496 lines
63 KiB
XML
496 lines
63 KiB
XML
<document ID-CLB-Dataset="56712" ID-DOI="http://dx.doi.org/10.3897/herpetozoa.35.e94920" ID-GBIF-Dataset="e8f01291-4827-458e-9575-32637e9020fb" ID-Pensoft-Pub="2682-955X-35-265" ID-Pensoft-UUID="E04369D7C9EB5294A6DD1780BD27542F" ID-ZooBank="29C2ED3FBC5E45E78E58FCC93B988455" ModsDocID="2682-955X-35-265" checkinTime="1671892101870" checkinUser="pensoft" docAuthor="Li, Mao-Liang, Ren, Jin-Long, Huang, Jun-Jie, Lyu, Zhi-Tong, Qi, Shuo, Jiang, Ke, Wang, Ying-Yong & Li, Jia-Tang" docDate="2022" docId="C3E61DC2F9B05389A33A6DE55B6983C7" docLanguage="en" docName="Herpetozoa 35: 265-282" docOrigin="Herpetozoa 35" docPubDate="2022-12-21" docSource="http://dx.doi.org/10.3897/herpetozoa.35.e94920" docTitle="Hebius sauteri" docType="treatment" docVersion="6" id="E04369D7C9EB5294A6DD1780BD27542F" lastPageNumber="265" masterDocId="E04369D7C9EB5294A6DD1780BD27542F" masterDocTitle="On the validity of Hebius sauteri maximus (Malnate, 1962) (Squamata, Natricidae), with the redescription of H. maximus comb. nov. and H. sauteri (Boulenger, 1909)" masterLastPageNumber="282" masterPageNumber="265" pageNumber="265" updateTime="1732848484904" updateUser="ExternalLinkService">
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<mods:title id="F3DC316266D1B95B721C00F3E2C7E738">On the validity of Hebius sauteri maximus (Malnate, 1962) (Squamata, Natricidae), with the redescription of H. maximus comb. nov. and H. sauteri (Boulenger, 1909)</mods:title>
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<mods:namePart id="CEBAE15F2342D0F3C05B8445DDFC025E">Li, Mao-Liang</mods:namePart>
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<mods:affiliation id="EFC780B22C9582934965FC6D26CAFB09">College of Life Science and Technology, Huazhong Agricultural University, Wuhan 430070, Hubei, China & Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
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<mods:namePart id="6B69266246D8D70B710AA2686F125F93">Ren, Jin-Long</mods:namePart>
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<mods:affiliation id="6C3B2F6B48C6065AC615928115C37462">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
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<mods:namePart id="A1E4D8BEE015F419E36BFB0123784FD0">Huang, Jun-Jie</mods:namePart>
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<mods:affiliation id="F12C373B3D747DEF7A24757714455E5B">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
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<mods:namePart id="20BE86C4586E80C8ADC5A709B104C780">Lyu, Zhi-Tong</mods:namePart>
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<mods:affiliation id="390737D395BB6BAB995C15CEA2230AAD">School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
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<mods:namePart id="6A036DFDCD334298C4230BB71272E8D7">Qi, Shuo</mods:namePart>
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<mods:namePart id="B16CAE3E2E783610EF1FD299B9FDBEDD">Jiang, Ke</mods:namePart>
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<mods:affiliation id="7456887EA1C1AF9BAED27D09DE901606">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
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<mods:namePart id="6482945FE123A1F3DA9B509E9E1EDAEF">Wang, Ying-Yong</mods:namePart>
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<mods:affiliation id="3DC7317A0CA5190A54A6760FA7B83A4F">School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
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<mods:namePart id="C96882D16F1F89CB1E230C8C667E37AA">Li, Jia-Tang</mods:namePart>
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<mods:affiliation id="BDD22345A89C73B5B0E2EED495129D42">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China & School of Ecology, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
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<mods:nameIdentifier id="870409FDA7FDB2729B32729A82AAA8D5" type="email">lijt@cib.ac.cn</mods:nameIdentifier>
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<mods:title id="73AD47E1BDCA1853D38F57DDAEBFC02E">Herpetozoa</mods:title>
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<mods:part id="59AFBB92A1BB6168F5B68B2237EB3066">
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<mods:date id="5040EDA7B240C98925FC6E84285A8AE4">2022</mods:date>
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<mods:number id="21F829A8D4787FBEDCD17B287FBB72BE">2022-12-21</mods:number>
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<mods:number id="3D78FEDDA9DDC5DE524E83EB6FCF124B">35</mods:number>
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<mods:start id="AB937372FC0D200B47F23D31991229BD">265</mods:start>
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<mods:end id="1ACD46CCEF220B139DE677302569EF45">282</mods:end>
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<mods:url id="B24AD92CED007EBCC4C6EF9B34B1D8FF">http://dx.doi.org/10.3897/herpetozoa.35.e94920</mods:url>
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<mods:identifier id="231D9CBA8AE9C1DFB69C069738292140" type="DOI">http://dx.doi.org/10.3897/herpetozoa.35.e94920</mods:identifier>
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<treatment id="C3E61DC2F9B05389A33A6DE55B6983C7" ID-GBIF-Taxon="207956806" LSID="urn:lsid:plazi:treatment:C3E61DC2F9B05389A33A6DE55B6983C7" httpUri="http://treatment.plazi.org/id/C3E61DC2F9B05389A33A6DE55B6983C7" lastPageNumber="265" pageId="0" pageNumber="265" scope_class="Squamata">
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<subSubSection id="52BC7918B17ADE62BB74622AEFBACB9A" pageId="0" pageNumber="265" type="nomenclature">
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<paragraph id="860534A5A78C426CCDEE773ED2400FCC" pageId="0" pageNumber="265">
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<taxonomicName id="4EA33EE925C8CD12B583AF901B1A8897" LSID="C3E61DC2-F9B0-5389-A33A-6DE55B6983C7" authority="(Boulenger, 1909)" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">Hebius sauteri (Boulenger, 1909)</taxonomicName>
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</paragraph>
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<subSubSection id="F1A4A00666B1673D4F50673ADB32D987" pageId="0" pageNumber="265" type="description">
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<paragraph id="4D09874420803AF6A3623BA6B1D927AF" pageId="0" pageNumber="265">
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<figureCitation id="C66A67E243EB8A301B4103B55C1DF564" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Hebius maximus comb. nov. and re-defined distribution of H. sauteri in China based on museum collections, previous studies, and this study (Triangle: Hebius maximus comb. nov., round: H. sauteri sauteri, square: H. sauteri bourreti. Type localities are colored red)." figureDoi="10.3897/herpetozoa.35.e94920.figure2" httpUri="https://binary.pensoft.net/fig/785721" pageId="0" pageNumber="265">Figs 2</figureCitation>
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<figureCitation id="5A356BCF9DC432F52F4015EAA4EBD0FE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between Hebius sauteri and Hebius maximus comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang. A-C. Hebius maximus comb. nov. (A. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China; B. CIB 8482, Bijie City, Guizhou, China; C. CIB 8458, Yingjing County, Sichuan, China) D-F. H. sauteri (CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China)." figureDoi="10.3897/herpetozoa.35.e94920.figure3" httpUri="https://binary.pensoft.net/fig/785722" pageId="0" pageNumber="265">, 3</figureCitation>
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<figureCitation id="45B4593294F63EE1A498AB90945364AA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hebius sauteri. A. CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; B. CIB 118517, Xinyong Village, Dehua County, Fujian, China. Photographs by Jin-Long Ren." figureDoi="10.3897/herpetozoa.35.e94920.figure4" httpUri="https://binary.pensoft.net/fig/785723" pageId="0" pageNumber="265">, 4</figureCitation>
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<figureCitation id="47CDD8BEA9B058853E07474AB5C75A3D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Preserved specimen of H. sauteri (CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang." figureDoi="10.3897/herpetozoa.35.e94920.figure5" httpUri="https://binary.pensoft.net/fig/785724" pageId="0" pageNumber="265">, 5</figureCitation>
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<figureCitation id="6711F286AF691C194FD8CE4341D7F4FF" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of the hemipenis of H. sauteri (SYS r 000323, Mt. Jinggang, Jiangxi Province, China), photos from left to right showing sulcate, asulcate, and apical sides of the hemipenis. Photographs by Jun-Jie Huang." figureDoi="10.3897/herpetozoa.35.e94920.figure6" httpUri="https://binary.pensoft.net/fig/785725" pageId="0" pageNumber="265">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="51D257B220EA71F2F47CD085D06530E0" pageId="0" pageNumber="265" type="reference_group">
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<paragraph id="F970148FCD1D548A88DED8E064E9D5AB" pageId="0" pageNumber="265">
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<taxonomicName id="ACA803F88E8497F3E6CD6B8B8A3D5F20" authorityName="Boulenger" authorityYear="1909" family="Colubridae" genus="Tropidonotus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tropidonotus sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">Tropidonotus sauteri</taxonomicName>
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Boulenger, 1909, Ann. Mag. Nat. Hist., London. ser.8, 4, 495. Type locality: Kosempo (= Chiahsien, Kaohsiung County), Taiwan, China.
