<documentID-CLB-Dataset="56712"ID-DOI="http://dx.doi.org/10.3897/herpetozoa.35.e94920"ID-GBIF-Dataset="e8f01291-4827-458e-9575-32637e9020fb"ID-Pensoft-Pub="2682-955X-35-265"ID-Pensoft-UUID="E04369D7C9EB5294A6DD1780BD27542F"ID-ZooBank="29C2ED3FBC5E45E78E58FCC93B988455"ModsDocID="2682-955X-35-265"checkinTime="1671892101870"checkinUser="pensoft"docAuthor="Li, Mao-Liang, Ren, Jin-Long, Huang, Jun-Jie, Lyu, Zhi-Tong, Qi, Shuo, Jiang, Ke, Wang, Ying-Yong & Li, Jia-Tang"docDate="2022"docId="C3E61DC2F9B05389A33A6DE55B6983C7"docLanguage="en"docName="Herpetozoa 35: 265-282"docOrigin="Herpetozoa 35"docPubDate="2022-12-21"docSource="http://dx.doi.org/10.3897/herpetozoa.35.e94920"docTitle="Hebius sauteri"docType="treatment"docVersion="6"id="E04369D7C9EB5294A6DD1780BD27542F"lastPageNumber="265"masterDocId="E04369D7C9EB5294A6DD1780BD27542F"masterDocTitle="On the validity of Hebius sauteri maximus (Malnate, 1962) (Squamata, Natricidae), with the redescription of H. maximus comb. nov. and H. sauteri (Boulenger, 1909)"masterLastPageNumber="282"masterPageNumber="265"pageNumber="265"updateTime="1732848484904"updateUser="ExternalLinkService">
<mods:titleid="F3DC316266D1B95B721C00F3E2C7E738">On the validity of Hebius sauteri maximus (Malnate, 1962) (Squamata, Natricidae), with the redescription of H. maximus comb. nov. and H. sauteri (Boulenger, 1909)</mods:title>
<mods:affiliationid="EFC780B22C9582934965FC6D26CAFB09">College of Life Science and Technology, Huazhong Agricultural University, Wuhan 430070, Hubei, China & Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:affiliationid="6C3B2F6B48C6065AC615928115C37462">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:affiliationid="F12C373B3D747DEF7A24757714455E5B">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:affiliationid="390737D395BB6BAB995C15CEA2230AAD">School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
<mods:affiliationid="482ACDCECC964BC66849E17A57F40FE1">School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
<mods:affiliationid="7456887EA1C1AF9BAED27D09DE901606">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:affiliationid="3DC7317A0CA5190A54A6760FA7B83A4F">School of Life Sciences, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
<mods:affiliationid="BDD22345A89C73B5B0E2EED495129D42">Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu, Chengdu 610041, Sichuan, China & School of Ecology, Sun Yat-Sen University, Guangzhou 510275, Guangdong, China</mods:affiliation>
<figureCitationid="C66A67E243EB8A301B4103B55C1DF564"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Distribution of Hebius maximus comb. nov. and re-defined distribution of H. sauteri in China based on museum collections, previous studies, and this study (Triangle: Hebius maximus comb. nov., round: H. sauteri sauteri, square: H. sauteri bourreti. Type localities are colored red)."figureDoi="10.3897/herpetozoa.35.e94920.figure2"httpUri="https://binary.pensoft.net/fig/785721"pageId="0"pageNumber="265">Figs 2</figureCitation>
<figureCitationid="5A356BCF9DC432F52F4015EAA4EBD0FE"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between Hebius sauteri and Hebius maximus comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang. A-C. Hebius maximus comb. nov. (A. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China; B. CIB 8482, Bijie City, Guizhou, China; C. CIB 8458, Yingjing County, Sichuan, China) D-F. H. sauteri (CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China)."figureDoi="10.3897/herpetozoa.35.e94920.figure3"httpUri="https://binary.pensoft.net/fig/785722"pageId="0"pageNumber="265">, 3</figureCitation>
