcol="3" gridrow="11" pageId="7" pageNumber="78">790–986</td>
</tr>
<tr id="81BB34B0FF99000DFF29DC42FB6C2F9D" box="[159,1184,1148,1169]" gridrow="12" pageId="7" pageNumber="78" rowspan-1="1" rowspan-2="1" rowspan-3="1">
<th id="C26A5DCCFF99000DFF29DC42FEB52F9D" box="[159,377,1148,1169]" gridcol="0" gridrow="12" pageId="7" pageNumber="78">Vertebral count (n=19)</th>
</tr>
<tr id="81BB34B0FF99000DFF29DC99FB6C2FB0" box="[159,1184,1191,1212]" gridrow="13" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DC99FEB52FB0" box="[159,377,1191,1212]" gridcol="0" gridrow="13" pageId="7" pageNumber="78">Predorsal vertebrae</th>
<td id="C26A5DCCFF99000DFD45DC99FCDB2FB0" box="[755,791,1191,1212]" gridcol="1" gridrow="13" pageId="7" pageNumber="78">19</td>
<td id="C26A5DCCFF99000DFC05DC99FC112FB0" box="[947,989,1191,1212]" gridcol="2" gridrow="13" pageId="7" pageNumber="78">18.4</td>
<td id="C26A5DCCFF99000DFBFDDC99FB6C2FB0" box="[1099,1184,1191,1212]" gridcol="3" gridrow="13" pageId="7" pageNumber="78">15–23</td>
</tr>
<tr id="81BB34B0FF99000DFF29DCEAFB6C2FE5" box="[159,1184,1236,1257]" gridrow="14" pageId="7" pageNumber="78">
<th id="C26A5DCCFF99000DFF29DCEAFEB52FE5" box="[159,377,1236,1257]" gridcol="0" gridrow="14" pageId="7" pageNumber="78">Preanal vertebrae</th>
<td id="C26A5DCCFF99000DFD45DCEAFCDB2FE5" box="[755,791,1236,1257]" gridcol="1" gridrow="14" pageId="7" pageNumber="78">65</td>
<td id="C26A5DCCFF99000DFC05DCEAFC112FE5" box="[947,989,1236,1257]" gridcol="2" gridrow="14" pageId="7" pageNumber="78">63.9</td>
<td id="C26A5DCCFF99000DFBFDDCEAFB6C2FE5" box="[1099,1184,1236,1257]" gridcol="3" gridrow="14" pageId="7" pageNumber="78">61–66</td>
</tr>
</table>
</paragraph>
<paragraph id="3F3436F2FF99FFF5FF71DDBFFEBF2D42" blockId="7.[151,1437,1409,2013]" pageId="7" pageNumber="78">
<emphasis id="0DFFEAE0FF99FFF5FF71DDBFFC882E96" bold="true" box="[199,836,1409,1434]" pageId="7" pageNumber="78">
Counts and measurements of the
<typeStatus id="E0308850FF99FFF5FDDEDDBFFD032E96" box="[616,719,1409,1434]" pageId="7" pageNumber="78" type="holotype">holotype</typeStatus>
(in mm).
</emphasis>
Total length 559; head 51.4; trunk 197.6; tail 310; predorsal distance 87; pectoral-fin length 10.0; pectoral-fin base 4.4; body depth
<emphasis id="0DFFEAE0FF99FFF5FBFDDD98FBA02EB1" box="[1099,1132,1446,1469]" italics="true" pageId="7" pageNumber="78">ca.</emphasis>
18 at gill openings; body width
<emphasis id="0DFFEAE0FF99FFF5FF68DDF4FECC2EED" box="[222,256,1482,1505]" italics="true" pageId="7" pageNumber="78">ca.</emphasis>
14.5 at gill openings; body depth at anus
<emphasis id="0DFFEAE0FF99FFF5FD6BDDF4FD332EED" box="[733,767,1482,1505]" italics="true" pageId="7" pageNumber="78">ca.</emphasis>
18.5; body width at anus
<emphasis id="0DFFEAE0FF99FFF5FB91DDF4FB852EED" box="[1063,1097,1482,1505]" italics="true" pageId="7" pageNumber="78">ca.</emphasis>
16.5; body depth at branchial basket
<emphasis id="0DFFEAE0FF99FFF5FF53DDD0FECB2D09" box="[229,263,1518,1541]" italics="true" pageId="7" pageNumber="78">ca.</emphasis>
20; snout 10.6; tip of snout to rictus 19.4; snout overhang beyond tip of lower jaw 1.3; eye diameter 6.3; interorbital width 5.8; gill opening height 6.7; isthmus width 10.8. Lateral-line pores 67, total pores 154. Vertebral formula 19-65-160.