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</paragraph>
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||
<paragraph id="CA76AC3A2A2F287B0C5D5DF4E66D856B" pageId="0" pageNumber="265">
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<taxonomicName id="571B74CDD36FFA77CFFD156C4BE90A6C" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" baseAuthorityYear="1909" family="Colubridae" genus="Natrix" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Natrix sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">Natrix sauteri</taxonomicName>
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-
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<bibRefCitation id="D2D5627ADAF59FB797D6F17E69A87396" author="Mell, R" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="265" refId="B29" refString="Mell, R, 1931. List of Chinese snakes. Lingnan Science Journal, Canton 8: 199-219." title="List of Chinese snakes. Lingnan Science Journal, Canton 8: 199 - 219." year="1931">Mell 1931</bibRefCitation>
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[1929]: 204;
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||
<bibRefCitation id="13770A82378529256BAE6C13444C72B2" author="Moichiro, M" journalOrPublisher="Daiichi-Shobo, Tokyo" pageId="0" pageNumber="265" refId="B30" refString="Moichiro, M, 1931. A Monograph of the Snakes of Japan. Daiichi-Shobo, Tokyo" title="A Monograph of the Snakes of Japan." year="1931">Moichiro 1931</bibRefCitation>
|
||
: 30;
|
||
<bibRefCitation id="9FCD82C719DEA8D27357C1614EA38F55" author="Bourret, R" journalOrPublisher="Bulletin Generale de l'Instruction Publique" pageId="0" pageNumber="265" pagination="149 - 157" refId="B2" refString="Bourret, R, 1934a. Notes herpetologiques sur l'Indochine francaise. II. Sur quelques serpents des montagnes du Tonkin. Bulletin Generale de l'Instruction Publique 13 (8): 149 - 157" title="Notes herpetologiques sur l'Indochine francaise. II. Sur quelques serpents des montagnes du Tonkin." volume="13" year="1934 a">Bourret 1934a</bibRefCitation>
|
||
: 4;
|
||
<bibRefCitation id="4547B278991DD0DBF3244A6C71868C0E" author="Bourret, R" journalOrPublisher="Bulletin Generale de l'Instruction Publique" pageId="0" pageNumber="265" pagination="289 - 296" refId="B4" refString="Bourret, R, 1935. Notes herpetologiques sur l'Indochine francaise. XI. Sur quelques serpents recoltes en 1934. Bulletin Generale de l'Instruction Publique 14 (9): 289 - 296" title="Notes herpetologiques sur l'Indochine francaise. XI. Sur quelques serpents recoltes en 1934." volume="14" year="1935">Bourret 1935</bibRefCitation>
|
||
: 2;
|
||
<bibRefCitation id="1A99A7DDFCDEEEDAA1C76A18D5A1617A" author="Pope, CH" journalOrPublisher="The American Museum of Natural History, New York" pageId="0" pageNumber="265" refId="B37" refString="Pope, CH, 1935. The reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia (Vol. X). The American Museum of Natural History, New York" title="The reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia (Vol. X)." year="1935">Pope 1935</bibRefCitation>
|
||
: 125 (in part);
|
||
<bibRefCitation id="4C8505D093C1AD46675523C363CE1E11" author="Smith, MA" journalOrPublisher="Taylor and Francis, London" pageId="0" pageNumber="265" refId="B44" refString="Smith, MA, 1943. The fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese sub-region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London" title="The fauna of British India, Ceylon and Burma, including the whole of the Indo-Chinese sub-region. Reptilia and Amphibia. 3 (Serpentes)." year="1943">Smith 1943</bibRefCitation>
|
||
: 287.
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</paragraph>
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||
<paragraph id="C6A7986ACE69209EC5880A0619EFE1ED" pageId="0" pageNumber="265">
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||
<taxonomicName id="C989E9719A4B01BE852C245C4B345AEE" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" baseAuthorityYear="1909" family="Colubridae" genus="Amphiesma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphiesma sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">Amphiesma sauteri</taxonomicName>
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||
-
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<bibRefCitation id="46D5FA891A5D8DAA77FE587FDFA28033" author="Malnate, EV" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="265" pagination="41 - 71" refId="B26" refString="Malnate, EV, 1960. Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae. Proceedings of the Academy of Natural Sciences of Philadelphia 112 (3): 41 - 71" title="Systematic division and evolution of the colubrid snake genus Natrix, with comments on the subfamily Natricinae." volume="112" year="1960">Malnate 1960</bibRefCitation>
|
||
: 51;
|
||
<bibRefCitation id="DC895470E8C684622FD414F87B3197A6" author="Zhao, EM" journalOrPublisher="Molecular Ecology Resources" pageId="0" pageNumber="265" refId="B57" refString="Zhao, EM, Adler, K, 1993. Herpetology of China. Society for the Study of Amphibians and Reptiles, Contributions to Herpetology 10: 522." title="Herpetology of China. Society for the Study of Amphibians and Reptiles, Contributions to Herpetology 10: 522." year="1993">Zhao and Adler 1993</bibRefCitation>
|
||
: 228 (in part);
|
||
<bibRefCitation id="8422F2C13552032FC54B7253FAFE0A94" author="Xie, RG" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="265" pagination=": 82" refId="B54" refString="Xie, RG, Liu, ZH, Qiu, SM, Zheng, WR, Nie, JX, Ma, WH, 1998. Surveys of snakes in Deqing County, Guangdong Province, China. Russian Journal of Herpetology 5 (1): 82" title="Surveys of snakes in Deqing County, Guangdong Province, China." volume="5" year="1998">Xie et al. 1998</bibRefCitation>
|
||
: 82;
|
||
<bibRefCitation id="C95E15D343C41B7D86412416C3A86E68" author="Zhao, EM" journalOrPublisher="Science Press, Beijing" pageId="0" pageNumber="265" refId="B60" refString="Zhao, EM, Huang, MH, Zong, Y, 1998. Fauna Sinica, Reptilia, Squamata. Serpentes (Vol. 3). Science Press, Beijing" title="Fauna Sinica, Reptilia, Squamata. Serpentes (Vol. 3)." year="1998">Zhao et al. 1998</bibRefCitation>
|
||
: 80 (in part);
|
||
<bibRefCitation id="4A7CDE22CAFFBED929353EB88A68D887" author="Orlov, NL" journalOrPublisher="Russian Journal of Herpetology" pageId="0" pageNumber="265" pagination="69 - 80" refId="B34" refString="Orlov, NL, Murphy, RW, Papenfuss, TJ, 2000. List of snakes of Tam-Dao Mountain ridge (Tonkin, Vietnam). Russian Journal of Herpetology 7 (1): 69 - 80" title="List of snakes of Tam-Dao Mountain ridge (Tonkin, Vietnam)." volume="7" year="2000">Orlov et al. 2000</bibRefCitation>
|
||
: 72;
|
||
<bibRefCitation id="9D128ED70CF17FF766DC40A0F222CBB7" DOI="https://doi.org/10.11646/zootaxa.1462.1.2" author="David, P" journalOrPublisher="Zootaxa" pageId="0" pageNumber="265" pagination="41 - 60" refId="B7" refString="David, P, Bain, RH, Nguyen, QT, Orlov, N L, Vogel, G, Vu, T, Ziegler, T, 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41 - 60, DOI: https://doi.org/10.11646/zootaxa.1462.1.2" title="A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae)." url="https://doi.org/10.11646/zootaxa.1462.1.2" volume="1462" year="2007">David et al. 2007</bibRefCitation>
|
||
: 60 (in part);
|
||
<bibRefCitation id="FA016DDC0CBE6FF736507AF22EF92526" author="Nguyen, VS" journalOrPublisher="Edition Chimaira, Frankfurt am Main, Germany" pageId="0" pageNumber="265" refId="B32" refString="Nguyen, VS, Ho, TC, Nguyen, QT, 2009. Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main, Germany" title="Herpetofauna of Vietnam." year="2009">Nguyen et al. 2009</bibRefCitation>
|
||
: 357; Xiang et al. 2012: 230;
|
||
<bibRefCitation id="8260107ED2FB13D29051573883D9B00B" author="Wallach, V" journalOrPublisher="CRC Press Boca Raton, Florida, USA" pageId="0" pageNumber="265" refId="B49" refString="Wallach, V, Williams, KL, Boundy, J, 2014. Snakes of the world: a catalogue of living and extinct species. CRC Press Boca Raton, Florida, USA" title="Snakes of the world: a catalogue of living and extinct species." year="2014">Wallach et al. 2014</bibRefCitation>
|
||
: 33 (in part).
