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<document id="870C3A42357091E2B639FDD4DF3F2AA3" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1626381872300" checkinUser="felipe" docAuthor="Oláh, János, Johanson, Kjell Arne &amp; Barnard, Peter C." docDate="2008" docId="627D87E1FF94F770FF7EFDBCFD247A9A" docLanguage="en" docName="zt01738p171.pdf" docOrigin="Zootaxa 1738" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Cheumatopsyche andohahela Olah &amp; Johanson 2008, new species" docType="treatment" docVersion="2" lastPageNumber="107" masterDocId="9E44FF99FFFEF71AFFE9FF9BFFD07D08" masterDocTitle="Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)" masterLastPageNumber="171" masterPageNumber="1" pageNumber="107" updateTime="1699011095215" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="7ADBEBAF4BAD6D6A5E8E257CF0F929D1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B7AF57D3825F4587FEAA96E8F4B7445B">
<mods:title id="83CBCA1B59741B5955D23E1215717B37">Revision of the Oriental and Afrotropical species of Cheumatopsyche Wallengren (Hydropsychidae, Trichoptera)</mods:title>
</mods:titleInfo>
<mods:name id="F7BAFE026F3E50D3EAB15ABD90759454" type="personal">
<mods:role id="68072ECC28F75098788CC4AE862989C2">
<mods:roleTerm id="E35BF59B5568CAB8E7D3F60774BD6298">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="05BA07EE40A9FD241572CBD727554791">Oláh, János</mods:namePart>
</mods:name>
<mods:name id="F0F766C7F0FBC6DDEF940F5BF952318D" type="personal">
<mods:role id="719995036E29553CD9B90475CACA6460">
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</mods:role>
<mods:namePart id="8B3DA8AA79B42023E74EA54475CD7BE1">Johanson, Kjell Arne</mods:namePart>
</mods:name>
<mods:name id="2B706D1DE16304ED3621D374A05B08E9" type="personal">
<mods:role id="AEBE1174DBEC7CA5277B47A2FA72D221">
<mods:roleTerm id="759B966A52D03460C77BAC3FE845071D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="387C3AEFB2F31463DF10DAC7F19A26E8">Barnard, Peter C.</mods:namePart>
</mods:name>
<mods:typeOfResource id="2133F7D8FC4935F6C20D9DCB22954AA4">text</mods:typeOfResource>
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<mods:titleInfo id="CD9E623A14ABB40E355DAD86CC8DE3F9">
<mods:title id="AFC0A630156795553C10D56BA4B95346">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="AAF95097F9F27C3E116A9EC0767A710B">
<mods:date id="5C649F16B1259849220BE2E54A3139CF">2008</mods:date>
<mods:detail id="5AF6EF8930615B2C1343941FA215D414" type="pubDate">
<mods:number id="847E21F4D7D287E7C2F8403C1D2694A6">2008-03-31</mods:number>
</mods:detail>
<mods:detail id="0A700147AB6FDFAF419133F333F481EA" type="volume">
<mods:number id="F9CDED6816218020ABDB5FA7802A0CA1">1738</mods:number>
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<mods:start id="FB5BF61A66C8698F34754AA5BC8D15F7">1</mods:start>
<mods:end id="E1819AA54BF6CD2DF10D54E4615BF104">171</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="44AF6CB68EEA701B6997CDF00C6060C6">journal article</mods:classification>
<mods:identifier id="7EB13BE722AF546FD3B5F88800757740" type="ISSN">1175­5334</mods:identifier>
</mods:mods>