</paragraph>
</subSubSection>
<subSubSection id="77916579FF99FFFBFF71DE67FD382A68" lastPageId="9" lastPageNumber="80" pageId="7" pageNumber="78" type="description">
<paragraph id="3F3436F2FF99FFF5FF71DE67FCAD2CD1" blockId="7.[151,1437,1409,2013]" pageId="7" pageNumber="78">
<emphasis id="0DFFEAE0FF99FFF5FF71DE67FE952D7E" bold="true" box="[199,345,1625,1650]" pageId="7" pageNumber="78">Description.</emphasis>
Body moderately elongate (
<figureCitation id="A7B02A77FF99FFF5FD2ADE67FD2D2D7E" box="[668,737,1625,1650]" captionStart="FIGURE 4" captionStartId="6.[151,244,1933,1954]" captionTargetBox="[388,1198,201,1884]" captionTargetId="figure@6.[388,1198,190,1904]" captionTargetPageId="6" captionText="FIGURE 4. Ophichthus bicolor sp. nov. A. NMMB-P 18103, holotype, 559 mm TL, preserved. B. NMMB-P 22212, paratype, 696 mm TL, fresh. C. NMMB-P 17544, paratype, 701 mm TL, fresh. Bars indicate the origins of the dorsal fin (left) and anal fin (right)." httpUri="https://zenodo.org/record/233210/files/figure.png" pageId="7" pageNumber="78">Fig. 4</figureCitation>
), subcircular to posterior portion of tail, then becoming more compressed, its depth at gill openings
<quantity id="F8739B17FF99FFF5FDE2DE40FD762D9A" box="[596,698,1661,1686]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" metricValueMax="9.652" metricValueMin="6.096" pageId="7" pageNumber="78" unit="in" value="31.0" valueMax="38.0" valueMin="24.0">24–38 in</quantity>
TL. Branchial basket slightly expanded. Head 3.3–4.0 in trunk. Head and trunk 2.1–2.3 and head
<quantity id="F8739B17FF99FFF5FD91DE9FFD642DB6" box="[551,680,1697,1722]" metricMagnitude="-1" metricUnit="m" metricValue="2.6543" metricValueMax="2.9463999999999997" metricValueMin="2.3622" pageId="7" pageNumber="78" unit="in" value="10.45" valueMax="11.6" valueMin="9.3">9.3–11.6 in</quantity>
TL. Snout rounded, moderately acute when viewed from above; underside of snout not bisected by a groove. Lower jaw included, its tip reaching to middle of anterior nostril tube. Upper jaw moderately elongated, rictus well behind a vertical from posterior margin of eye. Eye moderate, slightly behind center of upper jaw,
<quantity id="F8739B17FF99FFF5FE45DF30FDA72C2A" box="[499,619,1805,1830]" metricMagnitude="-2" metricUnit="m" metricValue="8.763" metricValueMax="9.652" metricValueMin="7.874" pageId="7" pageNumber="78" unit="in" value="3.45" valueMax="3.8" valueMin="3.1">3.1–3.8 in</quantity>
upper jaw and
<quantity id="F8739B17FF99FFF5FC82DF33FC692C2A" box="[820,933,1805,1830]" metricMagnitude="-1" metricUnit="m" metricValue="2.0955" metricValueMax="2.54" metricValueMin="1.651" pageId="7" pageNumber="78" unit="in" value="8.25" valueMax="10.0" valueMin="6.5">6.5–10 in</quantity>
head. Anterior nostrils tubular, extending ventrolaterally from snout at
<emphasis id="0DFFEAE0FF99FFF5FE56DF0BFDCD2C40" box="[480,513,1845,1868]" italics="true" pageId="7" pageNumber="78">ca.</emphasis>
80o, reaching below upper lip but not reaching tip of chin when directed forward. Posterior nostril a hole above upper lip, covered by a flap that extends below edge of mouth. Barbels absent from upper lip. Dorsal-fin origin well behind pectoral fin by more than 2 pectoral-fin lengths. Median fins low but obvious, ending approximately an eye diameter before the bluntly pointed tail tip. Pectoral fins pointed, not elongate and lanceolate, the longest rays slightly above mid fin.