<figureCitationid="45B4593294F63EE1A498AB90945364AA"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Hebius sauteri. A. CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; B. CIB 118517, Xinyong Village, Dehua County, Fujian, China. Photographs by Jin-Long Ren."figureDoi="10.3897/herpetozoa.35.e94920.figure4"httpUri="https://binary.pensoft.net/fig/785723"pageId="0"pageNumber="265">, 4</figureCitation>
<figureCitationid="47CDD8BEA9B058853E07474AB5C75A3D"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Preserved specimen of H. sauteri (CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang."figureDoi="10.3897/herpetozoa.35.e94920.figure5"httpUri="https://binary.pensoft.net/fig/785724"pageId="0"pageNumber="265">, 5</figureCitation>
<figureCitationid="6711F286AF691C194FD8CE4341D7F4FF"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Photographs of the hemipenis of H. sauteri (SYS r 000323, Mt. Jinggang, Jiangxi Province, China), photos from left to right showing sulcate, asulcate, and apical sides of the hemipenis. Photographs by Jun-Jie Huang."figureDoi="10.3897/herpetozoa.35.e94920.figure6"httpUri="https://binary.pensoft.net/fig/785725"pageId="0"pageNumber="265">, 6</figureCitation>
<taxonomicNameid="571B74CDD36FFA77CFFD156C4BE90A6C"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"baseAuthorityYear="1909"family="Colubridae"genus="Natrix"higherTaxonomySource="treatment-meta"kingdom="Animalia"lsidName="Natrix sauteri"order="Squamata"pageId="0"pageNumber="265"rank="species"species="sauteri">Natrix sauteri</taxonomicName>
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<taxonomicNameid="C989E9719A4B01BE852C245C4B345AEE"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"baseAuthorityYear="1909"family="Colubridae"genus="Amphiesma"higherTaxonomySource="treatment-meta"kingdom="Animalia"lsidName="Amphiesma sauteri"order="Squamata"pageId="0"pageNumber="265"rank="species"species="sauteri">Amphiesma sauteri</taxonomicName>
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<taxonomicNameid="14ACE034BA3C2C8598A5B22D9CF5E3CA"lsidName="H. s. subsp. sauteri"pageId="0"pageNumber="265"rank="subSpecies"species="s."subSpecies="sauteri">
<emphasisid="C73D3B2D8B89353DFEC4AEE136720267"italics="true"pageId="0"pageNumber="265">H. s. sauteri</emphasis>
can be distinguished from its congeners by the following features: (1) body size small to medium, maximum ToL 455 mm; (2) dorsal scales rows 17-17-17, all weekly keeled or smooth on outmost one or two rows; (3) ventral scales 116-133, subcaudals 60-86, paired; (4) maxillary teeth 23-27, last two or three distinctly enlarged teeth, without diastema; (5) preocular single, occasionally two; (6) supralabials 6-8, 2nd or 2nd-3rd border the loreal; (7) infralabials 6-9, mostly edged with black on their posterior border; (8) single white to orange stripe extends from
<paragraphid="849B2490C215F2E743743C7166E70EB5"pageId="0"pageNumber="265">Head moderately distinct from neck; body size small to medium, ToL 145-455 mm (282-370 mm in males, 145-455 mm in females), TL/ToL 0.24-0.33 (0.27-0.31 in males, 0.24-0.33 in females). Eye large, pupil round.</paragraph>
Body scale in 17-17-17 rows, weekly keeled except outer 1-4 rows smooth anteriorly, outer 0-2 rows smooth at midbody and outer 0-1 row smooth posteriorly; ventrals 116-133 (125-130 in males, 116-133 in females); subcaudals 60-86 (60-86 in females, 65-84 in males), paired.