|
||
</paragraph>
|
||
<paragraph id="9E5F01E0111A55E4013AB95CA2B9331D" pageId="0" pageNumber="265">
|
||
<taxonomicName id="4E6756C0BE5507FDDC99878B957E3F3A" authority="sauteri" family="Colubridae" genus="Amphiesma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphiesma sauteri subsp. sauteri" order="Squamata" pageId="0" pageNumber="265" rank="subSpecies" species="sauteri" subSpecies="sauteri">Amphiesma sauteri sauteri</taxonomicName>
|
||
-
|
||
<bibRefCitation id="EBCE5AC1CB3814DD841F2DA0C6C6A892" author="Malnate, EV" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="265" pagination="251 - 299" refId="B27" refString="Malnate, EV, 1962. The relationships of five species of the Asiatic natricine snake genus Amphiesma. Proceedings of the Academy of Natural Sciences of Philadelphia 114 (8): 251 - 299" title="The relationships of five species of the Asiatic natricine snake genus Amphiesma." volume="114" year="1962">Malnate 1962</bibRefCitation>
|
||
: 272.
|
||
</paragraph>
|
||
<paragraph id="800F263C07F229991863CB5B5FF4D0FF" pageId="0" pageNumber="265">
|
||
<taxonomicName id="42AD38B44023FA21E46A085B62D7AF8C" family="Colubridae" genus="Amphiesma" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Amphiesma sauteri subsp. bourreti" order="Squamata" pageId="0" pageNumber="265" rank="subSpecies" species="sauteri" subSpecies="bourreti">Amphiesma sauteri bourreti</taxonomicName>
|
||
-
|
||
<bibRefCitation id="130007CFEBB8FA821974ABD2CF80F296" author="Malnate, EV" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="265" pagination="251 - 299" refId="B27" refString="Malnate, EV, 1962. The relationships of five species of the Asiatic natricine snake genus Amphiesma. Proceedings of the Academy of Natural Sciences of Philadelphia 114 (8): 251 - 299" title="The relationships of five species of the Asiatic natricine snake genus Amphiesma." volume="114" year="1962">Malnate 1962</bibRefCitation>
|
||
: 273.
|
||
</paragraph>
|
||
<paragraph id="A75AE6A68963A7CAB66999365CEF4537" pageId="0" pageNumber="265">
|
||
<taxonomicName id="4037154462E941BD908337BF73B942EF" authorityName="Van Denburgh" authorityYear="1909" family="Colubridae" genus="Natrix" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Natrix copei" order="Squamata" pageId="0" pageNumber="265" rank="species" species="copei">Natrix copei</taxonomicName>
|
||
<bibRefCitation id="A73E2B2E1B0C1097C495BD938212FD04" author="Van Denburgh, J" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="0" pageNumber="265" pagination="49 - 56" refId="B48" refString="Van Denburgh, J, 1909. New and previously unrecorded species of reptiles and amphibians from the island of Formosa. Proceedings of the California Academy of Sciences 4 (3): 49 - 56" title="New and previously unrecorded species of reptiles and amphibians from the island of Formosa." volume="4" year="1909">Van Denburgh 1909</bibRefCitation>
|
||
, Proc. California Acad. Sci. 3(3): 52-53. Type locality: "Kanshirei, Formosa" (= Guanziling, Tainan County, Taiwan, China)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="6BAF9523C665362F48B85B11E5A9C374" pageId="0" pageNumber="265" type="types">
|
||
<paragraph id="3B4B62907899CE7A4A0918276EC773C2" pageId="0" pageNumber="265">Types.</paragraph>
|
||
<paragraph id="0A8B3366E3FDFD5E77BE5587791D642D" pageId="0" pageNumber="265">
|
||
<emphasis id="2B8F24453847C0BC725C673DF2222F1C" bold="true" italics="true" pageId="0" pageNumber="265">Syntypes</emphasis>
|
||
. NHMUK (formerly BMNH) 1946.1.13.38-39, a male and a female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E342C9E2C724BBD5029385C58097769F" pageId="0" pageNumber="265" type="referred specimens">
|
||
<paragraph id="2251C8B26D5A94AC6B292E1728059EEB" pageId="0" pageNumber="265">Referred specimens.</paragraph>
|
||
<paragraph id="61D9BB5E9B62A4BD716A24B3C0D83A06" pageId="0" pageNumber="265">
|
||
19 specimens of
|
||
<taxonomicName id="14ACE034BA3C2C8598A5B22D9CF5E3CA" lsidName="H. s. subsp. sauteri" pageId="0" pageNumber="265" rank="subSpecies" species="s." subSpecies="sauteri">
|
||
<emphasis id="C73D3B2D8B89353DFEC4AEE136720267" italics="true" pageId="0" pageNumber="265">H. s. sauteri</emphasis>
|
||
</taxonomicName>
|
||
were examined in this study: CHINA (
|
||
<emphasis id="FD798FB54CDCFADF334413D5CB190713" italics="true" pageId="0" pageNumber="265">n</emphasis>
|
||
= 19) -
|
||
<emphasis id="7C9678149A01D24E951AAFE5609428AE" bold="true" pageId="0" pageNumber="265">Taiwan Prov.</emphasis>
|
||
CAS 18984, Pingtung County; CAS 18988 (paratype of
|
||
<taxonomicName id="A02D65FF425AB1CBFCCB68EFFC336E72" authorityName="Van Denburgh" authorityYear="1909" family="Colubridae" genus="Natrix" kingdom="Animalia" lsidName="Natrix copei" order="Squamata" pageId="0" pageNumber="265" rank="species" species="copei">
|
||
<emphasis id="51FE7815E2B5541FA2C12598553A419A" italics="true" pageId="0" pageNumber="265">Natrix copei</emphasis>
|
||
</taxonomicName>
|
||
Van Denburgh, 1909), Tainan City, Guanziling. -
|
||
<emphasis id="B94FE3D332DC093D5BDDFD80426718D6" bold="true" pageId="0" pageNumber="265">Fujian Prov.</emphasis>
|
||
CIB 8476-8479, CIB 78046, Dehua County, Daiyun; CIB 118517, Dehua County, Xinyong Village; CIB 8480, Wuyishan City, Xingcun Township. -
|
||
<emphasis id="9D51A4C65AC124763C9112E0413572EC" bold="true" pageId="0" pageNumber="265">Anhui Prov.</emphasis>
|
||
CIB 8483, Huangshan City. -
|
||
<emphasis id="0EF7F0E863DEB3E4733A446993E8EA15" bold="true" pageId="0" pageNumber="265">Jiangxi Prov.</emphasis>
|
||
SYS r000323, Jinggangshan City, Mt. Jinggang; SYS r001258, SYS r001266, Longnan County, Jiulianshan. -
|
||
<emphasis id="0EB12600E9A9C725545F5CBC1EFB6090" bold="true" pageId="0" pageNumber="265">Hainan Prov.</emphasis>
|
||
YBU 17001, Ledong Li Autonomous County, Jianfengling; CIB 8481. -
|
||
<emphasis id="131431D7927467549EBAC668B62C15ED" bold="true" pageId="0" pageNumber="265">Guangdong Prov.</emphasis>
|
||
SYS r001150, Xinyi City, Sihe Township; CIB 118516, Dongguan City, Yinpingshan. -
|
||
<emphasis id="6F788D512FCD8949D34D117F0315A699" bold="true" pageId="0" pageNumber="265">Guizhou Prov.</emphasis>
|
||
SYS r000275, Libo County, Maolan National Natural Reserve. -
|
||
<emphasis id="8AEE11E3F50980B294D60B53F1C42124" bold="true" pageId="0" pageNumber="265">Hunan Prov.</emphasis>