</paragraph>
<paragraph id="3F3436F2FF96FFFAFF71D8A9FEF92A69" blockId="8.[151,1437,151,536]" pageId="8" pageNumber="79">
Head pores small but apparent (
<figureCitation id="A7B02A77FF96FFFAFD9BD8A9FDBE2BBC" box="[557,626,151,176]" captionStart="FIGURE 5" captionStartId="8.[151,244,955,976]" captionTargetBox="[261,1342,598,938]" captionTargetId="figure@8.[250,1342,587,943]" captionTargetPageId="8" captionText="FIGURE 5. Semi-diagrammatic illustration of lateral head of Ophichthus bicolor McCosker &amp; Ho, sp. nov., from the holotype. Arrows point to the frontal pore (left) and median pore of supratemporal series (right)." httpUri="https://zenodo.org/record/233211/files/figure.png" pageId="8" pageNumber="79">Fig. 5</figureCitation>
). Single median interorbital and temporal pores. Supraorbital pores 1+4 (occasionally 1+3), infraorbital pores 4+2, lower jaw pores 5–7 (mainly 6), preopercular pores 2, supratemporal pores 3. Faint rows of minute sensory papillae are present along nape, along anterior margin of orbit, and in a horseshoe-shaped pattern around base of anterior nostril. Lateral-line pores apparent; 7–9 before gill opening in an arching sequence, 18–22 before dorsal-fin origin; 63–67 before anus,
<quantity id="F8739B17FF96FFFAFC10D919FBE92A4C" box="[934,1061,295,320]" metricMagnitude="0" metricUnit="m" metricValue="3.9497" metricValueMax="4.013199999999999" metricValueMin="3.8862" pageId="8" pageNumber="79" unit="in" value="155.5" valueMax="158.0" valueMin="153.0">153–158 in</quantity>
total, the last
<emphasis id="0DFFEAE0FF96FFFAFB75D917FB292A4C" box="[1219,1253,297,320]" italics="true" pageId="8" pageNumber="79">ca.</emphasis>
2 eye diameters before tail tip.
</paragraph>
<paragraph id="3F3436F2FF96FFFAFF71D951FB482914" blockId="8.[151,1437,151,536]" pageId="8" pageNumber="79">
Teeth (
<figureCitation id="A7B02A77FF96FFFAFEA1D951FE912A84" box="[279,349,367,392]" captionStart="FIGURE 6" captionStartId="8.[151,244,1773,1794]" captionTargetBox="[259,1336,1064,1751]" captionTargetId="figure@8.[253,1345,1057,1752]" captionTargetPageId="8" captionText="FIGURE 6. Semi-diagrammatic illustration of dentition of Ophichthus bicolor McCosker &amp; Ho, sp. nov., from the holotype. A. Upper Jaw. B. Lower Jaw" httpUri="https://zenodo.org/record/233212/files/figure.png" pageId="8" pageNumber="79">Fig. 6</figureCitation>
) fang-like, none enlarged, conical, slightly retrorse, and not closely spaced. An intermaxillary rosette of 4–5 teeth followed by an intermaxillary patch of 3 pairs followed by 6–8 uniserial vomerine teeth which decrease slightly in size posteriorly. Maxillary teeth biserial,
<quantity id="F8739B17FF96FFFAFCD7D986FC752ADC" box="[865,953,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.54" metricValueMin="1.778" pageId="8" pageNumber="79" unit="in" value="8.5" valueMax="10.0" valueMin="7.0">7–10 in</quantity>
inner tooth row and
<quantity id="F8739B17FF96FFFAFB01D986FAD02ADC" box="[1207,1308,439,464]" metricMagnitude="-1" metricUnit="m" metricValue="3.175" metricValueMax="3.556" metricValueMin="2.794" pageId="8" pageNumber="79" unit="in" value="12.5" valueMax="14.0" valueMin="11.0">11–14 in</quantity>
outer row. Mandibular teeth uniserial posteriorly; an anterior patch of 3 pairs of teeth at symphysis, followed by 1–3 pairs of teeth; outer row of 13–17 widely spaced teeth on each side, decreasing in size posteriorly.