Rostral broad, visible from above; internasals paired, nearly triangular, curved outwards, not in contact with loreal; prefrontals paired, nearly in a diamond shape, wider than long, border on loreal laterally, preocular and supraocular posteriorly; frontal hexagonal, longer than wide, embedded into parietal posteriorly; parietal paired, much longer than wide; nasals wider than high, divided below nostril, borders 1st-2nd supralabials; loreal single, rarely divided (right side of CIB 8481), borders 2nd or 2nd-3rd supralabials, not entering orbit; preocular 1-2; supraocular single, narrowed anteriorly, much wider than high, not in contact with parietal; postoculars 2-3, rarely 4 (left side of CIB 8483); supralabials 6-8, 5th, 6th or 7th highest, 3rd -4th entering orbit, rarely 2nd-3rd (right side of CIB 8480), 3rd only (both sides of CIB 8479) or 4th-5th (left side of CIB 8477); infralabials 6-9, first pair in contact behind the mental, IfL 1-3, 1-4 or 1-5 border the anterior chin shields; temporals variable, in two or three rows, 1-2 + 1-3 + 0/2-3; chin shields paired, posterior pair longer (Figs
<figureCitationid="CA1665ECBAEACB84932173A5E9383BEE"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between Hebius sauteri and Hebius maximus comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang. A-C. Hebius maximus comb. nov. (A. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China; B. CIB 8482, Bijie City, Guizhou, China; C. CIB 8458, Yingjing County, Sichuan, China) D-F. H. sauteri (CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China)."figureDoi="10.3897/herpetozoa.35.e94920.figure3"httpUri="https://binary.pensoft.net/fig/785722"pageId="0"pageNumber="265">3</figureCitation>
<figureCitationid="57B29337670EFE96B1834B822340565A"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Preserved specimen of H. sauteri (CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang."figureDoi="10.3897/herpetozoa.35.e94920.figure5"httpUri="https://binary.pensoft.net/fig/785724"pageId="0"pageNumber="265">5</figureCitation>
<tableCitationid="8BCB7C14828F75384AFC62212FB2AF66"captionStart="Table 4"captionStartId="T4"captionText="Table 4. Morphological characters of H. maximus and H. sauteri. For specimens with asymmetrical cephalic scales, the scale count was given as " left / right ", " - " indicates missing data."httpUri="http://table.plazi.org/id/CEF25E8772CA7CD1980C3D0E96F6D81F"pageId="0"pageNumber="265"tableUuid="CEF25E8772CA7CD1980C3D0E96F6D81F">4</tableCitation>
The description of the hemipenis is based on an adult specimen (SYS r000323) from Mt. Jinggang, Jiangxi, southeastern China. Left side of hemipenis fully everted, right side remained
The everted hemipenis is thin and short, Y-shaped, shallowly bilobed. Both sulcate and asulcate surfaces densely ornamented with spinules, a large basal hook present at proximal part.
<figureCitationid="C43E691CD7AEFB26DFD0894425435607"captionStart="Figure 6"captionStartId="F6"captionText="Figure 6. Photographs of the hemipenis of H. sauteri (SYS r 000323, Mt. Jinggang, Jiangxi Province, China), photos from left to right showing sulcate, asulcate, and apical sides of the hemipenis. Photographs by Jun-Jie Huang."figureDoi="10.3897/herpetozoa.35.e94920.figure6"httpUri="https://binary.pensoft.net/fig/785725"pageId="0"pageNumber="265">6</figureCitation>
Based on two adult female specimens from Guangdong (CIB 118516) and Fujian (CIB 118517), China. Dorsal surface of the head reddish-brown with irregular grayish-green spots, a pair of elliptic orangish spots symmetrically present on inner sides of parietals, just separated by parietal sutures; ventral side of head whitish; 1st-5th supralabials white with black edges posteriorly, 6th or 6-7th brown on their upper part; infralabials white with black edge between each scale; an orange stripe extends from
to dorsal side of neck, 1-2 scales wide; dorsal surface of body and tail dark brown to reddish-brown, top with or without an ill-defined dark olive stripe scattered with black spots extends from neck to the end, a reddish-brown lateral streak on 4th-6th body scale rows extends from neck to tail with series of orange spots separated by 2-3 scales above; ventral and subcaudal sides cream white, with black spots on both ends of the ventral scale forming a chain-like pattern on the ventral view; sclera brown (Fig.