|
||
SYS r001766, Guidong County, Bamianshan.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A8643B71BAE5EAECD1FED869EDBD833E" pageId="0" pageNumber="265" type="diagnosis">
|
||
<paragraph id="8FF3E6DA883A508BC0FBA3144F2D7C94" pageId="0" pageNumber="265">Diagnosis.</paragraph>
|
||
<paragraph id="A0ABA3A5C147CD0A1E869B4ED53B2A05" pageId="0" pageNumber="265">
|
||
<taxonomicName id="AE573977115A4C52EE9EDA05E7B73E18" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="B1623DA27000E415B55C36C1EFBB97DF" italics="true" pageId="0" pageNumber="265">Hebius sauteri</emphasis>
|
||
</taxonomicName>
|
||
can be distinguished from its congeners by the following features: (1) body size small to medium, maximum ToL 455 mm; (2) dorsal scales rows 17-17-17, all weekly keeled or smooth on outmost one or two rows; (3) ventral scales 116-133, subcaudals 60-86, paired; (4) maxillary teeth 23-27, last two or three distinctly enlarged teeth, without diastema; (5) preocular single, occasionally two; (6) supralabials 6-8, 2nd or 2nd-3rd border the loreal; (7) infralabials 6-9, mostly edged with black on their posterior border; (8) single white to orange stripe extends from
|
||
<emphasis id="8D67B24B78AC43C82F88060761A24A79" italics="true" pageId="0" pageNumber="265">angulus oris</emphasis>
|
||
to neck; (9) dorsal body brown to reddish brown, with or without single series of white spots or short band on lateral.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B00372E5AA67118AD01883953E87B266" pageId="0" pageNumber="265" type="description">
|
||
<paragraph id="564B112D044F34AB75D84624A6F06EBF" pageId="0" pageNumber="265">Description.</paragraph>
|
||
<paragraph id="849B2490C215F2E743743C7166E70EB5" pageId="0" pageNumber="265">Head moderately distinct from neck; body size small to medium, ToL 145-455 mm (282-370 mm in males, 145-455 mm in females), TL/ToL 0.24-0.33 (0.27-0.31 in males, 0.24-0.33 in females). Eye large, pupil round.</paragraph>
|
||
<paragraph id="7374046F441D1F12D3DC20A9C1A52DDE" pageId="0" pageNumber="265">
|
||
<emphasis id="79FDE666150C972C76F0D2BA0409B84C" bold="true" pageId="0" pageNumber="265">
|
||
<emphasis id="1FA713DC9733D198ED17F3581D6CFA56" italics="true" pageId="0" pageNumber="265">Dentition</emphasis>
|
||
.
|
||
</emphasis>
|
||
Maxillary teeth 23-27 without diastema, gradually enlarged posteriorly, last two or three distinctly enlarged.
|
||
</paragraph>
|
||
<paragraph id="A34908485A2F7BBA4A9CE75FAAD5118F" pageId="0" pageNumber="265">
|
||
<emphasis id="053138080A6164991F0FFD78A5C7D0A2" bold="true" pageId="0" pageNumber="265">
|
||
<emphasis id="0907B5B7C506EE4B98000A141515EBFE" italics="true" pageId="0" pageNumber="265">Body scalation</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body scale in 17-17-17 rows, weekly keeled except outer 1-4 rows smooth anteriorly, outer 0-2 rows smooth at midbody and outer 0-1 row smooth posteriorly; ventrals 116-133 (125-130 in males, 116-133 in females); subcaudals 60-86 (60-86 in females, 65-84 in males), paired.
|
||
</paragraph>
|
||
<paragraph id="B376EA4F5D21094E19D2702FB38A5187" pageId="0" pageNumber="265">
|
||
<emphasis id="7EF81A91C67A4D0CB47C37787D50E1B8" bold="true" pageId="0" pageNumber="265">
|
||
<emphasis id="E23F82826D5D192FED7482B77A575CF4" italics="true" pageId="0" pageNumber="265">Head scalation</emphasis>
|
||
.
|
||
</emphasis>
|
||
Rostral broad, visible from above; internasals paired, nearly triangular, curved outwards, not in contact with loreal; prefrontals paired, nearly in a diamond shape, wider than long, border on loreal laterally, preocular and supraocular posteriorly; frontal hexagonal, longer than wide, embedded into parietal posteriorly; parietal paired, much longer than wide; nasals wider than high, divided below nostril, borders 1st-2nd supralabials; loreal single, rarely divided (right side of CIB 8481), borders 2nd or 2nd-3rd supralabials, not entering orbit; preocular 1-2; supraocular single, narrowed anteriorly, much wider than high, not in contact with parietal; postoculars 2-3, rarely 4 (left side of CIB 8483); supralabials 6-8, 5th, 6th or 7th highest, 3rd -4th entering orbit, rarely 2nd-3rd (right side of CIB 8480), 3rd only (both sides of CIB 8479) or 4th-5th (left side of CIB 8477); infralabials 6-9, first pair in contact behind the mental, IfL 1-3, 1-4 or 1-5 border the anterior chin shields; temporals variable, in two or three rows, 1-2 + 1-3 + 0/2-3; chin shields paired, posterior pair longer (Figs
|
||
<figureCitation id="CA1665ECBAEACB84932173A5E9383BEE" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between Hebius sauteri and Hebius maximus comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang. A-C. Hebius maximus comb. nov. (A. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China; B. CIB 8482, Bijie City, Guizhou, China; C. CIB 8458, Yingjing County, Sichuan, China) D-F. H. sauteri (CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China)." figureDoi="10.3897/herpetozoa.35.e94920.figure3" httpUri="https://binary.pensoft.net/fig/785722" pageId="0" pageNumber="265">3</figureCitation>
|
||
,
|
||
<figureCitation id="57B29337670EFE96B1834B822340565A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Preserved specimen of H. sauteri (CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang." figureDoi="10.3897/herpetozoa.35.e94920.figure5" httpUri="https://binary.pensoft.net/fig/785724" pageId="0" pageNumber="265">5</figureCitation>
|
||
, Table
|
||
<tableCitation id="8BCB7C14828F75384AFC62212FB2AF66" captionStart="Table 4" captionStartId="T4" captionText="Table 4. Morphological characters of H. maximus and H. sauteri. For specimens with asymmetrical cephalic scales, the scale count was given as " left / right ", " - " indicates missing data." httpUri="http://table.plazi.org/id/CEF25E8772CA7CD1980C3D0E96F6D81F" pageId="0" pageNumber="265" tableUuid="CEF25E8772CA7CD1980C3D0E96F6D81F">4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="860335B854ABCFA82B665F3B016D7B92" pageId="0" pageNumber="265">
|
||
<emphasis id="A75C68B2413C671306D5FC32C76B77FF" bold="true" pageId="0" pageNumber="265">
|
||
<emphasis id="35DB769AED6FB48418BE76AC8DE5F73E" italics="true" pageId="0" pageNumber="265">Hemipenis</emphasis>
|
||
.