</paragraph>
<caption id="6BF4667AFF96FFFAFF21DB85FC1F28FF" httpUri="https://zenodo.org/record/233211/files/figure.png" pageId="8" pageNumber="79" targetBox="[261,1342,598,938]" targetPageId="8">
<paragraph id="3F3436F2FF96FFFAFF21DB85FC1F28FF" blockId="8.[151,1437,955,1011]" pageId="8" pageNumber="79">
<emphasis id="0DFFEAE0FF96FFFAFF21DB85FEC328DC" bold="true" box="[151,271,955,976]" pageId="8" pageNumber="79">FIGURE 5.</emphasis>
Semi-diagrammatic illustration of lateral head of
<taxonomicName id="F88B4D71FF96FFFAFCB1DB85FBB828DC" authority="McCosker &amp; Ho" authorityName="McCosker &amp; Ho" box="[775,1140,955,976]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="8" pageNumber="79" phylum="Chordata" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="0DFFEAE0FF96FFFAFCB1DB85FC0828DC" box="[775,964,955,976]" italics="true" pageId="8" pageNumber="79">Ophichthus bicolor</emphasis>
McCosker &amp; Ho
</taxonomicName>
,
<taxonomicNameLabel id="16CC579BFF96FFFAFB32DB85FB1828DC" box="[1156,1236,955,976]" pageId="8" pageNumber="79" rank="species">
<emphasis id="0DFFEAE0FF96FFFAFB32DB85FB1828DC" bold="true" box="[1156,1236,955,976]" pageId="8" pageNumber="79">sp. nov.</emphasis>
</taxonomicNameLabel>
, from the holotype. Arrows point to the frontal pore (left) and median pore of supratemporal series (right).
</paragraph>
</caption>
<caption id="6BF4667AFF96FFFAFF21DED3FE422C2C" httpUri="https://zenodo.org/record/233212/files/figure.png" pageId="8" pageNumber="79" targetBox="[259,1336,1064,1751]" targetPageId="8">
<paragraph id="3F3436F2FF96FFFAFF21DED3FE422C2C" blockId="8.[151,1436,1773,1824]" pageId="8" pageNumber="79">
<emphasis id="0DFFEAE0FF96FFFAFF21DED3FEC32C0E" bold="true" box="[151,271,1773,1794]" pageId="8" pageNumber="79">FIGURE 6.</emphasis>
Semi-diagrammatic illustration of dentition of
<taxonomicName id="F88B4D71FF96FFFAFD5FDED3FB9B2C0E" authority="McCosker &amp; Ho" authorityName="McCosker &amp; Ho" box="[745,1111,1773,1794]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="8" pageNumber="79" phylum="Chordata" rank="species" species="bicolor" status="sp. nov.">
<emphasis id="0DFFEAE0FF96FFFAFD5FDED3FC692C0E" box="[745,933,1773,1794]" italics="true" pageId="8" pageNumber="79">Ophichthus bicolor</emphasis>
McCosker &amp; Ho
</taxonomicName>
,
<taxonomicNameLabel id="16CC579BFF96FFFAFBD2DED3FB782C0E" box="[1124,1204,1773,1794]" pageId="8" pageNumber="79" rank="species">
<emphasis id="0DFFEAE0FF96FFFAFBD2DED3FB782C0E" bold="true" box="[1124,1204,1773,1794]" pageId="8" pageNumber="79">sp. nov.</emphasis>
</taxonomicNameLabel>
, from the holotype. A. Upper Jaw. B. Lower Jaw
</paragraph>
</caption>
<paragraph id="3F3436F2FF96FFFBFF71DF5DFE742A4C" blockId="8.[151,1437,1891,2024]" lastBlockId="9.[151,1437,151,1005]" lastPageId="9" lastPageNumber="80" pageId="8" pageNumber="79">
Coloration when fresh (
<figureCitation id="A7B02A77FF96FFFAFE65DF5DFDDB2C77" box="[467,535,1891,1916]" captionStart="FIGURE 4" captionStartId="6.[151,244,1933,1954]" captionTargetBox="[388,1198,201,1884]" captionTargetId="figure@6.[388,1198,190,1904]" captionTargetPageId="6" captionText="FIGURE 4. Ophichthus bicolor sp. nov. A. NMMB-P 18103, holotype, 559 mm TL, preserved. B. NMMB-P 22212, paratype, 696 mm TL, fresh. C. NMMB-P 17544, paratype, 701 mm TL, fresh. Bars indicate the origins of the dorsal fin (left) and anal fin (right)." httpUri="https://zenodo.org/record/233210/files/figure.png" pageId="8" pageNumber="79">Fig. 4</figureCitation>