<treatment id="627D87E1FF94F770FF7EFDBCFD247A9A" LSID="urn:lsid:plazi:treatment:627D87E1FF94F770FF7EFDBCFD247A9A" httpUri="http://treatment.plazi.org/id/627D87E1FF94F770FF7EFDBCFD247A9A" lastPageNumber="107" pageId="106" pageNumber="107">
<subSubSection id="A2CE657CFF94F770FF7EFDBCFCAE7F49" box="[151,894,551,577]" pageId="106" pageNumber="107" type="nomenclature">
<paragraph id="EA6B36F7FF94F770FF7EFDBCFCAE7F49" blockId="106.[151,894,551,618]" box="[151,894,551,577]" pageId="106" pageNumber="107">
<heading id="B123819BFF94F770FF7EFDBCFCAE7F49" bold="true" box="[151,894,551,577]" fontSize="11" level="1" pageId="106" pageNumber="107" reason="1">
<emphasis id="D8A0EAE5FF94F770FF7EFDBCFCAE7F49" bold="true" box="[151,894,551,577]" pageId="106" pageNumber="107">
<taxonomicName id="2DD44D74FF94F770FF7EFDBCFD307F49" authority="Olah &amp; Johanson" authorityName="Olah &amp; Johanson" authorityYear="2008" box="[151,736,551,577]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="andohahela" status="sp. nov.">
<emphasis id="D8A0EAE5FF94F770FF7EFDBCFE2C7F49" bold="true" box="[151,508,551,577]" italics="true" pageId="106" pageNumber="107">Cheumatopsyche andohahela</emphasis>
Oláh &amp; Johanson
</taxonomicName>
,
<taxonomicNameLabel id="C393579EFF94F770FD07FDBCFCAE7F49" box="[750,894,551,577]" pageId="106" pageNumber="107" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF94F770FF7EFDCBFBA87A12" pageId="106" pageNumber="107" type="description">
<paragraph id="EA6B36F7FF94F770FF7EFDCBFEE37F62" blockId="106.[151,894,551,618]" box="[151,307,592,618]" pageId="106" pageNumber="107">
<figureCitation id="72EF2A72FF94F770FF7EFDCBFEE37F62" box="[151,307,592,618]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260–263</figureCitation>
</paragraph>
<paragraph id="EA6B36F7FF94F770FF7EFD3BFAC97E8A" blockId="106.[151,1436,672,1938]" pageId="106" pageNumber="107">
This species is similar to
<emphasis id="D8A0EAE5FF94F770FE55FD3BFDED7FB1" box="[444,573,672,697]" italics="true" pageId="106" pageNumber="107">
<taxonomicName id="2DD44D74FF94F770FE55FD3BFDE97FB1" authority="Oláh &amp; Johanson &amp; Barnard, 2008" authorityName="Oláh &amp; Johanson &amp; Barnard" authorityYear="2008" box="[444,569,672,697]" class="Insecta" family="Hydropsychidae" genus="Cheumatopsyche" kingdom="Animalia" order="Trichoptera" pageId="106" pageNumber="107" phylum="Arthropoda" rank="species" species="padaha" status="sp. nov.">C. padaha</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="C393579EFF94F770FDADFD3BFD007FB2" box="[580,720,672,698]" pageId="106" pageNumber="107" rank="species">new species</taxonomicNameLabel>
but easily distinguished by the presence of strongly modified harpagones. The upper branches of harpagones are basally broad, with a needle-sharp, dorsal tip being almost right-angled to the dorsal margin of the proximal part of harpagones, easily seen in lateral view. The lower branches of harpagones are broad, forming the blunt apices in ventral and dorsal view. The ventral branch is also covered by short, strong setae. The setose dorsobasal part of segment X is present but forms a pair of small, setose protuberances at the intersegmental dorsal region. The species is very large and dark.