|
||
</emphasis>
|
||
The description of the hemipenis is based on an adult specimen (SYS r000323) from Mt. Jinggang, Jiangxi, southeastern China. Left side of hemipenis fully everted, right side remained
|
||
<emphasis id="1687E213C267789DA363C697AB039C37" italics="true" pageId="0" pageNumber="265">in situ</emphasis>
|
||
for description of retracted condition.
|
||
</paragraph>
|
||
<paragraph id="D26E53F7BE7071AD02AE9C8C176097B8" pageId="0" pageNumber="265">
|
||
The everted hemipenis is thin and short, Y-shaped, shallowly bilobed. Both sulcate and asulcate surfaces densely ornamented with spinules, a large basal hook present at proximal part.
|
||
<taxonomicName id="2C46FE4D0B8FAFC6BF1FA42D5331537A" class="Magnoliopsida" family="Crassulaceae" genus="Sulcus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sulcus spermaticus" order="Saxifragales" pageId="0" pageNumber="265" phylum="Tracheophyta" rank="species" species="spermaticus">
|
||
<emphasis id="62556E923E8ECDE380F127A5DFD061B9" italics="true" pageId="0" pageNumber="265">Sulcus spermaticus</emphasis>
|
||
</taxonomicName>
|
||
single, extending to base of inner right lobe where it takes a centripetal position.
|
||
<taxonomicName id="BACD02319C9623A0CC48DE6FCA6DBD7B" authorityName="R.T.Clausen" authorityYear="1979" class="Magnoliopsida" family="Crassulaceae" genus="Sulcus" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Sulcus" order="Saxifragales" pageId="0" pageNumber="265" phylum="Tracheophyta" rank="genus">Sulcus</taxonomicName>
|
||
lip highly developed and raised, walls covered with spinules (Fig.
|
||
<figureCitation id="C43E691CD7AEFB26DFD0894425435607" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Photographs of the hemipenis of H. sauteri (SYS r 000323, Mt. Jinggang, Jiangxi Province, China), photos from left to right showing sulcate, asulcate, and apical sides of the hemipenis. Photographs by Jun-Jie Huang." figureDoi="10.3897/herpetozoa.35.e94920.figure6" httpUri="https://binary.pensoft.net/fig/785725" pageId="0" pageNumber="265">6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E980640B1ACEBCFED31D2112626980B4" pageId="0" pageNumber="265">
|
||
<emphasis id="E2EDEC92444755C95B5393EAEAE3E4A8" italics="true" pageId="0" pageNumber="265">In situ</emphasis>
|
||
hemipenis extends to 7thSC with its crotch extending to 6thSC; crotch of
|
||
<emphasis id="C60F9D30C1A3D00211541CBD5155F0F5" italics="true" pageId="0" pageNumber="265">m. retractor penis magnus</emphasis>
|
||
extending to 8thSC, origin of
|
||
<emphasis id="F1CBA57F494ECC4E60B5CC0883A2689C" italics="true" pageId="0" pageNumber="265">m. retractor penis magnus</emphasis>
|
||
at level of 20thSC.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CFE93ECA381B691BC1C5821281C5A201" pageId="0" pageNumber="265" type="coloration in life">
|
||
<paragraph id="C044E25B2EF0BFC54FDE45BC392958E2" pageId="0" pageNumber="265">Coloration in life.</paragraph>
|
||
<paragraph id="FFCBABC27A9B204864B3843D56407AFE" pageId="0" pageNumber="265">
|
||
Based on two adult female specimens from Guangdong (CIB 118516) and Fujian (CIB 118517), China. Dorsal surface of the head reddish-brown with irregular grayish-green spots, a pair of elliptic orangish spots symmetrically present on inner sides of parietals, just separated by parietal sutures; ventral side of head whitish; 1st-5th supralabials white with black edges posteriorly, 6th or 6-7th brown on their upper part; infralabials white with black edge between each scale; an orange stripe extends from
|
||
<emphasis id="D47F9793394484DD8C9030779B466BE7" italics="true" pageId="0" pageNumber="265">angulus oris</emphasis>
|
||
to dorsal side of neck, 1-2 scales wide; dorsal surface of body and tail dark brown to reddish-brown, top with or without an ill-defined dark olive stripe scattered with black spots extends from neck to the end, a reddish-brown lateral streak on 4th-6th body scale rows extends from neck to tail with series of orange spots separated by 2-3 scales above; ventral and subcaudal sides cream white, with black spots on both ends of the ventral scale forming a chain-like pattern on the ventral view; sclera brown (Fig.
|
||
<figureCitation id="670F1D6781FBCF06FA9E2ABD707D98A7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Hebius sauteri. A. CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; B. CIB 118517, Xinyong Village, Dehua County, Fujian, China. Photographs by Jin-Long Ren." figureDoi="10.3897/herpetozoa.35.e94920.figure4" httpUri="https://binary.pensoft.net/fig/785723" pageId="0" pageNumber="265">4</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="9C1BDD6CF7114622292433369EED01D3" doi="10.3897/herpetozoa.35.e94920.figure2" httpUri="https://binary.pensoft.net/fig/785721" pageId="0" pageNumber="265" start="Figure 2" startId="F2">
|
||
<paragraph id="79D0FA140C513FF09FAED437EF48D831" pageId="0" pageNumber="265">
|
||
<emphasis id="906114FA56642AE178F5982D76F4570C" bold="true" pageId="0" pageNumber="265">Figure 2.</emphasis>
|
||
Distribution of
|
||
<taxonomicName id="D30A6B68FDB93DB1D62E33BD9C107240" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Malnate" baseAuthorityYear="1962" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius maximus" order="Squamata" pageId="0" pageNumber="265" rank="species" species="maximus">
|
||
<emphasis id="5A7E3B89231661D49ADA45218059DA8C" italics="true" pageId="0" pageNumber="265">Hebius maximus</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. and re-defined distribution of
|
||
<taxonomicName id="D3EE176CE61D2089F31AF7201AD91E86" lsidName="H. sauteri" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="2E0ADA565AC045574E9B6FCD61402CC4" italics="true" pageId="0" pageNumber="265">H. sauteri</emphasis>
|
||
</taxonomicName>
|
||
in China based on museum collections, previous studies, and this study (Triangle:
|
||
<taxonomicName id="3E0759DCCF32B829E624B307F038537D" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Malnate" baseAuthorityYear="1962" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius maximus" order="Squamata" pageId="0" pageNumber="265" rank="species" species="maximus">
|
||
<emphasis id="839B737300CF4DF5E958DEF3C8AF8E7B" italics="true" pageId="0" pageNumber="265">Hebius maximus</emphasis>
|
||
</taxonomicName>
|
||
comb. nov., round:
|
||
<taxonomicName id="4E77A360E38A157E27A4BA4C8BBA1ED4" authority="sauteri" lsidName="H. sauteri subsp. sauteri" pageId="0" pageNumber="265" rank="subSpecies" species="sauteri" subSpecies="sauteri">
|
||
<emphasis id="0DF7950AFE7551747B49B68A5E3684F2" italics="true" pageId="0" pageNumber="265">H. sauteri sauteri</emphasis>
|
||
</taxonomicName>
|
||
, square:
|
||
<taxonomicName id="D313CDDF1B92452C952ECF11E6DF6EA8" lsidName="H. sauteri subsp. bourreti" pageId="0" pageNumber="265" rank="subSpecies" species="sauteri" subSpecies="bourreti">
|
||
<emphasis id="4D035CCB7DAB54CAEE3EB362EE9D3E9F" italics="true" pageId="0" pageNumber="265">H. sauteri bourreti</emphasis>
|
||
</taxonomicName>
|
||
. Type localities are colored red).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A88854C392E7D748DFEF80EE454A505F" doi="10.3897/herpetozoa.35.e94920.figure3" httpUri="https://binary.pensoft.net/fig/785722" pageId="0" pageNumber="265" start="Figure 3" startId="F3">
|
||
<paragraph id="58C1A4BB362FDB9466B27A6566E62AB5" pageId="0" pageNumber="265">
|
||
<emphasis id="B90F7608DB2D290F9B85C4E6E003CAFC" bold="true" pageId="0" pageNumber="265">Figure 3.</emphasis>
|
||
Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between
|
||
<taxonomicName id="284016E39C8587B24EE4F9472B38523F" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="5DE6EE158AC6764B29CD260A93618404" italics="true" pageId="0" pageNumber="265">Hebius sauteri</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="DA4CE7B3FD7EB4807EF06D569C2AD56B" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Malnate" baseAuthorityYear="1962" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius maximus" order="Squamata" pageId="0" pageNumber="265" rank="species" species="maximus">
|
||
<emphasis id="72496BCF5081123B3C87ACE512C58886" italics="true" pageId="0" pageNumber="265">Hebius maximus</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang.