) yellowish-brown dorsally, strongly contrasting with the white throat and belly, extending to the anus. Dorsal and anal fins yellowish or pinkish with narrow brown margins. Pectoral fin yellowish or pinkish, with a brown patch at its base. Nostrils white, contrasting with brown snout and upper lips. Coloration in preservative yellowish-brown dorsally, strongly contrasting with white throat and belly extending to anus. A yellowish-orange band flanks pale anal fin. Dorsal fin white, flanked by narrow brown bands. Pectoral fin pale, a brown smudge centrally behind base. Nostrils white, contrasting with brown snout and upper lip. Head brown dorsally, fading behind dorsal-fin origin. Lower jaw with brown bands overlying mandibular pore series. Gums pale. Palate and tongue with a speckling of minute dark spots. Peritoneum pale with fine dark speckling along upper wall. Tail tip white.
</paragraph>
<paragraph id="3F3436F2FF97FFFBFF71D972FD382A68" blockId="9.[151,1437,151,1005]" box="[199,756,332,357]" pageId="9" pageNumber="80">
<emphasis id="0DFFEAE0FF97FFFBFF71D972FF322A69" bold="true" box="[199,254,332,357]" pageId="9" pageNumber="80">Size.</emphasis>
The largest known specimen is
<quantity id="F8739B17FF97FFFBFDD1D972FD082A68" box="[615,708,332,356]" metricMagnitude="-1" metricUnit="m" metricValue="9.27" pageId="9" pageNumber="80" unit="mm" value="927.0">927 mm</quantity>
TL.
</paragraph>
</subSubSection>
<subSubSection id="77916579FF97FFFBFF71D951FA512A84" box="[199,1437,367,392]" pageId="9" pageNumber="80" type="etymology">
<paragraph id="3F3436F2FF97FFFBFF71D951FA512A84" blockId="9.[151,1437,151,1005]" box="[199,1437,367,392]" pageId="9" pageNumber="80">
<emphasis id="0DFFEAE0FF97FFFBFF71D951FE822A84" bold="true" box="[199,334,367,392]" pageId="9" pageNumber="80">Etymology.</emphasis>
From Latin, in reference to its sharply contrasting coloration, to be treated as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="77916579FF97FFFBFF71D9AAFDE42ADC" pageId="9" pageNumber="80" type="distribution">
<paragraph id="3F3436F2FF97FFFBFF71D9AAFDE42ADC" blockId="9.[151,1437,151,1005]" pageId="9" pageNumber="80">
<emphasis id="0DFFEAE0FF97FFFBFF71D9AAFEAC2AA1" bold="true" box="[199,352,404,429]" pageId="9" pageNumber="80">Distribution.</emphasis>
Known from the
<typeStatus id="E0308850FF97FFFBFD82D9ABFDA82AA1" box="[564,612,405,429]" pageId="9" pageNumber="80">type</typeStatus>
series, collected from off Dong-gang, SW
<collectingCountry id="479C7662FF97FFFBFBEAD9AAFB622AA1" box="[1116,1198,404,429]" name="Taiwan" pageId="9" pageNumber="80">Taiwan</collectingCountry>
and off Taitung, SE
<collectingCountry id="479C7662FF97FFFBFF21D989FF202ADC" box="[151,236,439,464]" name="Taiwan" pageId="9" pageNumber="80">Taiwan</collectingCountry>
, from
<emphasis id="0DFFEAE0FF97FFFBFE82D987FE9A2ADC" box="[308,342,441,464]" italics="true" pageId="9" pageNumber="80">ca.</emphasis>
<quantity id="F8739B17FF97FFFBFEEAD986FE172AC3" box="[348,475,440,464]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="9" pageNumber="80" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200–400 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="77916579FF97FFFBFF71D9E2FD5428E0" pageId="9" pageNumber="80" type="discussion">
