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<mods:title id="56C589C0F0A3FF8E06C8764214FF9DC8">Intersexual Arms Race and the Evolution of the Sphragis in Pteronymia Butterflies</mods:title>
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<mods:namePart id="65B51B48D91D87A55089B6451BAE40C6">Carvalho, Ana Paula S.</mods:namePart>
<mods:affiliation id="DDAEEB75835DD7176A14F9F783D4DDD5">Department of Entomology and Nematology, University of Florida, 1881 Natural Area Drive, Gainesville, FL 32608,</mods:affiliation>
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<mods:namePart id="8B67020C94F5817D03008A57DCF5A730">Mota, Luísa L.</mods:namePart>
<mods:affiliation id="91F9CF0D436EA4713ADEC1A734293E24">Departamento de Biologia Animal, Rua Monteiro Lobato 255, Instituto de Biologia, Universidade Estadual de Campinas, Campinas, SP 13083 - 862, Brazil, and</mods:affiliation>
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<mods:namePart id="543E1747B8F1E3118562C88A3CB9CAA7">Kawahara, Akito Y.</mods:namePart>
<mods:affiliation id="288282B67B268FA31467C4867D33D21F">Department of Entomology and Nematology, University of Florida, 1881 Natural Area Drive, Gainesville, FL 32608,</mods:affiliation>
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<subSubSection id="F59ED1ED4041C37AFF10FAEAFE34C43B" box="[146,468,1369,1389]" pageId="4" pageNumber="5" type="nomenclature">
<paragraph id="BD3B82664041C37AFF10FAEAFE34C43B" blockId="4.[146,468,1369,1389]" box="[146,468,1369,1389]" pageId="4" pageNumber="5">
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<taxonomicName id="7A84F9E54041C37AFF10FAEAFE34C43B" ID-CoL="6MPP3" authority="(Weeks, 1901)" baseAuthorityName="Weeks" baseAuthorityYear="1901" box="[146,468,1369,1389]" class="Insecta" family="Nymphalidae" genus="Pteronymia" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="andreas">
<emphasis id="8FF05E744041C37AFF10FAEAFEA9C43B" box="[146,329,1369,1389]" italics="true" pageId="4" pageNumber="5">Pteronymia andreas</emphasis>
<emphasis id="8FF05E744041C37AFECDFAE9FE34C43B" bold="true" box="[335,468,1369,1389]" pageId="4" pageNumber="5">(Weeks, 1901)</emphasis>
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<subSubSection id="F59ED1ED4041C37AFF30FAC7FDC7C65E" pageId="4" pageNumber="5" type="description">
<paragraph id="BD3B82664041C37AFF30FAC7FF1EC4DD" blockId="4.[146,771,1399,1800]" box="[178,254,1399,1419]" pageId="4" pageNumber="5">
<figureCitation id="25BF9EE34041C37AFF30FAC7FF1EC4DD" box="[178,254,1399,1419]" captionStart="Fig" captionStartId="5.[146,173,1466,1483]" captionTargetBox="[314,1273,148,1429]" captionTargetId="figure-218@5.[312,1275,147,1430]" captionTargetPageId="5" captionText="Fig. 25. The sterigma and sphragis of P. andreas (2), P. calgiria (3), P. obscuratus (4), and P.latilla (5). A: ventral, sphragis absent; B: ventral,sphragis present; C: lateral, sphragis present; D: P. obscuratus with a spermatophore externally connected to the sphragis." figureDoi="http://doi.org/10.5281/zenodo.4551935" httpUri="https://zenodo.org/record/4551935/files/figure.png" pageId="4" pageNumber="5">Figure 2</figureCitation>
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<paragraph id="BD3B82664041C37AFF30FA24FE0DC493" blockId="4.[146,771,1399,1800]" pageId="4" pageNumber="5">
<emphasis id="8FF05E744041C37AFF30FA24FE82C4FE" box="[178,354,1428,1448]" italics="true" pageId="4" pageNumber="5">Females examined:</emphasis>
22. 1 with a complete sphragis; 12 bearing incomplete or amorphous sphragides.
</paragraph>
<paragraph id="BD3B82664041C37AFF30FA60FE9AC76D" blockId="4.[146,771,1399,1800]" pageId="4" pageNumber="5">
<emphasis id="8FF05E744041C37AFF30FA60FEC3C4B4" box="[178,291,1488,1506]" italics="true" pageId="4" pageNumber="5">Description:</emphasis>
Coloration—golden-brown to yellowish off-white. Shape—plate-like. Structure—mostly hollow. Scales—found in some specimens, but not ordered, probably accidentally attached. Complexity category—2.
</paragraph>
<paragraph id="BD3B82664041C37AFF30F9F5FDC7C65E" blockId="4.[146,771,1399,1800]" pageId="4" pageNumber="5">
<emphasis id="8FF05E744041C37AFF30F9F5FEFBC70E" box="[178,283,1605,1624]" italics="true" pageId="4" pageNumber="5">Comments:</emphasis>
The complete sphragis that we observed was likely deformed due to the way it was attached to the sterigma. Nearly all (12/13, 92%) of the sphragides observed for this species were incomplete. Incomplete sphragides were in the form of a plate, and always completely internalized inside the bowl-like sterigma. The amount of sphragidal material was variable, but often it only covered the ostium bursae and immediate surroundings.
</paragraph>
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