|
||
<emphasis id="31F56AAAF2B82EBBC63659E9E509D8AC" bold="true" pageId="0" pageNumber="265">A-C.</emphasis>
|
||
<taxonomicName id="8B73260699AE7DDFB4141B4B68F118BD" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Malnate" baseAuthorityYear="1962" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius maximus" order="Squamata" pageId="0" pageNumber="265" rank="species" species="maximus">
|
||
<emphasis id="1C3AEDF5C5748AC18CE80A9312226A02" italics="true" pageId="0" pageNumber="265">Hebius maximus</emphasis>
|
||
</taxonomicName>
|
||
comb. nov. (
|
||
<emphasis id="F738EA623F83004FA5B294007292FDE1" bold="true" pageId="0" pageNumber="265">A</emphasis>
|
||
. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China;
|
||
<emphasis id="2DDCFE2520B90589EE41275701F4A6C4" bold="true" pageId="0" pageNumber="265">B.</emphasis>
|
||
CIB 8482, Bijie City, Guizhou, China;
|
||
<emphasis id="AFAEED43AA315A467C65E99ADD6D62C1" bold="true" pageId="0" pageNumber="265">C.</emphasis>
|
||
CIB 8458, Yingjing County, Sichuan, China)
|
||
<emphasis id="7F166AC83F85F1BF483297E1FBC1EC6D" bold="true" pageId="0" pageNumber="265">D-F.</emphasis>
|
||
<taxonomicName id="4CF93F098BDC975142A73DF4C43CDA01" lsidName="H. sauteri" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="84A1C7236BE8F86DFBECD140D1EF4A69" italics="true" pageId="0" pageNumber="265">H. sauteri</emphasis>
|
||
</taxonomicName>
|
||
(CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CADD961FC55AA64867E80C8659DBF68F" doi="10.3897/herpetozoa.35.e94920.figure4" httpUri="https://binary.pensoft.net/fig/785723" pageId="0" pageNumber="265" start="Figure 4" startId="F4">
|
||
<paragraph id="8F40C0560B5BF51CF3DE7D3F9AAF08F5" pageId="0" pageNumber="265">
|
||
<emphasis id="7CE009E334E3FD5974A6B5CB8A8595F1" bold="true" pageId="0" pageNumber="265">Figure 4.</emphasis>
|
||
<taxonomicName id="A19DF58D5F65D9688C5C2DD0B940484B" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="C10E5A3CDC4486D6310CFF00489A403F" italics="true" pageId="0" pageNumber="265">Hebius sauteri</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="45D150A6434CCA7C76DEE26A72B0B1D2" bold="true" pageId="0" pageNumber="265">A.</emphasis>
|
||
CIB 118516, Yinpingshan, Dongguan City, Guangdong, China;
|
||
<emphasis id="0146E4541FBE3CC084B7B44E4D01630C" bold="true" pageId="0" pageNumber="265">B.</emphasis>
|
||
CIB 118517, Xinyong Village, Dehua County, Fujian, China. Photographs by Jin-Long Ren.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="F6568C406F683E7C82C8F15FCA6FAA4D" doi="10.3897/herpetozoa.35.e94920.figure5" httpUri="https://binary.pensoft.net/fig/785724" pageId="0" pageNumber="265" start="Figure 5" startId="F5">
|
||
<paragraph id="6A7C318BDBFEAC122F458F92715BC157" pageId="0" pageNumber="265">
|
||
<emphasis id="4AC0509FA85B3B98E3E1A0CC1CCA3C73" bold="true" pageId="0" pageNumber="265">Figure 5.</emphasis>
|
||
Preserved specimen of
|
||
<taxonomicName id="6C35E8F7060860C5F6E9F8F4F903B635" lsidName="H. sauteri" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="171B4BAE24A6CF44BFA077425E401727" italics="true" pageId="0" pageNumber="265">H. sauteri</emphasis>
|
||
</taxonomicName>
|
||
(CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E8C5C9482469788B63C974DEFB078A0D" doi="10.3897/herpetozoa.35.e94920.figure6" httpUri="https://binary.pensoft.net/fig/785725" pageId="0" pageNumber="265" start="Figure 6" startId="F6">
|
||
<paragraph id="C495F89A463D4F186318450119323E4D" pageId="0" pageNumber="265">
|
||
<emphasis id="5AE469FC71C8D6A4941F607AC88490A7" bold="true" pageId="0" pageNumber="265">Figure 6.</emphasis>
|
||
Photographs of the hemipenis of
|
||
<taxonomicName id="2612FB94F0E08EB435591F4F0DECA75F" lsidName="H. sauteri" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="9770299ACAC355F4CAE5DD7870ED86F6" italics="true" pageId="0" pageNumber="265">H. sauteri</emphasis>
|
||
</taxonomicName>
|
||
(SYS r000323, Mt. Jinggang, Jiangxi Province, China), photos from left to right showing sulcate, asulcate, and apical sides of the hemipenis. Photographs by Jun-Jie Huang.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="3D26223993423B8C7BF22B1C07A851F2" pageId="0" pageNumber="265" type="coloration in preservative">
|
||
<paragraph id="421EC718EE810C3DB6BC8003F55D90E4" pageId="0" pageNumber="265">Coloration in preservative.</paragraph>
|
||
<paragraph id="6916E673B664055D49E62EAFF849FBEF" pageId="0" pageNumber="265">
|
||
Largely the same as in life except dorsal side of head brown to dark brown, scattered with small black spots; ventral side of head cream white; a pale yellow stripe extends from
|
||
<emphasis id="93591A82B2F67796A02FF72B8C5B0766" italics="true" pageId="0" pageNumber="265">angulus oris</emphasis>
|
||
to dorsal side of neck, 1-2 scales wide; dorsal of body and tail brown to dark brown, a lighter lateral streak on 4th-6th body scale rows presents or not, covered with series of cream white spots separated by 2-3 scales; ventral and subcaudal sides cream to pale white; sclera black (Figs
|
||
<figureCitation id="DC01DD24C547C671BC9C002EEC429993" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between Hebius sauteri and Hebius maximus comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang. A-C. Hebius maximus comb. nov. (A. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China; B. CIB 8482, Bijie City, Guizhou, China; C. CIB 8458, Yingjing County, Sichuan, China) D-F. H. sauteri (CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China)." figureDoi="10.3897/herpetozoa.35.e94920.figure3" httpUri="https://binary.pensoft.net/fig/785722" pageId="0" pageNumber="265">3</figureCitation>
|
||
,
|
||
<figureCitation id="25BEB723E50299CC78A796A87B4A4478" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Preserved specimen of H. sauteri (CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang." figureDoi="10.3897/herpetozoa.35.e94920.figure5" httpUri="https://binary.pensoft.net/fig/785724" pageId="0" pageNumber="265">5</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="96CC0E9479B35ADD367A197E148AA452" pageId="0" pageNumber="265" type="etymology">
|
||
<paragraph id="6F99C70DF84E6247725E6E65D7A8CF94" pageId="0" pageNumber="265">Etymology.</paragraph>