</paragraph>
<paragraph id="EA6B36F7FF94F770FF2EFC0BFD8D797A" blockId="106.[151,1436,672,1938]" pageId="106" pageNumber="107">Male. Body and wings brown with brown pubescence; wings without spots or pattern. Maxillary palp segment I shortest, segment II longer, segments III and IV equally long, segment V as long as sum of segments I– IV. Head dorsum brown, with 9 similarly coloured warts and six small wart spots. Antennae about 1.2 times longer than forewings. Proepisternum without swollen setal wart. Protarsal claws asymmetrical, laterally flanked by setal bundle. Forewing length 16.0 mm, hind wing length 10.0 mm. Forewing crossveins m-cu and cu tangental. Hind wing fork I present.</paragraph>
<paragraph id="EA6B36F7FF94F770FF2EFB1BFBA87A12" blockId="106.[151,1436,672,1938]" pageId="106" pageNumber="107">
Male genitalia. Segment IX fused annularly; sternum about 2 times longer than tergum (
<figureCitation id="72EF2A72FF94F770FB0AFB1BFA9B7992" box="[1251,1355,1152,1178]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
); anterior margins of segment IX rounded; apical lobe on posterolateral margins well developed, located immediately above coxopodite bases; spine row on posterior margins of segment IX complete; spines long and irregularly bent on dorsum, straight, spines on apical lobe shortest. Intersegmental step between segment IX and segment X occupied by pair of setose protuberances (
<figureCitation id="72EF2A72FF94F770FCA8FABBFC797832" box="[833,937,1312,1338]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
). Segment X long, distal half curving dorsad; ventral margin strongly convex; apex blunt in lateral view (
<figureCitation id="72EF2A72FF94F770FC77FAD3FBD8786A" box="[926,1032,1352,1378]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
), simple; tongue-shaped in dorsal view. Large apicoventral setal lobes dominating distal half of segment X; completely fused with segment X, except at apices. Setaless mesocaudal lobe reduced. Dorsal interlobular gap narrow, tongue-shaped, deep. Setose areas present at dorsobasal part of segment X, forming pair of small protuberances visible in both lateral and dorsal view (
<figureCitation id="72EF2A72FF94F770FE81FA73FDDF7B0A" box="[360,527,1512,1538]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260, 261</figureCitation>
). Transverse sutures not visible. Longitudinal sutures running along ventral margin of segment (
<figureCitation id="72EF2A72FF94F770FE7AF98BFDD07B22" box="[403,512,1552,1578]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
). Lateral setose areas forming pair of elevated, vertically elongating warts located centrally on segment X (
<figureCitation id="72EF2A72FF94F770FDF3F9A3FD527B5A" box="[538,642,1592,1618]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
). Coxopodites robust, basal half thinner than distal half (
<figureCitation id="72EF2A72FF94F770FACEF9A3FA5F7B5A" box="[1319,1423,1592,1618]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
), strongly broadening from mid-length; slightly exceeding apex of segment X. Harpagones with bilobed apices (
<figureCitation id="72EF2A72FF94F770FF76F913FEDA7BAA" box="[159,266,1672,1698]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 262</figureCitation>
); upper branches forming needle-sharpened dorsal processes being almost right-angled to dorsal margin of proximal part of harpagones (
<figureCitation id="72EF2A72FF94F770FD80F92BFD017BC2" box="[617,721,1712,1738]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
); both lower branches blunt, rounded, covered with short and stout setae (
<figureCitation id="72EF2A72FF94F770FEC3F943FE467BFA" box="[298,406,1752,1778]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 260</figureCitation>
). Phallotheca (
<figureCitation id="72EF2A72FF94F770FDA3F943FD627BFA" box="[586,690,1752,1778]" captionStart="FIGURES 260–263" captionStartId="107.[151,269,1611,1635]" captionTargetBox="[170,1391,209,1563]" captionTargetId="figure-120@107.[151,1436,194,1587]" captionTargetPageId="107" captionText="FIGURES 260–263. Cheumatopsyche andohahela Oláh &amp; Johanson, new species, holotype. 260 — genitalia, lateral; 261 — genitalia, dorsal; 262 — gonocoxite, ventral; 263 — phallus, lateral." pageId="106" pageNumber="107">Fig. 263</figureCitation>
) with small phallobase; slightly curving posterad, sclerotised endothecal process circular; phallotremal sclerites broad and rounded in lateral view.