<paragraph id="3F3436F2FF97FFFBFF71D9E2FE522818" blockId="9.[151,1437,151,1005]" pageId="9" pageNumber="80">
<emphasis id="0DFFEAE0FF97FFFBFF71D9E2FEF72AF9" bold="true" box="[199,315,476,501]" pageId="9" pageNumber="80">Remarks.</emphasis>
This new species would not be mistaken for most other
<taxonomicName id="F88B4D71FF97FFFBFC63D9E2FB9A2AF9" box="[981,1110,476,501]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="0DFFEAE0FF97FFFBFC63D9E2FB9A2AF9" box="[981,1110,476,501]" italics="true" pageId="9" pageNumber="80">Ophichthus</emphasis>
</taxonomicName>
from
<collectingCountry id="479C7662FF97FFFBFB17D9E2FB3F2AF9" box="[1185,1267,476,501]" name="Taiwan" pageId="9" pageNumber="80">Taiwan</collectingCountry>
because of its posterior dorsal-fin origin, dentition, jaw length, eye position, and its distinctive body coloration. It is among the largest species of Taiwanese
<taxonomicName id="F88B4D71FF97FFFBFE5BDA1AFDA22931" box="[493,622,548,573]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="genus">
<emphasis id="0DFFEAE0FF97FFFBFE5BDA1AFDA22931" box="[493,622,548,573]" italics="true" pageId="9" pageNumber="80">Ophichthus</emphasis>
</taxonomicName>
, reaching nearly a meter in length. The new species is similar to
<taxonomicName id="F88B4D71FF97FFFBFA37DA1BFE55296C" authority="Asano, 1987" authorityName="Asano" authorityYear="1987" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="species" species="megalops">
<emphasis id="0DFFEAE0FF97FFFBFA37DA1BFECF296C" italics="true" pageId="9" pageNumber="80">
<collectingCountry id="479C7662FF97FFFBFA37DA1BFA542930" box="[1409,1432,549,572]" name="Iceland" pageId="9" pageNumber="80">O</collectingCountry>
. megalops
</emphasis>
<bibRefCitation id="5B1A4B03FF97FFFBFEBFDA76FE55296C" author="Asano" box="[265,409,584,608]" pageId="9" pageNumber="80" refString="Asano, H. (1987) A new ophichthid eel, Ophichthus megalops, from the Kumano-nada, Japan. Japanese Journal of Ichthyology, 34, 135 - 137." type="journal article" year="1987">Asano, 1987</bibRefCitation>
</taxonomicName>
, known from
<collectingCountry id="479C7662FF97FFFBFD82DA76FDB82953" box="[564,628,584,607]" name="Japan" pageId="9" pageNumber="80">Japan</collectingCountry>
and
<collectingCountry id="479C7662FF97FFFBFD1FDA79FD33296C" box="[681,767,583,608]" name="Taiwan" pageId="9" pageNumber="80">Taiwan</collectingCountry>
. It differs from
<taxonomicName id="F88B4D71FF97FFFBFC1BDA79FBD4296C" box="[941,1048,583,608]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="species" species="megalops">
<emphasis id="0DFFEAE0FF97FFFBFC1BDA79FBD4296C" box="[941,1048,583,608]" italics="true" pageId="9" pageNumber="80">megalops</emphasis>
</taxonomicName>
in the more anterior location of its dorsal-fin origin (15–19% of TL behind snout tip vs. 24–26%), its mean pre-dorsal vertebral number (18 vs. 29), in its preopercular pores (2 vs. 3), in its posterior anal-fin coloration (pale vs. black), and in having fewer maxillary teeth (8–11 inner row and 11–14 outer row vs. 25–30 inner row and 25–30 outer row teeth). They are quite similar in their tail and head proportions, total vertebral numbers (155–163 vs. 161–163), and in their body depth (25–31 vs. 25–29 times in TL).
</paragraph>
<paragraph id="3F3436F2FF97FFFBFF71DB21FD5428E0" blockId="9.[151,1437,151,1005]" pageId="9" pageNumber="80">
The new species is also similar to
<taxonomicName id="F88B4D71FF97FFFBFDDADB21FB022834" authority="Liu, Tang &amp; Zhang, 2010" authorityName="Liu, Tang &amp; Zhang" authorityYear="2010" box="[620,1230,799,824]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="species" species="retrodorsalis">
<emphasis id="0DFFEAE0FF97FFFBFDDADB21FC462834" box="[620,906,799,824]" italics="true" pageId="9" pageNumber="80">Ophichthus retrodorsalis</emphasis>
<bibRefCitation id="5B1A4B03FF97FFFBFC21DB21FB022834" author="Liu" box="[919,1230,799,824]" pageId="9" pageNumber="80" refString="Liu, D., Tang, W. &amp; Zhang, C. (2010) Ophichthus retrodorsalis sp. nov. In: Zhang, C. (Ed.), Fauna Sinica. Osteichthyes: Anguilliformes and Notacanthiformes. Science Press, Beijing, pp. 332 - 333." type="book chapter" year="2010">Liu, Tang &amp; Zhang, 2010</bibRefCitation>
</taxonomicName>
, known from the
<typeStatus id="E0308850FF97FFFBFF21DB7AFF352851" box="[151,249,836,861]" pageId="9" pageNumber="80" type="holotype">holotype</typeStatus>
collected from Fuzhou, Fujian Province,
<collectingCountry id="479C7662FF97FFFBFD64DB7AFCD52851" box="[722,793,836,861]" name="China" pageId="9" pageNumber="80">China</collectingCountry>
. It has a similar dorsal-fin origin and its dentition is also multiserial. One of us (HCH) examined the
<typeStatus id="E0308850FF97FFFBFD36DB59FD2E288C" box="[640,738,871,896]" pageId="9" pageNumber="80" type="holotype">holotype</typeStatus>
of
<taxonomicName id="F88B4D71FF97FFFBFCB1DB56FC75288C" box="[775,953,871,896]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="species" species="retrodorsalis">
<emphasis id="0DFFEAE0FF97FFFBFCB1DB56FC75288C" box="[775,953,871,896]" italics="true" pageId="9" pageNumber="80">
<collectingCountry id="479C7662FF97FFFBFCB1DB56FCD22873" box="[775,798,872,895]" name="Iceland" pageId="9" pageNumber="80">O</collectingCountry>
. retrodorsalis
</emphasis>
</taxonomicName>
and found it to differ from
<taxonomicName id="F88B4D71FF97FFFBFB5DDB56FA92288C" box="[1259,1374,871,896]" class="Actinopterygii" family="Ophichthidae" genus="Ophichthus" kingdom="Animalia" order="Anguilliformes" pageId="9" pageNumber="80" phylum="Chordata" rank="species" species="bicolor">
<emphasis id="0DFFEAE0FF97FFFBFB5DDB56FA92288C" box="[1259,1374,871,896]" italics="true" pageId="9" pageNumber="80">
<collectingCountry id="479C7662FF97FFFBFB5DDB56FACE2873" box="[1259,1282,872,895]" name="Iceland" pageId="9" pageNumber="80">O</collectingCountry>
. bicolor
</emphasis>
</taxonomicName>
in its coloration (uniformly light brown), pectoral fins (dark), body depth (50 times in TL at gill opening), head length (14.7 times in TL), eye position (posterior margin of orbit above rictus) and in its dentition (triserial throughout). We were unable to obtain its vertebral counts.
</paragraph>
</subSubSection>
</treatment>
</document>
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