|
||
<paragraph id="EFF5BCC066E49D1A19BD56EB189EE953" pageId="0" pageNumber="265">
|
||
<taxonomicName id="5759AAA804F8C67C2050296CB09FB0C5" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="EDBA81B6098F96CA3175FF1D8D475341" italics="true" pageId="0" pageNumber="265">Hebius sauteri</emphasis>
|
||
</taxonomicName>
|
||
was named after the collector of the type specimens, Hans Sauter (1871-1943), a German naturalist (
|
||
<bibRefCitation id="3CE4105888F53A3E0D61713BAEAFF35E" DOI="https://doi.org/10.1080/00222930908692704" author="Boulenger, GA" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="265" pagination="492 - 495" refId="B1" refString="Boulenger, GA, 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. The Annals and Magazine of Natural History 8 (4): 492 - 495, DOI: https://doi.org/10.1080/00222930908692704" title="Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa." url="https://doi.org/10.1080/00222930908692704" volume="8" year="1909">Boulenger 1909</bibRefCitation>
|
||
). We suggest that the Chinese common name of this species remains unchanged, as "Zōng
|
||
<normalizedToken id="18ACD73AD2B5BD7022FC70FCFE6C0B47" originalValue="Hēi">Hēi</normalizedToken>
|
||
<normalizedToken id="0C27DFD81BB704C52A2B13E3D9CD4350" originalValue="Fù">Fu</normalizedToken>
|
||
<normalizedToken id="7421A72120373AA5F72FA110D8601910" originalValue="Liàn">Lian</normalizedToken>
|
||
<normalizedToken id="05D28CFA82902FE374F0351E7819987A" originalValue="Shé">She</normalizedToken>
|
||
(
|
||
<normalizedToken id="F997FE27ECF3F652B1F02DCD08B3EFDA" originalValue="棕黑腹链蛇)”">棕黑腹链蛇)"</normalizedToken>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4E46C091BDDE157307D4081D317CE8CF" pageId="0" pageNumber="265" type="natural history">
|
||
<paragraph id="8AF803C5C584923BC24D7D7636C973F3" pageId="0" pageNumber="265">Natural history.</paragraph>
|
||
<paragraph id="833E7568753498DE041CA951467A72E5" pageId="0" pageNumber="265">
|
||
This species inhabits flatlands and mountain regions up to 1,500 m and prefers terrestrial microhabitat such as grassland, woodland, and bushes near water bodies; diurnal, occasionally found at night, preys include earthworms, slugs, snails and tadpoles; oviparous (this study;
|
||
<bibRefCitation id="9F68429525016E1CBC71E972D1A3FE5A" author="Pope, CH" journalOrPublisher="The American Museum of Natural History, New York" pageId="0" pageNumber="265" refId="B37" refString="Pope, CH, 1935. The reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia (Vol. X). The American Museum of Natural History, New York" title="The reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia (Vol. X)." year="1935">Pope 1935</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="A94C184121AECC65354157E0E8DD3725" author="Zhao, EM" journalOrPublisher="Anhui Science and Technology Publishing House, Hefei" pageId="0" pageNumber="265" refId="B62" refString="Zhao, EM, 2006. Snakes of China. Anhui Science and Technology Publishing House, Hefei" title="Snakes of China." year="2006">Zhao 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="2C7C7BEDEB91347D68D11498CC5BB917" author="Xiang, GS" journalOrPublisher="Owl Press, Taipei" pageId="0" pageNumber="265" refId="B53" refString="Xiang, GS, Li, PX, Yang, YR, 2009. Colored illustrations of amphibians and reptiles of Taiwan. Owl Press, Taipei" title="Colored illustrations of amphibians and reptiles of Taiwan." year="2009">Xiang et al. 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E7EA12C6435DAE67575724904A436711" pageId="0" pageNumber="265" type="distribution">
|
||
<paragraph id="5B0F198386D8BB9EBD5ECA6F34FF109F" pageId="0" pageNumber="265">Distribution.</paragraph>
|
||
<paragraph id="EFDE957A48E58BF736C7A790F8848562" pageId="0" pageNumber="265">
|
||
<taxonomicName id="CB6B3102EB3FD9B4EF5A286C00D2AF73" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="620A95B561759EB7ADA766457AC50725" italics="true" pageId="0" pageNumber="265">Hebius sauteri</emphasis>
|
||
</taxonomicName>
|
||
is currently known to be reported in Vietnam (Ngan Son, Lang Son, Tam Dao and Ba Vi), and China, including Taiwan (Kaohsiung County [type locality]), Fujian (Dehua County), Anhui (Huangshan County), Jiangxi (Jinggangshan City, Longnan County), Hunan (Yizhang County), Guangdong (Lianping County, Xinyi County, Deqing County, Ruyuan County, Dongguan City), Guangxi (Jinxiu County, Quanzhou County, Rongxian County, Beiliu County,
|
||
<normalizedToken id="E3171B384B25C5C57F55C24B38EF1FEF" originalValue="Long’an">Long'an</normalizedToken>
|
||
County), Guizhou (Libo County, Leishan County), Yunnan (Jinping County), Hainan (Qiongzhong County, Wuzhishan County, Baisha County, Lingshui County) (
|
||
<bibRefCitation id="23D936C570B10C4EAE7EE9437E2374FB" author="Deuve, J" journalOrPublisher="M., Paris" pageId="0" pageNumber="265" refId="B10" refString="Deuve, J, 1970. Serpents du Laos. Memoirs O.R.S.T.O. M., Paris" title="Serpents du Laos. Memoirs O. R. S. T. O." year="1970">Deuve 1970</bibRefCitation>
|
||
; Wu 1985;
|
||
<bibRefCitation id="5463AC94BDD92028DA465187A5E90114" author="Zhao, EM" journalOrPublisher="Anhui Science and Technology Publishing House, Hefei" pageId="0" pageNumber="265" refId="B62" refString="Zhao, EM, 2006. Snakes of China. Anhui Science and Technology Publishing House, Hefei" title="Snakes of China." year="2006">Zhao 2006</bibRefCitation>
|
||
; Yang et al. 2008;
|
||
<bibRefCitation id="71D1458EA646E5A025ABE9658EE72EAE" author="Nguyen, VS" journalOrPublisher="Edition Chimaira, Frankfurt am Main, Germany" pageId="0" pageNumber="265" refId="B32" refString="Nguyen, VS, Ho, TC, Nguyen, QT, 2009. Herpetofauna of Vietnam. Edition Chimaira, Frankfurt am Main, Germany" title="Herpetofauna of Vietnam." year="2009">Nguyen et al. 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="565EF43FB42BE5EA4CA7872B8E0DAF8A" author="Zhang, YX" journalOrPublisher="Guangxi Normal University Press, Guilin" pageId="0" pageNumber="265" refId="B59" refString="Zhang, YX, 2009. Herpetology in Guangxi. Guangxi Normal University Press, Guilin" title="Herpetology in Guangxi." year="2009">Zhang 2009</bibRefCitation>
|
||
; Li 2011; Shen 2014; this study) (Fig.