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF94F770FF2EF8BCFA857A4A" box="[199,1365,1831,1858]" pageId="106" pageNumber="107" type="materials_examined">
<paragraph id="EA6B36F7FF94F770FF2EF8BCFA857A4A" blockId="106.[151,1436,672,1938]" box="[199,1365,1831,1858]" pageId="106" pageNumber="107">
<materialsCitation id="5ABC3CAAFF94F770FF2EF8BCFA857A4A" box="[199,1365,1831,1858]" collectingDate="1956-01" collectionCode="MNHN" collectorName="R. Paulian" country="Madagascar" elevation="1800" location="Andohahela" pageId="106" pageNumber="107" specimenCount="1" typeStatus="holotype">
<emphasis id="D8A0EAE5FF94F770FF2EF8BCFDB47A49" bold="true" box="[199,612,1831,1857]" pageId="106" pageNumber="107">
<typeStatus id="356F8855FF94F770FF2EF8BCFEE97A49" box="[199,313,1831,1857]" pageId="106" pageNumber="107" type="holotype">Holotype</typeStatus>
male:
<collectingCountry id="92C37667FF94F770FE64F8BCFDB47A49" box="[397,612,1831,1857]" name="Madagascar" pageId="106" pageNumber="107">MADAGASCAR</collectingCountry>
</emphasis>
:
<location id="EF0B602CFF94F770FD9DF8B3FCD57A4A" LSID="urn:lsid:plazi:treatment:627D87E1FF94F770FF7EFDBCFD247A9A:EF0B602CFF94F770FD9DF8B3FCD57A4A" box="[628,773,1832,1858]" country="Madagascar" name="Andohahela" pageId="106" pageNumber="107">Andohahela</location>
,
<quantity id="2D2C9B12FF94F770FCF9F8B3FCBB7A4A" box="[784,875,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="106" pageNumber="107" unit="m" value="1800.0">
<elevation id="61F9D1C4FF94F770FCF9F8B3FCBB7A4A" box="[784,875,1832,1858]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="106" pageNumber="107" unit="m" value="1800.0">1800 m</elevation>
</quantity>
,
<date id="9E6A1037FF94F770FC9EF8B3FC117A4A" box="[887,961,1832,1858]" pageId="106" pageNumber="107" value="1956-01">
<collectingDate id="8E2EE9DFFF94F770FC9EF8B3FC117A4A" box="[887,961,1832,1858]" pageId="106" pageNumber="107" value="1956-01">i.1956</collectingDate>
</date>
[
<collectorName id="47215321FF94F770FC38F8B3FB807A4A" box="[977,1104,1832,1858]" pageId="106" pageNumber="107">R. Paulian</collectorName>
] (
<collectionCode id="8CC5AE32FF94F770FB8CF8B3FB137A4A" box="[1125,1219,1832,1858]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="106" pageNumber="107">MNHN</collectionCode>
, in alcohol).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF94F770FF2EF8D4FDD57A62" box="[199,517,1871,1898]" pageId="106" pageNumber="107" type="distribution">
<paragraph id="EA6B36F7FF94F770FF2EF8D4FDD57A62" blockId="106.[151,1436,672,1938]" box="[199,517,1871,1898]" pageId="106" pageNumber="107">
<emphasis id="D8A0EAE5FF94F770FF2EF8D4FEB97A61" bold="true" box="[199,361,1871,1897]" pageId="106" pageNumber="107">Distribution:</emphasis>
<collectingCountry id="92C37667FF94F770FE98F8CBFDD17A62" box="[369,513,1872,1898]" name="Madagascar" pageId="106" pageNumber="107">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="A2CE657CFF94F770FF2EF8ECFD247A9A" box="[199,756,1911,1938]" pageId="106" pageNumber="107" type="etymology">
<paragraph id="EA6B36F7FF94F770FF2EF8ECFD247A9A" blockId="106.[151,1436,672,1938]" box="[199,756,1911,1938]" pageId="106" pageNumber="107">
<emphasis id="D8A0EAE5FF94F770FF2EF8ECFE877A99" bold="true" box="[199,343,1911,1937]" pageId="106" pageNumber="107">Etymology:</emphasis>
from the name of the
<typeStatus id="356F8855FF94F770FDB5F8E3FD5E7A9A" box="[604,654,1912,1938]" pageId="106" pageNumber="107">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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