<figureCitationid="670F1D6781FBCF06FA9E2ABD707D98A7"captionStart="Figure 4"captionStartId="F4"captionText="Figure 4. Hebius sauteri. A. CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; B. CIB 118517, Xinyong Village, Dehua County, Fujian, China. Photographs by Jin-Long Ren."figureDoi="10.3897/herpetozoa.35.e94920.figure4"httpUri="https://binary.pensoft.net/fig/785723"pageId="0"pageNumber="265">4</figureCitation>
<taxonomicNameid="D30A6B68FDB93DB1D62E33BD9C107240"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Malnate"baseAuthorityYear="1962"class="Reptilia"family="Colubridae"genus="Hebius"kingdom="Animalia"lsidName="Hebius maximus"order="Squamata"pageId="0"pageNumber="265"rank="species"species="maximus">
<taxonomicNameid="3E0759DCCF32B829E624B307F038537D"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Malnate"baseAuthorityYear="1962"class="Reptilia"family="Colubridae"genus="Hebius"kingdom="Animalia"lsidName="Hebius maximus"order="Squamata"pageId="0"pageNumber="265"rank="species"species="maximus">
<taxonomicNameid="DA4CE7B3FD7EB4807EF06D569C2AD56B"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Malnate"baseAuthorityYear="1962"class="Reptilia"family="Colubridae"genus="Hebius"kingdom="Animalia"lsidName="Hebius maximus"order="Squamata"pageId="0"pageNumber="265"rank="species"species="maximus">
<taxonomicNameid="8B73260699AE7DDFB4141B4B68F118BD"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Malnate"baseAuthorityYear="1962"class="Reptilia"family="Colubridae"genus="Hebius"kingdom="Animalia"lsidName="Hebius maximus"order="Squamata"pageId="0"pageNumber="265"rank="species"species="maximus">
(SYS r000323, Mt. Jinggang, Jiangxi Province, China), photos from left to right showing sulcate, asulcate, and apical sides of the hemipenis. Photographs by Jun-Jie Huang.
Largely the same as in life except dorsal side of head brown to dark brown, scattered with small black spots; ventral side of head cream white; a pale yellow stripe extends from
to dorsal side of neck, 1-2 scales wide; dorsal of body and tail brown to dark brown, a lighter lateral streak on 4th-6th body scale rows presents or not, covered with series of cream white spots separated by 2-3 scales; ventral and subcaudal sides cream to pale white; sclera black (Figs
<figureCitationid="DC01DD24C547C671BC9C002EEC429993"captionStart="Figure 3"captionStartId="F3"captionText="Figure 3. Dorsal (left), lateral (middle) and ventral (right) of the head comparisons between Hebius sauteri and Hebius maximus comb. nov. Row D photographed by Kai Wang, others were photographed by Jun-Jie Huang. A-C. Hebius maximus comb. nov. (A. CIB 118635, Mt. Laojun, Pingshan County, Sichuan, China; B. CIB 8482, Bijie City, Guizhou, China; C. CIB 8458, Yingjing County, Sichuan, China) D-F. H. sauteri (CAS 18984, Pingtung County, Taiwan, China; CIB 118516, Yinpingshan, Dongguan City, Guangdong, China; CIB 118517, Daiyun, Dehua County, Fujian, China)."figureDoi="10.3897/herpetozoa.35.e94920.figure3"httpUri="https://binary.pensoft.net/fig/785722"pageId="0"pageNumber="265">3</figureCitation>
<figureCitationid="25BEB723E50299CC78A796A87B4A4478"captionStart="Figure 5"captionStartId="F5"captionText="Figure 5. Preserved specimen of H. sauteri (CAS 18988), from the type locality of the species, Guanziling, Tainan City, Taiwan, China. Not to scale. Photographs by Kai Wang."figureDoi="10.3897/herpetozoa.35.e94920.figure5"httpUri="https://binary.pensoft.net/fig/785724"pageId="0"pageNumber="265">5</figureCitation>
<bibRefCitationid="3CE4105888F53A3E0D61713BAEAFF35E"DOI="https://doi.org/10.1080/00222930908692704"author="Boulenger, GA"journalOrPublisher="The Annals and Magazine of Natural History"pageId="0"pageNumber="265"pagination="492 - 495"refId="B1"refString="Boulenger, GA, 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. The Annals and Magazine of Natural History 8 (4): 492 - 495, DOI: https://doi.org/10.1080/00222930908692704"title="Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa."url="https://doi.org/10.1080/00222930908692704"volume="8"year="1909">Boulenger 1909</bibRefCitation>
This species inhabits flatlands and mountain regions up to 1,500 m and prefers terrestrial microhabitat such as grassland, woodland, and bushes near water bodies; diurnal, occasionally found at night, preys include earthworms, slugs, snails and tadpoles; oviparous (this study;
<bibRefCitationid="9F68429525016E1CBC71E972D1A3FE5A"author="Pope, CH"journalOrPublisher="The American Museum of Natural History, New York"pageId="0"pageNumber="265"refId="B37"refString="Pope, CH, 1935. The reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia (Vol. X). The American Museum of Natural History, New York"title="The reptiles of China. Turtles, Crocodilians, Snakes, Lizards. Natural History of Central Asia (Vol. X)."year="1935">Pope 1935</bibRefCitation>
<bibRefCitationid="2C7C7BEDEB91347D68D11498CC5BB917"author="Xiang, GS"journalOrPublisher="Owl Press, Taipei"pageId="0"pageNumber="265"refId="B53"refString="Xiang, GS, Li, PX, Yang, YR, 2009. Colored illustrations of amphibians and reptiles of Taiwan. Owl Press, Taipei"title="Colored illustrations of amphibians and reptiles of Taiwan."year="2009">Xiang et al. 2009</bibRefCitation>
is currently known to be reported in Vietnam (Ngan Son, Lang Son, Tam Dao and Ba Vi), and China, including Taiwan (Kaohsiung County [type locality]), Fujian (Dehua County), Anhui (Huangshan County), Jiangxi (Jinggangshan City, Longnan County), Hunan (Yizhang County), Guangdong (Lianping County, Xinyi County, Deqing County, Ruyuan County, Dongguan City), Guangxi (Jinxiu County, Quanzhou County, Rongxian County, Beiliu County,
County), Guizhou (Libo County, Leishan County), Yunnan (Jinping County), Hainan (Qiongzhong County, Wuzhishan County, Baisha County, Lingshui County) (
<bibRefCitationid="23D936C570B10C4EAE7EE9437E2374FB"author="Deuve, J"journalOrPublisher="M., Paris"pageId="0"pageNumber="265"refId="B10"refString="Deuve, J, 1970. Serpents du Laos. Memoirs O.R.S.T.O. M., Paris"title="Serpents du Laos. Memoirs O. R. S. T. O."year="1970">Deuve 1970</bibRefCitation>
<bibRefCitationid="565EF43FB42BE5EA4CA7872B8E0DAF8A"author="Zhang, YX"journalOrPublisher="Guangxi Normal University Press, Guilin"pageId="0"pageNumber="265"refId="B59"refString="Zhang, YX, 2009. Herpetology in Guangxi. Guangxi Normal University Press, Guilin"title="Herpetology in Guangxi."year="2009">Zhang 2009</bibRefCitation>
<figureCitationid="51574675F7DDB91425DE180A881F0352"captionStart="Figure 2"captionStartId="F2"captionText="Figure 2. Distribution of Hebius maximus comb. nov. and re-defined distribution of H. sauteri in China based on museum collections, previous studies, and this study (Triangle: Hebius maximus comb. nov., round: H. sauteri sauteri, square: H. sauteri bourreti. Type localities are colored red)."figureDoi="10.3897/herpetozoa.35.e94920.figure2"httpUri="https://binary.pensoft.net/fig/785721"pageId="0"pageNumber="265">2</figureCitation>
<bibRefCitationid="90FD4282ADFEA196D22C850BC43A8852"author="Van Denburgh, J"journalOrPublisher="Proceedings of the California Academy of Sciences"pageId="0"pageNumber="265"pagination="49 - 56"refId="B48"refString="Van Denburgh, J, 1909. New and previously unrecorded species of reptiles and amphibians from the island of Formosa. Proceedings of the California Academy of Sciences 4 (3): 49 - 56"title="New and previously unrecorded species of reptiles and amphibians from the island of Formosa."volume="4"year="1909">Van Denburgh (1909)</bibRefCitation>
<bibRefCitationid="B2FCC0D8E0C0C8815E211476516EB888"DOI="https://doi.org/10.1080/00222930908692704"author="Boulenger, GA"journalOrPublisher="The Annals and Magazine of Natural History"pageId="0"pageNumber="265"pagination="492 - 495"refId="B1"refString="Boulenger, GA, 1909. Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa. The Annals and Magazine of Natural History 8 (4): 492 - 495, DOI: https://doi.org/10.1080/00222930908692704"title="Descriptions of four new frogs and a new snake discovered by Mr. H. Sauter in Formosa."url="https://doi.org/10.1080/00222930908692704"volume="8"year="1909">Boulenger (1909)</bibRefCitation>
<bibRefCitationid="BA2F5C5205EB2A7E3A301307300D653D"DOI="https://doi.org/10.5479/si.00963801.1731.91"author="Stejneger, LH"journalOrPublisher="Proceedings of the United States National Museum"pageId="0"pageNumber="265"pagination="91 - 114"refId="B46"refString="Stejneger, LH, 1910. The batrachians and reptiles of Formosa. Proceedings of the United States National Museum 38 (1731): 91 - 114, DOI: https://doi.org/10.5479/si.00963801.1731.91"title="The batrachians and reptiles of Formosa."url="https://doi.org/10.5479/si.00963801.1731.91"volume="38"year="1910">Stejneger 1910</bibRefCitation>
<bibRefCitationid="25DA238E6252C239DBAB1C00621A79AA"DOI="https://doi.org/10.5479/si.00963801.1731.91"author="Stejneger, LH"journalOrPublisher="Proceedings of the United States National Museum"pageId="0"pageNumber="265"pagination="91 - 114"refId="B46"refString="Stejneger, LH, 1910. The batrachians and reptiles of Formosa. Proceedings of the United States National Museum 38 (1731): 91 - 114, DOI: https://doi.org/10.5479/si.00963801.1731.91"title="The batrachians and reptiles of Formosa."url="https://doi.org/10.5479/si.00963801.1731.91"volume="38"year="1910">Stejneger (1910)</bibRefCitation>
<taxonomicNameid="09537BE472BD6EA973DE1B3077AA2723"authorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"authorityYear="2022"baseAuthorityName="Li & Ren & Huang & Lyu & Qi & Jiang & Wang & Li"baseAuthorityYear="1909"family="Colubridae"genus="Natrix"kingdom="Animalia"lsidName="Natrix sauteri"order="Squamata"pageId="0"pageNumber="265"rank="species"species="sauteri">
<bibRefCitationid="2109B10940D91F69DDCB4F1BE8153751"DOI="https://doi.org/10.5479/si.00963801.1731.91"author="Stejneger, LH"journalOrPublisher="Proceedings of the United States National Museum"pageId="0"pageNumber="265"pagination="91 - 114"refId="B46"refString="Stejneger, LH, 1910. The batrachians and reptiles of Formosa. Proceedings of the United States National Museum 38 (1731): 91 - 114, DOI: https://doi.org/10.5479/si.00963801.1731.91"title="The batrachians and reptiles of Formosa."url="https://doi.org/10.5479/si.00963801.1731.91"volume="38"year="1910">Stejneger (1910)</bibRefCitation>
<taxonomicNameid="3EC7B9940ABCF60BE68702CB97CF14BE"lsidName="H. s. subsp. bourreti"pageId="0"pageNumber="265"rank="subSpecies"species="s."subSpecies="bourreti">
<emphasisid="6B2396CA7275FC1517CD294A61D0D36F"italics="true"pageId="0"pageNumber="265">H. s. bourreti</emphasis>