|
||
<figureCitation id="51574675F7DDB91425DE180A881F0352" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of Hebius maximus comb. nov. and re-defined distribution of H. sauteri in China based on museum collections, previous studies, and this study (Triangle: Hebius maximus comb. nov., round: H. sauteri sauteri, square: H. sauteri bourreti. Type localities are colored red)." figureDoi="10.3897/herpetozoa.35.e94920.figure2" httpUri="https://binary.pensoft.net/fig/785721" pageId="0" pageNumber="265">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="EAE45D8FEC441CF17679202A573F3980" pageId="0" pageNumber="265" type="remarks">
|
||
<paragraph id="413D8A1806052DBD8C3BB89496A72AA4" pageId="0" pageNumber="265">Remarks.</paragraph>
|
||
<paragraph id="47B4CE3C77AF84ADE0ABC47C6A8BA059" pageId="0" pageNumber="265">
|
||
<taxonomicName id="E57FC0DBE4C5011BA63DB39EE9390DF4" authorityName="Van Denburgh" authorityYear="1909" family="Colubridae" genus="Natrix" kingdom="Animalia" lsidName="Natrix copei" order="Squamata" pageId="0" pageNumber="265" rank="species" species="copei">
|
||
<emphasis id="2EF9B8CBE684791EBB98E9E3DF6B9FAD" italics="true" pageId="0" pageNumber="265">Natrix copei</emphasis>
|
||
</taxonomicName>
|
||
(Van Denburgh, 1909) was described by
|
||
<bibRefCitation id="90FD4282ADFEA196D22C850BC43A8852" author="Van Denburgh, J" journalOrPublisher="Proceedings of the California Academy of Sciences" pageId="0" pageNumber="265" pagination="49 - 56" refId="B48" refString="Van Denburgh, J, 1909. New and previously unrecorded species of reptiles and amphibians from the island of Formosa. Proceedings of the California Academy of Sciences 4 (3): 49 - 56" title="New and previously unrecorded species of reptiles and amphibians from the island of Formosa." volume="4" year="1909">Van Denburgh (1909)</bibRefCitation>
|
||
based on specimen form
|
||
<normalizedToken id="9408BA3D69E007CAF76F8E67421B2F04" originalValue="“Kanshirei”">"Kanshirei"</normalizedToken>
|
||
(now Guanziling), Tainan County, Taiwan Province, China, the description of this species was published just 20 days after that of
|
||
<taxonomicName id="7969D0496B0E50C34F6D94A9DA971095" authorityName="Boulenger" authorityYear="1909" family="Colubridae" genus="Tropidonotus" kingdom="Animalia" lsidName="Tropidonotus sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="834E6A7E44F104A2206CDE90A0A33B8F" italics="true" pageId="0" pageNumber="265">Tropidonotus sauteri</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation id="B2FCC0D8E0C0C8815E211476516EB888" DOI="https://doi.org/10.1080/00222930908692704" author="Boulenger, GA" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="265" pagination="492 - 495" refId="B1" refString="Boulenger, GA, 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. The Annals and Magazine of Natural History 8 (4): 492 - 495, DOI: https://doi.org/10.1080/00222930908692704" title="Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa." url="https://doi.org/10.1080/00222930908692704" volume="8" year="1909">Boulenger (1909)</bibRefCitation>
|
||
(now
|
||
<taxonomicName id="7729B62CED6E172836F9617FCADD76FA" baseAuthorityName="Boulenger" baseAuthorityYear="1909" class="Reptilia" family="Colubridae" genus="Hebius" kingdom="Animalia" lsidName="Hebius sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="710BB64D9F0B5C7BC3220941DEC4B9F6" italics="true" pageId="0" pageNumber="265">Hebius sauteri</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation id="BA2F5C5205EB2A7E3A301307300D653D" DOI="https://doi.org/10.5479/si.00963801.1731.91" author="Stejneger, LH" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="265" pagination="91 - 114" refId="B46" refString="Stejneger, LH, 1910. The batrachians and reptiles of Formosa. Proceedings of the United States National Museum 38 (1731): 91 - 114, DOI: https://doi.org/10.5479/si.00963801.1731.91" title="The batrachians and reptiles of Formosa." url="https://doi.org/10.5479/si.00963801.1731.91" volume="38" year="1910">Stejneger 1910</bibRefCitation>
|
||
),
|
||
<bibRefCitation id="25DA238E6252C239DBAB1C00621A79AA" DOI="https://doi.org/10.5479/si.00963801.1731.91" author="Stejneger, LH" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="265" pagination="91 - 114" refId="B46" refString="Stejneger, LH, 1910. The batrachians and reptiles of Formosa. Proceedings of the United States National Museum 38 (1731): 91 - 114, DOI: https://doi.org/10.5479/si.00963801.1731.91" title="The batrachians and reptiles of Formosa." url="https://doi.org/10.5479/si.00963801.1731.91" volume="38" year="1910">Stejneger (1910)</bibRefCitation>
|
||
considered
|
||
<taxonomicName id="35BA8B99E8BC05B3B0AB122635E7C505" authorityName="Van Denburgh" authorityYear="1909" family="Colubridae" genus="Natrix" kingdom="Animalia" lsidName="Natrix copei" order="Squamata" pageId="0" pageNumber="265" rank="species" species="copei">
|
||
<emphasis id="F77D05CA2FAE666B6A2225EDAE5B04BB" italics="true" pageId="0" pageNumber="265">Natrix copei</emphasis>
|
||
</taxonomicName>
|
||
a synonym of
|
||
<taxonomicName id="09537BE472BD6EA973DE1B3077AA2723" authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" authorityYear="2022" baseAuthorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li" baseAuthorityYear="1909" family="Colubridae" genus="Natrix" kingdom="Animalia" lsidName="Natrix sauteri" order="Squamata" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="A33D9E7BF1AD26546078E2839E0CBE3E" italics="true" pageId="0" pageNumber="265">Natrix sauteri</emphasis>
|
||
</taxonomicName>
|
||
. We examined the paratype of
|
||
<taxonomicName id="98F48637153BD2637DA67B75CF6AE73A" authorityName="Van Denburgh" authorityYear="1909" family="Colubridae" genus="Natrix" kingdom="Animalia" lsidName="Natrix copei" order="Squamata" pageId="0" pageNumber="265" rank="species" species="copei">
|
||
<emphasis id="8D4AF2E7CA018344F807D89A958CE6AE" italics="true" pageId="0" pageNumber="265">Natrix copei</emphasis>
|
||
</taxonomicName>
|
||
, and its morphological characters are in the range of variation of
|
||
<taxonomicName id="F1B720A1FDAF90F376445E0B78A71DD9" lsidName="H. sauteri" pageId="0" pageNumber="265" rank="species" species="sauteri">
|
||
<emphasis id="4347F8210EB0AA0E25265CF6041A0BA4" italics="true" pageId="0" pageNumber="265">H. sauteri</emphasis>
|
||
</taxonomicName>
|
||
, which also supports the treatment of
|
||
<taxonomicName id="4B87F41C739184D93B32EC1D3F6158D3" authorityName="Van Denburgh" authorityYear="1909" family="Colubridae" genus="Natrix" kingdom="Animalia" lsidName="Natrix copei" order="Squamata" pageId="0" pageNumber="265" rank="species" species="copei">
|
||
<emphasis id="FA2BCBFB79818258E0C20E3D9DCF6318" italics="true" pageId="0" pageNumber="265">Natrix copei</emphasis>
|
||
</taxonomicName>
|
||
as an objective synonym by
|
||
<bibRefCitation id="2109B10940D91F69DDCB4F1BE8153751" DOI="https://doi.org/10.5479/si.00963801.1731.91" author="Stejneger, LH" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="265" pagination="91 - 114" refId="B46" refString="Stejneger, LH, 1910. The batrachians and reptiles of Formosa. Proceedings of the United States National Museum 38 (1731): 91 - 114, DOI: https://doi.org/10.5479/si.00963801.1731.91" title="The batrachians and reptiles of Formosa." url="https://doi.org/10.5479/si.00963801.1731.91" volume="38" year="1910">Stejneger (1910)</bibRefCitation>
|
||
. The taxonomic status of the subspecies
|
||
<taxonomicName id="3EC7B9940ABCF60BE68702CB97CF14BE" lsidName="H. s. subsp. bourreti" pageId="0" pageNumber="265" rank="subSpecies" species="s." subSpecies="bourreti">
|
||
<emphasis id="6B2396CA7275FC1517CD294A61D0D36F" italics="true" pageId="0" pageNumber="265">H. s. bourreti</emphasis>
|
||
</taxonomicName>
|
||
is discussed below.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |