392 lines
58 KiB
XML
392 lines
58 KiB
XML
<document id="1F6B9BE611BFE1D87F1677C438B39501" ID-CLB-Dataset="7175" ID-DOI="10.5252/geodiversitas2021v43a9" ID-GBIF-Dataset="04426a8e-c179-4af0-8e23-d72908e8b4cc" ID-ISSN="1638-9395" ID-Zenodo-Dep="4720776" ID-ZooBank="urn:lsid:zoobank.org:pub:11D0D852-39D7-449C-9EB3-C3D804114556" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1619445770811" checkinUser="felipe" docAuthor="Georgalis, Georgios L., Čerňanský, Andrej & Klembara, Jozef" docDate="2021" docId="03A1633BFF87FFE8338FFB2BFB843F01" docLanguage="en" docName="Geodiversitas.43.9.219-293.pdf" docOrigin="Geodiversitas 43 (9)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Palaeovaranus lismonimenos Georgalis & Čerňanský & Klembara 2021, n. sp." docType="treatment" docUuid="3C906259-DAB5-4B97-A1BB-7860695A76B8" docUuidSource="ZooBank" docVersion="10" lastPageNumber="258" masterDocId="FF981B43FFA3FFC1303EFFB8FA5E380D" masterDocTitle="Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material" masterLastPageNumber="293" masterPageNumber="219" pageNumber="253" updateTime="1732603017081" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="C05CD7348E87CBC96BDCCA5D4494A1E9">
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<mods:title id="381F5E91D4C3C0811CAB9A4B80F20D95">Osteological atlas of new lizards from the Phosphorites du Quercy (France), based on historical, forgotten, fossil material</mods:title>
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<mods:name id="12D3591C98E27E9EBEF698D4601DDCE8" type="personal">
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<mods:roleTerm id="B2E0BF10188B1A93C099C6B47F60289C">Author</mods:roleTerm>
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<mods:namePart id="BF2BB78694B51FA280EBB111F69D86FE">Georgalis, Georgios L.</mods:namePart>
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<mods:affiliation id="9B1CCE9634BA490530F6E8F51A6F7E45">Palaeontological Institute and Museum, University of Zurich, Karl Schmid-Strasse 4, 8006 Zurich (Switzerland) and Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia) and Department of Earth Sciences, University of Torino, Via Valperga Caluso 35, 10125 Turin (Italy)</mods:affiliation>
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<mods:nameIdentifier id="2A1DC5FB196106852AF8BEF1EEBF3D8D" type="email">georgios.georgalis@pim.uzh.ch</mods:nameIdentifier>
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<mods:name id="22243E96DEF919DBDF33583FECFE3041" type="personal">
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<mods:namePart id="24AD0540B6A921C854958725A3597EC9">Čerňanský, Andrej</mods:namePart>
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<mods:affiliation id="F2A0A5EB943C63A43E1AFF2F3FCE2FA2">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:namePart id="D94B2D9FC9047229BF2CDC5801EA3F54">Klembara, Jozef</mods:namePart>
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<mods:affiliation id="9D6231DD9A77A449F93AA304FF108101">Department of Ecology, Laboratory of Evolutionary Biology, Faculty of Natural Sciences, Comenius University in Bratislava, Mlynská dolina, 84215 Bratislava (Slovakia)</mods:affiliation>
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<mods:title id="B291F54196AFBBBE0BD24B080D5655CC">Geodiversitas</mods:title>
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<mods:part id="7033B9337028403A40B8232A693562A2">
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<mods:date id="0535D0FB1CF14F68535DAE24C391EDAA">2021</mods:date>
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<mods:number id="CEC708F680A6DBCB2CE57FF8C852395C">2021-04-22</mods:number>
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<treatment id="03A1633BFF87FFE8338FFB2BFB843F01" ID-DOI="http://doi.org/10.5281/zenodo.4721473" ID-GBIF-Taxon="181601113" ID-Zenodo-Dep="4721473" LSID="urn:lsid:zoobank.org:act:3C906259-DAB5-4B97-A1BB-7860695A76B8" httpUri="http://treatment.plazi.org/id/03A1633BFF87FFE8338FFB2BFB843F01" lastPageId="41" lastPageNumber="258" pageId="36" pageNumber="253" scope_family="Palaeovaranidae" scope_order="Squamata">
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<subSubSection id="C31281A6FF87FFE5338FFB2BFF753CA3" box="[945,1323,1171,1198]" pageId="36" pageNumber="253" type="nomenclature">
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<paragraph id="8BB7D22DFF87FFE5338FFB2BFF753CA3" blockId="36.[945,1323,1171,1230]" box="[945,1323,1171,1198]" pageId="36" pageNumber="253">
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<heading id="D0FF6541FF87FFE5338FFB2BFF753CA3" box="[945,1323,1171,1198]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="253" reason="2">
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<taxonomicName id="4C08A9AEFF87FFE5338FFB2BFEB73CA3" authority="Georgalis & Čerňanský & Klembara, 2021" authorityName="Georgalis & Čerňanský & Klembara" authorityYear="2021" box="[945,1257,1171,1198]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="253" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
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<emphasis id="B97C0E3FFF87FFE5338FFB2BFEB73CA3" bold="true" box="[945,1257,1171,1198]" italics="true" pageId="36" pageNumber="253">Palaeovaranus lismonimenos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24FB344FF87FFE534CEFB2CFF753CA3" box="[1264,1323,1172,1198]" pageId="36" pageNumber="253" rank="species">n. sp.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31281A6FF87FFE5338BFB0CFFF13D0A" pageId="36" pageNumber="253" type="description">
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<paragraph id="8BB7D22DFF87FFE5338BFB0CFF793CC3" blockId="36.[945,1323,1171,1230]" box="[949,1319,1204,1230]" pageId="36" pageNumber="253">
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(
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<figureCitation id="1333CEA8FF87FFE53380FB0CFE1C3CC3" box="[958,1090,1204,1230]" captionStart="FIG" captionStartId="43.[132,143,1820,1837]" captionTargetBox="[124,1472,185,1797]" captionTargetId="figure-78@43.[116,1472,247,1772]" captionTargetPageId="43" captionText="FIG. 37. — Palaeovaranus cayluxi Zittel, 1887-1890 and Palaeovaranus lismonimenos n. sp. A, C, virtual 3D models of parietal NHMW 2019/0048/0001 of Palaeovaranus cayluxi in dorsal (A) and ventral (C) views; B, D, virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in dorsal (B) and ventral (D) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720891" httpUri="https://zenodo.org/record/4720891/files/figure.png" pageId="36" pageNumber="253">Figs 37B, D</figureCitation>
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;
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<figureCitation id="1333CEA8FF87FFE53470FB0CFED13CC3" box="[1102,1167,1204,1230]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="44.[132,143,1323,1340]" captionStartId-1="45.[132,143,1318,1335]" captionStartId-2="46.[133,143,881,898]" captionTargetBox-0="[149,1430,217,1279]" captionTargetBox-1="[132,1455,181,1289]" captionTargetBox-2="[360,1236,153,859]" captionTargetId-0="figure-367@44.[145,1442,211,1284]" captionTargetId-1="figure-365@45.[132,1455,228,1276]" captionTargetId-2="figure-212@46.[472,1115,215,839]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionText-0="FIG. 38. — Palaeovaranus lismonimenos n. sp. Virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in posteroventral (A), dorsolateral (B), and anterodorsal (C) views. Scale bar: 5 mm." captionText-1="FIG. 39. — Palaeovaranus lismonimenos n. sp. Photographs of the holotype parietal NHMW 2019/0047/0001 in dorsal (A), ventral (B), and right lateral (C) views. Scale bar: 5 mm." captionText-2="FIG. 40. — Palaeovaranus lismonimenos n. sp. Close up photograph of the sculpturing pattern of the holotype parietal NHMW 2019/0047/0001 and the crenula- tion of its dorsolateral crests in dorsal view. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720893" figureDoi-1="http://doi.org/10.5281/zenodo.4720899" figureDoi-2="http://doi.org/10.5281/zenodo.4720901" httpUri-0="https://zenodo.org/record/4720893/files/figure.png" httpUri-1="https://zenodo.org/record/4720899/files/figure.png" httpUri-2="https://zenodo.org/record/4720901/files/figure.png" pageId="36" pageNumber="253">38-40</figureCitation>
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;
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<figureCitation id="1333CEA8FF87FFE534A3FB0CFE923CC3" box="[1181,1228,1204,1230]" captionStart="FIG" captionStartId="46.[132,143,1568,1585]" captionTargetBox="[264,1324,974,1525]" captionTargetId="figure-255@46.[358,1160,968,1533]" captionTargetPageId="46" captionText="FIG. 41. — Palaeovaranus cayluxi Zittel, 1887-1890 and Palaeovaranus lismonimenos n. sp. Transverse sections of 3D models of parietals. A, parietal NHMW 2019/0048/0001 of Palaeovaranus cayluxi; B, holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4720903" httpUri="https://zenodo.org/record/4720903/files/figure.png" pageId="36" pageNumber="253">41B</figureCitation>
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;
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<figureCitation id="1333CEA8FF87FFE534E6FB0CFEAA3CC3" box="[1240,1268,1204,1230]" captionStart="FIG" captionStartId="47.[132,143,1664,1681]" captionTargetBox="[122,1466,175,1637]" captionTargetId="figure-137@47.[122,1466,252,1642]" captionTargetPageId="47" captionText="FIG. 42. — Palaeovaranus lismonimenos n. sp. Parietals of juvenile individuals.A, B, photographs of parietal MNHN.F.QU17177 in dorsal (A) and ventral (B) views; C, D, photographs of parietal UM BFI 1873 in dorsal (C) and ventral (D) views. Both these specimens originally appeared as drawings in Augé (2005: Figs 194 and 195 respectively). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="36" pageNumber="253">42</figureCitation>
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;
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<figureCitation id="1333CEA8FF87FFE5353CFB0CFF403CC3" box="[1282,1310,1204,1230]" captionStart="FIG" captionStartId="48.[132,142,1821,1838]" captionTargetBox="[263,1345,214,1779]" captionTargetId="figure-93@48.[219,1369,201,1796]" captionTargetPageId="48" captionText="FIG. 43. — Palaeovaranus lismonimenos n. sp. frontals: A, B, photographs of frontal UM PRA 8 in dorsal (A) and ventral (B) views; C, D, photographs of frontal MNHN.F.QU17175 in dorsal (C) and ventral (D) views.Both these specimens originally appeared asdrawings inAugé (2005: figs196 and 189 respectively).Scale bars:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720911" httpUri="https://zenodo.org/record/4720911/files/figure.png" pageId="36" pageNumber="253">43</figureCitation>
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)
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</paragraph>
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<paragraph id="8BB7D22DFF87FFE53313FB4EFFF13D0A" blockId="36.[813,1455,1270,1287]" box="[813,1455,1270,1287]" pageId="36" pageNumber="253">
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<uri id="FF99DE2FFF87FFE53313FB4EFFF13D0A" box="[813,1455,1270,1287]" pageId="36" pageNumber="253">
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urn:lsid:zoobank.org:act:
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<uuid id="FFAEE8F8FF87FFE53438FB4EFFF13D0A" box="[1030,1455,1270,1287]" pageId="36" pageNumber="253">3C906259-DAB5-4B97-A1BB-7860695A76B8</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31281A6FF87FFE53313FA85F9BB3D62" pageId="36" pageNumber="253" type="materials_examined">
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<paragraph id="8BB7D22DFF87FFE53313FA85F9BB3D62" blockId="36.[813,1453,1341,1391]" pageId="36" pageNumber="253">
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<materialsCitation id="3B60D870FF87FFE53313FA85F9BB3D62" ID-GBIF-Occurrence="3091185307" collectionCode="NHMW" pageId="36" pageNumber="253" specimenCode="NHMW 2019" specimenCount="1" typeStatus="holotype">
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<typeStatus id="54B36C8FFF87FFE53313FA85F9FA3D5E" box="[813,932,1341,1364]" pageId="36" pageNumber="253" type="holotype">HOLOTYPE</typeStatus>
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. — An almost complete parietal (
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<specimenCode id="DBAE7A56FF87FFE53570FA85F93C3D62" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="36" pageNumber="253" type="Museum">NHMW 2019</specimenCode>
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/0047/0001).
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31281A6FF87FFE53313FA35FF533DF9" pageId="36" pageNumber="253" type="etymology">
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<paragraph id="8BB7D22DFF87FFE53313FA35FF533DF9" blockId="36.[811,1455,1420,1524]" pageId="36" pageNumber="253">
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ETYMOLOGY. — The new name originates from the Greek word “λησμονημένος” (“lismonimenos”) meaning “forgotten”, alluding to the fact that the
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<typeStatus id="54B36C8FFF87FFE533DEFA79FE683DD4" box="[992,1078,1473,1497]" pageId="36" pageNumber="253" type="holotype">holotype</typeStatus>
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specimen was forgotten and unnoticed inside a museum drawer for more than a century.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31281A6FF87FFE53313F9A9FFF03E53" pageId="36" pageNumber="253" type="materials_examined">
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<paragraph id="8BB7D22DFF87FFE53313F9A9FFF03E53" blockId="36.[811,1457,1553,1630]" pageId="36" pageNumber="253">
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<collectionCode id="ED194AE8FF87FFE53313F9A9F99C3E2A" box="[813,962,1553,1576]" pageId="36" pageNumber="253">TYPE LOCALITY</collectionCode>
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. — Imprecisely known locality, Phosphorites du Quercy, Department of Lot or Tarn-et-Garonne, Occitanie, southern
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<collectingCountry id="F31F92BDFF87FFE53315F9FEF9323E50" box="[811,876,1606,1629]" name="France" pageId="36" pageNumber="253">France</collectingCountry>
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; probably late Eocene, around
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<collectionCode id="ED194AE8FF87FFE534BDF9FEFEF93E50" box="[1155,1191,1606,1629]" country="0" name="Mohonk Preserve, Inc." pageId="36" pageNumber="253">MP</collectionCode>
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17 (see Distribution below).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31281A6FF87FFE43313F9C3F95B3F8F" lastPageId="37" lastPageNumber="254" pageId="36" pageNumber="253" type="diagnosis">
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<paragraph id="8BB7D22DFF87FFE53313F9C3FEBB3FE6" blockId="36.[811,1456,1659,2027]" pageId="36" pageNumber="253">
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DIAGNOSIS. — The parietal of
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<taxonomicName id="4C08A9AEFF87FFE5345BF9C3FF2F3E9E" authority="Georgalis & Čerňanský & Klembara, 2021" authorityName="Georgalis & Čerňanský & Klembara" authorityYear="2021" box="[1125,1393,1659,1683]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="253" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
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<emphasis id="B97C0E3FFF87FFE5345BF9C3FF2F3E9E" box="[1125,1393,1659,1683]" italics="true" pageId="36" pageNumber="253">Palaeovaranus lismonimenos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24FB344FF87FFE53546F9C3FFEE3E9F" box="[1400,1456,1659,1682]" pageId="36" pageNumber="253" rank="species">n. sp.</taxonomicNameLabel>
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differs from that of
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<taxonomicName id="4C08A9AEFF87FFE533CCF92DFE9F3EA0" authorityName="Zittel" authorityYear="1887" box="[1010,1217,1685,1709]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="36" pageNumber="253" phylum="Chordata" rank="species" species="cayluxi">
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<emphasis id="B97C0E3FFF87FFE533CCF92DFE9F3EA0" box="[1010,1217,1685,1709]" italics="true" pageId="36" pageNumber="253">Palaeovaranus cayluxi</emphasis>
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</taxonomicName>
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by the following distinguished features and the combination of features: 1) the dorsolateral crests are rather distinct, extending posterolaterally and dorsally, and their margins are distinctly crenulated; 2) the median crest is short (shorter than the length of the median triangular field measured in the median plane) and its posterior tip fits between the anterolateral processes of the triangular median field; 3) the anterior ends of the dorsolateral crests extend to the tips of the anterolateral processes; 4) the ornamentation of the parietal consists of small, more or less densely arranged mounds having more or less distinctly developed crest; and 5) the anterior margin of the parietal fossa lies anterior to the level of the junctions of the anterolateral margins of the supratemporal processes with the parietal plate.
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</paragraph>
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<caption id="DF7782A5FF86FFE430BAF960FB363F0D" ID-DOI="http://doi.org/10.5281/zenodo.4720869" ID-Zenodo-Dep="4720869" httpUri="https://zenodo.org/record/4720869/files/figure.png" pageId="37" pageNumber="254" startId="37.[132,143,1752,1769]" targetBox="[137,1448,216,1711]" targetPageId="37">
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<paragraph id="8BB7D22DFF86FFE430BAF960FB363F0D" blockId="37.[132,1455,1752,1792]" pageId="37" pageNumber="254">
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FIG. 30. —
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<taxonomicName id="4C08A9AEFF86FFE430DEF960FB0A3EE4" authorityName="Sullivan" authorityYear="1979" box="[224,340,1752,1769]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="254" phylum="Chordata" rank="tribe" tribe="Melanosaurini">Melanosaurini</taxonomicName>
|
|
indet. Photographs of presacral vertebra NHMW 2019/0094/0001 in anterior (
|
|
<emphasis id="B97C0E3FFF86FFE433DCF960F9B13EE4" bold="true" box="[994,1007,1752,1769]" pageId="37" pageNumber="254">A</emphasis>
|
|
), posterior (
|
|
<emphasis id="B97C0E3FFF86FFE4346AF960FE3F3EE4" bold="true" box="[1108,1121,1752,1769]" pageId="37" pageNumber="254">B</emphasis>
|
|
), dorsal (
|
|
<emphasis id="B97C0E3FFF86FFE43491F960FEE33EE4" bold="true" box="[1199,1213,1752,1769]" pageId="37" pageNumber="254">C</emphasis>
|
|
), ventral (
|
|
<emphasis id="B97C0E3FFF86FFE43531F960FF433EE4" bold="true" box="[1295,1309,1752,1769]" pageId="37" pageNumber="254">D</emphasis>
|
|
), and right lateral (
|
|
<emphasis id="B97C0E3FFF86FFE430B7F957FACB3F0D" bold="true" box="[137,149,1775,1792]" pageId="37" pageNumber="254">E</emphasis>
|
|
) views. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BB7D22DFF86FFE430BAF88BF95B3F8F" blockId="37.[130,775,1842,1922]" pageId="37" pageNumber="254">
|
|
REFERRED SPECIMENS. —
|
|
<materialsCitation id="3B60D870FF86FFE4314AF88AF95B3F8F" ID-GBIF-Occurrence="3091185305" pageId="37" pageNumber="254">A complete parietal (MNHN.F.QU17177) and a partial parietal (UM BFI 1873), both from juvenile individuals. Tentatively also, two frontals (MNHN.F.QU17175 and UM PRA 8).</materialsCitation>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C31281A6FF86FFE730BAF824FBF63EF3" lastPageId="38" lastPageNumber="255" pageId="37" pageNumber="254" type="distribution">
|
|
<paragraph id="8BB7D22DFF86FFE430BAF824F9593FE6" blockId="37.[131,775,1947,2027]" pageId="37" pageNumber="254">
|
|
DISTRIBUTION. — The
|
|
<typeStatus id="54B36C8FFF86FFE43161F823FBED3FBE" box="[351,435,1947,1971]" pageId="37" pageNumber="254" type="holotype">holotype</typeStatus>
|
|
parietal NHMW 2019/0047/0001, the referred parietal MNHN.F.QU17177, and the tentatively referred frontal MNHN.F.QU17175, all originate from impre-
|
|
</paragraph>
|
|
<paragraph id="8BB7D22DFF86FFE73313F88AFBF63EF3" blockId="37.[811,1457,1842,2027]" lastBlockId="38.[131,775,1713,1790]" lastPageId="38" lastPageNumber="255" pageId="37" pageNumber="254">
|
|
cisely known localities in the Department of Lot or that of Tarnet-Garonne, within the Phosphorites du Quercy. The referred parietal UM BFI 1873 originates from the late Eocene (MP 17) of Bouffie (= La Bouffie), Quercy (Department of Lot), while the tentatively referred frontal UM PRA 8 originates from the coeval, late Eocene (MP 17) locality of Les Pradigues, also in Quercy (but in the Department of Tarn-et-Garonne). Accordingly, we here sug- gest that the specimens with imprecise locality date (including the
|
|
<typeStatus id="54B36C8FFF85FFE730BDF973FA803EEE" box="[131,222,1739,1763]" pageId="38" pageNumber="255" type="holotype">holotype</typeStatus>
|
|
), originate also from late Eocene locality(ies), potentially also around the MP 17 stage.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<caption id="DF7782A5FF85FFE730BAF9F5FEE33E79" ID-DOI="http://doi.org/10.5281/zenodo.4720871" ID-Zenodo-Dep="4720871" httpUri="https://zenodo.org/record/4720871/files/figure.png" pageId="38" pageNumber="255" startId="38.[132,143,1613,1630]" targetBox="[140,1451,215,1571]" targetPageId="38">
|
|
<paragraph id="8BB7D22DFF85FFE730BAF9F5FEE33E79" blockId="38.[132,1455,1613,1652]" pageId="38" pageNumber="255">
|
|
FIG. 31. —
|
|
<taxonomicName id="4C08A9AEFF85FFE730E2F9F5FB0E3E53" authorityName="Sullivan" authorityYear="1979" box="[220,336,1613,1630]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="38" pageNumber="255" phylum="Chordata" rank="tribe" tribe="Melanosaurini">Melanosaurini</taxonomicName>
|
|
indet.
|
|
<emphasis id="B97C0E3FFF85FFE731B8F9F5FBF83E53" bold="true" box="[390,422,1613,1630]" pageId="38" pageNumber="255">A -E</emphasis>
|
|
, photographs of presacral vertebra NHMW 2019/0094/0002 in anterior (
|
|
<emphasis id="B97C0E3FFF85FFE733CDF9F5FE5E3E53" bold="true" box="[1011,1024,1613,1630]" pageId="38" pageNumber="255">A</emphasis>
|
|
), dorsal (
|
|
<emphasis id="B97C0E3FFF85FFE73474F9F5FE093E53" bold="true" box="[1098,1111,1613,1630]" pageId="38" pageNumber="255">B</emphasis>
|
|
), right lateral (
|
|
<emphasis id="B97C0E3FFF85FFE734F7F9F5FE893E53" bold="true" box="[1225,1239,1613,1630]" pageId="38" pageNumber="255">C</emphasis>
|
|
), posterior (
|
|
<emphasis id="B97C0E3FFF85FFE73506F9F5FF183E53" bold="true" box="[1336,1350,1613,1630]" pageId="38" pageNumber="255">D</emphasis>
|
|
), and ventral (
|
|
<emphasis id="B97C0E3FFF85FFE730B7F9DBFACB3E79" bold="true" box="[137,149,1635,1652]" pageId="38" pageNumber="255">E</emphasis>
|
|
) views;
|
|
<emphasis id="B97C0E3FFF85FFE730E7F9DBFABA3E79" bold="true" box="[217,228,1635,1652]" pageId="38" pageNumber="255">F</emphasis>
|
|
,
|
|
<emphasis id="B97C0E3FFF85FFE730D0F9DBFAA23E79" bold="true" box="[238,252,1635,1652]" pageId="38" pageNumber="255">G</emphasis>
|
|
, photographs of presacral vertebra NHMW 2019/0094/0003 in posterior (
|
|
<emphasis id="B97C0E3FFF85FFE7335EF9DBF9353E79" bold="true" box="[864,875,1635,1652]" pageId="38" pageNumber="255">F</emphasis>
|
|
) and ventral (
|
|
<emphasis id="B97C0E3FFF85FFE733E2F9DBF9B43E79" bold="true" box="[988,1002,1635,1652]" pageId="38" pageNumber="255">G</emphasis>
|
|
) views. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<subSubSection id="C31281A6FF85FFE630BAF88AFB113E81" lastPageId="39" lastPageNumber="256" pageId="38" pageNumber="255" type="description">
|
|
<paragraph id="8BB7D22DFF85FFE730BAF88AFB4B3F47" blockId="38.[132,776,1842,2028]" box="[132,277,1842,1868]" pageId="38" pageNumber="255">DESCRIPTION</paragraph>
|
|
<paragraph id="8BB7D22DFF85FFE630BAF8E9F80D3D80" blockId="38.[132,776,1842,2028]" lastBlockId="39.[130,777,1299,1677]" lastPageId="39" lastPageNumber="256" pageId="38" pageNumber="255">
|
|
<emphasis id="B97C0E3FFF85FFE730BAF8E9F9593F61" box="[132,775,1873,1900]" italics="true" pageId="38" pageNumber="255">
|
|
<typeStatus id="54B36C8FFF85FFE730BAF8E9FA823F66" box="[132,220,1873,1899]" pageId="38" pageNumber="255" type="holotype">Holotype</typeStatus>
|
|
<specimenCode id="DBAE7A56FF85FFE730DCF8EAFB283F61" box="[226,374,1874,1900]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="255" type="Museum">NHMW 2019</specimenCode>
|
|
/0047/0001 (
|
|
<figureCitation id="1333CEA8FF85FFE7323FF8EAF8243F66" box="[513,634,1874,1900]" captionStart="FIG" captionStartId="43.[132,143,1820,1837]" captionTargetBox="[124,1472,185,1797]" captionTargetId="figure-78@43.[116,1472,247,1772]" captionTargetPageId="43" captionText="FIG. 37. — Palaeovaranus cayluxi Zittel, 1887-1890 and Palaeovaranus lismonimenos n. sp. A, C, virtual 3D models of parietal NHMW 2019/0048/0001 of Palaeovaranus cayluxi in dorsal (A) and ventral (C) views; B, D, virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in dorsal (B) and ventral (D) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720891" httpUri="https://zenodo.org/record/4720891/files/figure.png" pageId="38" pageNumber="255">Figs 37B, D</figureCitation>
|
|
;
|
|
<figureCitation id="1333CEA8FF85FFE732B9F8EAF8983F61" box="[647,710,1874,1900]" captionStart-0="FIG" captionStart-1="FIG" captionStart-2="FIG" captionStartId-0="44.[132,143,1323,1340]" captionStartId-1="45.[132,143,1318,1335]" captionStartId-2="46.[133,143,881,898]" captionTargetBox-0="[149,1430,217,1279]" captionTargetBox-1="[132,1455,181,1289]" captionTargetBox-2="[360,1236,153,859]" captionTargetId-0="figure-367@44.[145,1442,211,1284]" captionTargetId-1="figure-365@45.[132,1455,228,1276]" captionTargetId-2="figure-212@46.[472,1115,215,839]" captionTargetPageId-0="44" captionTargetPageId-1="45" captionTargetPageId-2="46" captionText-0="FIG. 38. — Palaeovaranus lismonimenos n. sp. Virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in posteroventral (A), dorsolateral (B), and anterodorsal (C) views. Scale bar: 5 mm." captionText-1="FIG. 39. — Palaeovaranus lismonimenos n. sp. Photographs of the holotype parietal NHMW 2019/0047/0001 in dorsal (A), ventral (B), and right lateral (C) views. Scale bar: 5 mm." captionText-2="FIG. 40. — Palaeovaranus lismonimenos n. sp. Close up photograph of the sculpturing pattern of the holotype parietal NHMW 2019/0047/0001 and the crenula- tion of its dorsolateral crests in dorsal view. Scale bar: 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4720893" figureDoi-1="http://doi.org/10.5281/zenodo.4720899" figureDoi-2="http://doi.org/10.5281/zenodo.4720901" httpUri-0="https://zenodo.org/record/4720893/files/figure.png" httpUri-1="https://zenodo.org/record/4720899/files/figure.png" httpUri-2="https://zenodo.org/record/4720901/files/figure.png" pageId="38" pageNumber="255">38-40</figureCitation>
|
|
;
|
|
<figureCitation id="1333CEA8FF85FFE732EDF8EAF8A13F61" box="[723,767,1874,1900]" captionStart="FIG" captionStartId="46.[132,143,1568,1585]" captionTargetBox="[264,1324,974,1525]" captionTargetId="figure-255@46.[358,1160,968,1533]" captionTargetPageId="46" captionText="FIG. 41. — Palaeovaranus cayluxi Zittel, 1887-1890 and Palaeovaranus lismonimenos n. sp. Transverse sections of 3D models of parietals. A, parietal NHMW 2019/0048/0001 of Palaeovaranus cayluxi; B, holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. Scale bar: 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4720903" httpUri="https://zenodo.org/record/4720903/files/figure.png" pageId="38" pageNumber="255">41B</figureCitation>
|
|
)
|
|
</emphasis>
|
|
The
|
|
<typeStatus id="54B36C8FFF85FFE7308BF8C9FB4A3F86" box="[181,276,1905,1931]" pageId="38" pageNumber="255" type="holotype">holotype</typeStatus>
|
|
<specimenCode id="DBAE7A56FF85FFE73122F8CAFBE33F86" box="[284,445,1905,1932]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="38" pageNumber="255" type="Museum">NHMW 2019</specimenCode>
|
|
/0047/0001 has a length of its parietal table
|
|
<quantity id="4CF07FC8FF85FFE73129F829FB253FA1" box="[279,379,1937,1964]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="38" pageNumber="255" unit="mm" value="14.5">14.5 mm</quantity>
|
|
(measured in mid-line). The parietal table is anteroposteriorly elongate (
|
|
<figureCitation id="1333CEA8FF85FFE7322AF809F8253FC1" box="[532,635,1969,1996]" captionStart="FIG" captionStartId="43.[132,143,1820,1837]" captionTargetBox="[124,1472,185,1797]" captionTargetId="figure-78@43.[116,1472,247,1772]" captionTargetPageId="43" captionText="FIG. 37. — Palaeovaranus cayluxi Zittel, 1887-1890 and Palaeovaranus lismonimenos n. sp. A, C, virtual 3D models of parietal NHMW 2019/0048/0001 of Palaeovaranus cayluxi in dorsal (A) and ventral (C) views; B, D, virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in dorsal (B) and ventral (D) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720891" httpUri="https://zenodo.org/record/4720891/files/figure.png" pageId="38" pageNumber="255">Figs 37B</figureCitation>
|
|
;
|
|
<figureCitation id="1333CEA8FF85FFE732B7F809F8E43FC1" box="[649,698,1969,1996]" captionStart="FIG" captionStartId="44.[132,143,1323,1340]" captionTargetBox="[149,1430,217,1279]" captionTargetId="figure-367@44.[145,1442,211,1284]" captionTargetPageId="44" captionText="FIG. 38. — Palaeovaranus lismonimenos n. sp. Virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in posteroventral (A), dorsolateral (B), and anterodorsal (C) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720893" httpUri="https://zenodo.org/record/4720893/files/figure.png" pageId="38" pageNumber="255">38B</figureCitation>
|
|
;
|
|
<figureCitation id="1333CEA8FF85FFE732F6F809F8A43FC6" box="[712,762,1969,1995]" captionStart="FIG" captionStartId="45.[132,143,1318,1335]" captionTargetBox="[132,1455,181,1289]" captionTargetId="figure-365@45.[132,1455,228,1276]" captionTargetPageId="45" captionText="FIG. 39. — Palaeovaranus lismonimenos n. sp. Photographs of the holotype parietal NHMW 2019/0047/0001 in dorsal (A), ventral (B), and right lateral (C) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720899" httpUri="https://zenodo.org/record/4720899/files/figure.png" pageId="38" pageNumber="255">39A</figureCitation>
|
|
). The anterolateral process is slender. The dorsolateral crest is distinctly developed. It extends posterolaterally and dorsally. Its margin is distinctly crenulated. The crenulation consists of several more or less shallow and long notches. The anterior end of the dorsolateral crest reaches the tip of the anterolateral process. The dorsolateral crests and the anterior margin of the parietal limit a triangular field. Its deepest portion is pierced by the parietal foramen. The foramen lies in about the midlength of the anterior half of the parietal table. The surface of the triangular field is covered by small, but distinct mounds, most of them bearing the longitudinal crests on their surfaces. The median crest is short and its posterior pointed portion fits between the two anteriorly pointed anterolateral processes of the triangular median field (
|
|
<figureCitation id="1333CEA8FF84FFE631F5FAEBF8723D60" box="[459,556,1363,1389]" captionStart="FIG" captionStartId="45.[132,143,1318,1335]" captionTargetBox="[132,1455,181,1289]" captionTargetId="figure-365@45.[132,1455,228,1276]" captionTargetPageId="45" captionText="FIG. 39. — Palaeovaranus lismonimenos n. sp. Photographs of the holotype parietal NHMW 2019/0047/0001 in dorsal (A), ventral (B), and right lateral (C) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720899" httpUri="https://zenodo.org/record/4720899/files/figure.png" pageId="39" pageNumber="256">Fig. 39A</figureCitation>
|
|
). The supratemporal fossa is large and is inclined ventrolaterally.
|
|
</paragraph>
|
|
<caption id="DF7782A5FF84FFE630BAFB09FBE33CD4" ID-DOI="http://doi.org/10.5281/zenodo.4720873" ID-Zenodo-Dep="4720873" httpUri="https://zenodo.org/record/4720873/files/figure.png" pageId="39" pageNumber="256" startId="39.[132,143,1201,1218]" targetBox="[132,1455,211,1163]" targetPageId="39">
|
|
<paragraph id="8BB7D22DFF84FFE630BAFB09FBE33CD4" blockId="39.[132,1455,1201,1241]" pageId="39" pageNumber="256">
|
|
FIG. 32. —
|
|
<taxonomicName id="4C08A9AEFF84FFE630E1FB09FB0D3CCF" authorityName="Sullivan" authorityYear="1979" box="[223,339,1201,1218]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="256" phylum="Chordata" rank="tribe" tribe="Melanosaurini">Melanosaurini</taxonomicName>
|
|
indet. Photographs of sacral vertebra NHMW 2019/0094/0004 in anterior (
|
|
<emphasis id="B97C0E3FFF84FFE633FAFB09F98F3CCF" bold="true" box="[964,977,1201,1218]" pageId="39" pageNumber="256">A</emphasis>
|
|
), posterior (
|
|
<emphasis id="B97C0E3FFF84FFE6340BFB09FE1C3CCF" bold="true" box="[1077,1090,1201,1218]" pageId="39" pageNumber="256">B</emphasis>
|
|
), right lateral (
|
|
<emphasis id="B97C0E3FFF84FFE63487FB09FE993CCF" bold="true" box="[1209,1223,1201,1218]" pageId="39" pageNumber="256">C</emphasis>
|
|
), dorsal (
|
|
<emphasis id="B97C0E3FFF84FFE6352AFB09FF7C3CCF" bold="true" box="[1300,1314,1201,1218]" pageId="39" pageNumber="256">D</emphasis>
|
|
), ventral (
|
|
<emphasis id="B97C0E3FFF84FFE6354AFB09FFDE3CCF" bold="true" box="[1396,1408,1201,1218]" pageId="39" pageNumber="256">E</emphasis>
|
|
), and left lateral (
|
|
<emphasis id="B97C0E3FFF84FFE630DEFB70FAB53CD4" bold="true" box="[224,235,1224,1241]" pageId="39" pageNumber="256">F</emphasis>
|
|
) views. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BB7D22DFF84FFE630A5FA2BFB113E81" blockId="39.[130,777,1299,1677]" pageId="39" pageNumber="256">
|
|
The ventral surface is smooth (
|
|
<figureCitation id="1333CEA8FF84FFE631D4FA2BF80E3DA3" box="[490,592,1427,1454]" captionStart="FIG" captionStartId="43.[132,143,1820,1837]" captionTargetBox="[124,1472,185,1797]" captionTargetId="figure-78@43.[116,1472,247,1772]" captionTargetPageId="43" captionText="FIG. 37. — Palaeovaranus cayluxi Zittel, 1887-1890 and Palaeovaranus lismonimenos n. sp. A, C, virtual 3D models of parietal NHMW 2019/0048/0001 of Palaeovaranus cayluxi in dorsal (A) and ventral (C) views; B, D, virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in dorsal (B) and ventral (D) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720891" httpUri="https://zenodo.org/record/4720891/files/figure.png" pageId="39" pageNumber="256">Figs 37D</figureCitation>
|
|
;
|
|
<figureCitation id="1333CEA8FF84FFE63263FA2BF8D33DA0" box="[605,653,1427,1453]" captionStart="FIG" captionStartId="44.[132,143,1323,1340]" captionTargetBox="[149,1430,217,1279]" captionTargetId="figure-367@44.[145,1442,211,1284]" captionTargetPageId="44" captionText="FIG. 38. — Palaeovaranus lismonimenos n. sp. Virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in posteroventral (A), dorsolateral (B), and anterodorsal (C) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720893" httpUri="https://zenodo.org/record/4720893/files/figure.png" pageId="39" pageNumber="256">38A</figureCitation>
|
|
;
|
|
<figureCitation id="1333CEA8FF84FFE632A7FA2BF8973DA3" box="[665,713,1427,1454]" captionStart="FIG" captionStartId="45.[132,143,1318,1335]" captionTargetBox="[132,1455,181,1289]" captionTargetId="figure-365@45.[132,1455,228,1276]" captionTargetPageId="45" captionText="FIG. 39. — Palaeovaranus lismonimenos n. sp. Photographs of the holotype parietal NHMW 2019/0047/0001 in dorsal (A), ventral (B), and right lateral (C) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720899" httpUri="https://zenodo.org/record/4720899/files/figure.png" pageId="39" pageNumber="256">39B</figureCitation>
|
|
). The anterior margin of the parietal fossa lies anterior to the level of the junctions of the anterolateral margins of the supratemporal processes with the parietal plate. The ventral cranial crest is rather low and runs closely medially to the lateral margin of the parietal table. The postfoveal crest passes along the medial margin of the root portion of the supratemporal process (
|
|
<figureCitation id="1333CEA8FF84FFE630DCF9CAFB1D3E81" box="[226,323,1650,1676]" captionStart="FIG" captionStartId="44.[132,143,1323,1340]" captionTargetBox="[149,1430,217,1279]" captionTargetId="figure-367@44.[145,1442,211,1284]" captionTargetPageId="44" captionText="FIG. 38. — Palaeovaranus lismonimenos n. sp. Virtual 3D models of the holotype parietal NHMW 2019/0047/0001 of Palaeovaranus lismonimenos n. sp. in posteroventral (A), dorsolateral (B), and anterodorsal (C) views. Scale bar: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720893" httpUri="https://zenodo.org/record/4720893/files/figure.png" pageId="39" pageNumber="256">Fig. 38A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C31281A6FF84FFE830BAF90BFB843F01" lastPageId="41" lastPageNumber="258" pageId="39" pageNumber="256" type="discussion">
|
|
<paragraph id="8BB7D22DFF84FFE630BAF90BFAB63EC6" blockId="39.[130,777,1715,2028]" box="[132,232,1715,1741]" pageId="39" pageNumber="256">REMARKS</paragraph>
|
|
<paragraph id="8BB7D22DFF84FFE630BAF96AFE483F81" blockId="39.[130,777,1715,2028]" lastBlockId="39.[810,1457,1299,2028]" pageId="39" pageNumber="256">
|
|
The two smaller specimens MNHN.F.QU17177 and UM
|
|
<specimenCode id="DBAE7A56FF84FFE630B8F94AFAB03F01" box="[134,238,1778,1804]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="39" pageNumber="256">BFI 1873</specimenCode>
|
|
(
|
|
<figureCitation id="1333CEA8FF84FFE630C5F94AFB183F01" box="[251,326,1778,1804]" captionStart="FIG" captionStartId="47.[132,143,1664,1681]" captionTargetBox="[122,1466,175,1637]" captionTargetId="figure-137@47.[122,1466,252,1642]" captionTargetPageId="47" captionText="FIG. 42. — Palaeovaranus lismonimenos n. sp. Parietals of juvenile individuals.A, B, photographs of parietal MNHN.F.QU17177 in dorsal (A) and ventral (B) views; C, D, photographs of parietal UM BFI 1873 in dorsal (C) and ventral (D) views. Both these specimens originally appeared as drawings in Augé (2005: Figs 194 and 195 respectively). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="39" pageNumber="256">Fig. 42</figureCitation>
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|
), which we herein assign to
|
|
<taxonomicName id="4C08A9AEFF84FFE6324DF94AFB503F21" authority="Georgalis & Čerňanský & Klembara, 2021" authorityName="Georgalis & Čerňanský & Klembara" authorityYear="2021" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="256" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
|
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<emphasis id="B97C0E3FFF84FFE6324DF94AFB503F21" italics="true" pageId="39" pageNumber="256">Palaeovaranus lismonimenos</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A24FB344FF84FFE6312AF8AAFB103F21" box="[276,334,1810,1836]" pageId="39" pageNumber="256" rank="species">n. sp.</taxonomicNameLabel>
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were originally described by
|
|
<bibRefCitation id="EF99AFDCFF84FFE63241F8AAF9593F21" author="AUGE M." box="[639,775,1809,1836]" pageId="39" pageNumber="256" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé (2005)</bibRefCitation>
|
|
who also provided drawings of both (his Figs 194 and 195 respectively) and referred them to as “
|
|
<taxonomicName id="4C08A9AEFF84FFE6321EF8EAFAFC3F86" authorityName="" (Auge" authorityYear="2005" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="39" pageNumber="256" phylum="Chordata" rank="species" species="eucarinatus">
|
|
<emphasis id="B97C0E3FFF84FFE6321EF8EAFAFC3F86" italics="true" pageId="39" pageNumber="256">Necrosaurus eucarinatus</emphasis>
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</taxonomicName>
|
|
” (see Discussion below about the status of this name). MNHN.F.QU17177 has a length of the parietal table
|
|
<quantity id="4CF07FC8FF84FFE632E6F829FAEE3FC1" metricMagnitude="-2" metricUnit="m" metricValue="1.15" pageId="39" pageNumber="256" unit="mm" value="11.5">11.5 mm</quantity>
|
|
(measured in the mid-line) (
|
|
<figureCitation id="1333CEA8FF84FFE631EAF809F8143FC1" box="[468,586,1969,1996]" captionStart="FIG" captionStartId="47.[132,143,1664,1681]" captionTargetBox="[122,1466,175,1637]" captionTargetId="figure-137@47.[122,1466,252,1642]" captionTargetPageId="47" captionText="FIG. 42. — Palaeovaranus lismonimenos n. sp. Parietals of juvenile individuals.A, B, photographs of parietal MNHN.F.QU17177 in dorsal (A) and ventral (B) views; C, D, photographs of parietal UM BFI 1873 in dorsal (C) and ventral (D) views. Both these specimens originally appeared as drawings in Augé (2005: Figs 194 and 195 respectively). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="39" pageNumber="256">Fig. 42A, B</figureCitation>
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); thus, it is smaller than the
|
|
<typeStatus id="54B36C8FFF84FFE630DAF869FB1D3FE6" box="[228,323,2001,2027]" pageId="39" pageNumber="256" type="holotype">holotype</typeStatus>
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parietal
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<specimenCode id="DBAE7A56FF84FFE6319FF86AF81C3FE6" box="[417,578,2001,2028]" collectionCode="NHMW" country="Austria" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="39" pageNumber="256" type="Museum">NHMW 2019</specimenCode>
|
|
/0047/0001 (
|
|
<quantity id="4CF07FC8FF84FFE632EDF869F9033D23" box="[723,861,1300,2028]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="39" pageNumber="256" unit="mm" value="14.5">14.5 mm</quantity>
|
|
). The right supratemporal process is completely preserved in MNHN.F.QU17177; it extends posterolaterally (
|
|
<figureCitation id="1333CEA8FF84FFE63570FA8BF91E3D63" captionStart="FIG" captionStartId="47.[132,143,1664,1681]" captionTargetBox="[122,1466,175,1637]" captionTargetId="figure-137@47.[122,1466,252,1642]" captionTargetPageId="47" captionText="FIG. 42. — Palaeovaranus lismonimenos n. sp. Parietals of juvenile individuals.A, B, photographs of parietal MNHN.F.QU17177 in dorsal (A) and ventral (B) views; C, D, photographs of parietal UM BFI 1873 in dorsal (C) and ventral (D) views. Both these specimens originally appeared as drawings in Augé (2005: Figs 194 and 195 respectively). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="39" pageNumber="256">Fig. 42A, B</figureCitation>
|
|
). The dorsolateral crests of the specimen MNHN.F.QU17177 bear slightly developed crenulations and do not meet in the mid-line, as is also the case in the still smaller specimen UM
|
|
<specimenCode id="DBAE7A56FF84FFE63312FA0BF9CA3DC0" box="[812,916,1459,1485]" collectionCode="UM" country="Germany" name="University of Marburg" pageId="39" pageNumber="256">BFI 1873</specimenCode>
|
|
(
|
|
<figureCitation id="1333CEA8FF84FFE6339AFA0BFE773DC0" box="[932,1065,1459,1485]" captionStart="FIG" captionStartId="47.[132,143,1664,1681]" captionTargetBox="[122,1466,175,1637]" captionTargetId="figure-137@47.[122,1466,252,1642]" captionTargetPageId="47" captionText="FIG. 42. — Palaeovaranus lismonimenos n. sp. Parietals of juvenile individuals.A, B, photographs of parietal MNHN.F.QU17177 in dorsal (A) and ventral (B) views; C, D, photographs of parietal UM BFI 1873 in dorsal (C) and ventral (D) views. Both these specimens originally appeared as drawings in Augé (2005: Figs 194 and 195 respectively). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="39" pageNumber="256">Fig. 42C, D</figureCitation>
|
|
). In this smallest specimen UM
|
|
<specimenCode id="DBAE7A56FF84FFE635B8FA0BF9393DE0" collectionCode="UM" country="Germany" name="University of Marburg" pageId="39" pageNumber="256">BFI 1873</specimenCode>
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, the ornamentation is weakly developed and the dorsolateral crests are still more distantly placed one to another than in MNHN.F.QU17177. We may interpret this here by the hypothesis that during ontogenetic growth, the dorsolateral crests move one to another and finally fuse together in about their posterior portions to form a median crest. As a consequence, large anterior and small posterior median triangular fields are produced (
|
|
<figureCitation id="1333CEA8FF84FFE634BEF90AFE933EC1" box="[1152,1229,1714,1740]" captionStart="FIG" captionStartId="47.[132,143,1664,1681]" captionTargetBox="[122,1466,175,1637]" captionTargetId="figure-137@47.[122,1466,252,1642]" captionTargetPageId="47" captionText="FIG. 42. — Palaeovaranus lismonimenos n. sp. Parietals of juvenile individuals.A, B, photographs of parietal MNHN.F.QU17177 in dorsal (A) and ventral (B) views; C, D, photographs of parietal UM BFI 1873 in dorsal (C) and ventral (D) views. Both these specimens originally appeared as drawings in Augé (2005: Figs 194 and 195 respectively). Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720907" httpUri="https://zenodo.org/record/4720907/files/figure.png" pageId="39" pageNumber="256">Fig. 42</figureCitation>
|
|
). If so, the anatomy of three different size stages presented and discussed herein represent the first evidence of the medial movement of the dorsolateral crests to their final fusion in the median plane in adult specimens. We suppose the same process of the origin of the similar morphology of the dorsal surface of parietal in
|
|
<taxonomicName id="4C08A9AEFF84FFE63312F8C9FE503F86" authorityName="Zittel" authorityYear="1887" box="[812,1038,1905,1931]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="39" pageNumber="256" phylum="Chordata" rank="species" species="cayluxi">
|
|
<emphasis id="B97C0E3FFF84FFE63312F8C9FE503F86" box="[812,1038,1905,1931]" italics="true" pageId="39" pageNumber="256">Palaeovaranus cayluxi</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="8BB7D22DFF84FFE6337DF829FFF13FE1" blockId="39.[810,1457,1299,2028]" pageId="39" pageNumber="256">
|
|
Although there is no palaeovaranid frontal material in the NHMW collection, there have been previously described such elements from the Phosphorites du Quercy (
|
|
<figureCitation id="1333CEA8FF84FFE6356CF869FFC13FE6" box="[1362,1439,2001,2027]" captionStart="FIG" captionStartId="48.[132,142,1821,1838]" captionTargetBox="[263,1345,214,1779]" captionTargetId="figure-93@48.[219,1369,201,1796]" captionTargetPageId="48" captionText="FIG. 43. — Palaeovaranus lismonimenos n. sp. frontals: A, B, photographs of frontal UM PRA 8 in dorsal (A) and ventral (B) views; C, D, photographs of frontal MNHN.F.QU17175 in dorsal (C) and ventral (D) views.Both these specimens originally appeared asdrawings inAugé (2005: figs196 and 189 respectively).Scale bars:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720911" httpUri="https://zenodo.org/record/4720911/files/figure.png" pageId="39" pageNumber="256">Fig. 43</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="DF7782A5FF8BFFE930BAF812FB133FE4" ID-DOI="http://doi.org/10.5281/zenodo.4720875" ID-Zenodo-Dep="4720875" httpUri="https://zenodo.org/record/4720875/files/figure.png" pageId="40" pageNumber="257" startId="40.[132,143,1962,1979]" targetBox="[132,1455,207,1870]" targetPageId="40">
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|
<paragraph id="8BB7D22DFF8BFFE930BAF812FB133FE4" blockId="40.[132,1456,1962,2025]" pageId="40" pageNumber="257">
|
|
FIG. 33. —
|
|
<taxonomicName id="4C08A9AEFF8BFFE930DEF812FB703FB6" authorityName="Gray" authorityYear="1825" box="[224,302,1962,1979]" class="Reptilia" family="Anguidae" kingdom="Animalia" order="Squamata" pageId="40" pageNumber="257" phylum="Chordata" rank="subFamily" subFamily="Anguinae">Anguinae</taxonomicName>
|
|
indet.
|
|
<emphasis id="B97C0E3FFF8BFFE93156F812FBD93FB6" bold="true" box="[360,391,1962,1979]" pageId="40" pageNumber="257">A -F</emphasis>
|
|
, photographs of presacral vertebra NHMW 2019/0093/0001 in anterior (
|
|
<emphasis id="B97C0E3FFF8BFFE933D8F812F9AD3FB6" bold="true" box="[998,1011,1962,1979]" pageId="40" pageNumber="257">A</emphasis>
|
|
), posterior (
|
|
<emphasis id="B97C0E3FFF8BFFE93466F812FE3B3FB6" bold="true" box="[1112,1125,1962,1979]" pageId="40" pageNumber="257">B</emphasis>
|
|
), right lateral (
|
|
<emphasis id="B97C0E3FFF8BFFE934E3F812FEB53FB6" bold="true" box="[1245,1259,1962,1979]" pageId="40" pageNumber="257">C</emphasis>
|
|
), dorsal (
|
|
<emphasis id="B97C0E3FFF8BFFE93507F812FF193FB6" bold="true" box="[1337,1351,1962,1979]" pageId="40" pageNumber="257">D</emphasis>
|
|
), ventral (
|
|
<emphasis id="B97C0E3FFF8BFFE935A7F812FFFB3FB6" bold="true" box="[1433,1445,1962,1979]" pageId="40" pageNumber="257">E</emphasis>
|
|
), and left lateral (
|
|
<emphasis id="B97C0E3FFF8BFFE93138F879FB4F3FDF" bold="true" box="[262,273,1985,2002]" pageId="40" pageNumber="257">F</emphasis>
|
|
) views;
|
|
<emphasis id="B97C0E3FFF8BFFE93168F879FB273FDF" bold="true" box="[342,377,1985,2002]" pageId="40" pageNumber="257">G -K</emphasis>
|
|
, photographs of presacral vertebra NHMW 2019/0093/0002 in anterior (
|
|
<emphasis id="B97C0E3FFF8BFFE933E7F879F9B93FDF" bold="true" box="[985,999,1985,2002]" pageId="40" pageNumber="257">G</emphasis>
|
|
), posterior (
|
|
<emphasis id="B97C0E3FFF8BFFE93472F879FE043FDF" bold="true" box="[1100,1114,1985,2002]" pageId="40" pageNumber="257">H</emphasis>
|
|
), left lateral (
|
|
<emphasis id="B97C0E3FFF8BFFE934F9F879FE933FDF" bold="true" box="[1223,1229,1985,2002]" pageId="40" pageNumber="257">I</emphasis>
|
|
), dorsal (
|
|
<emphasis id="B97C0E3FFF8BFFE93524F879FF7A3FDF" bold="true" box="[1306,1316,1985,2002]" pageId="40" pageNumber="257">J</emphasis>
|
|
), and ventral (
|
|
<emphasis id="B97C0E3FFF8BFFE935A3F879FFF43FDF" bold="true" box="[1437,1450,1985,2002]" pageId="40" pageNumber="257">K</emphasis>
|
|
) views. Scale bar: 2 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<caption id="DF7782A5FF8AFFE830BAFA86FFEF3D42" ID-DOI="http://doi.org/10.5281/zenodo.4720877" ID-Zenodo-Dep="4720877" box="[132,1457,1342,1359]" httpUri="https://zenodo.org/record/4720877/files/figure.png" pageId="41" pageNumber="258" startId="41.[132,143,1342,1359]" targetBox="[132,1455,182,1310]" targetPageId="41">
|
|
<paragraph id="8BB7D22DFF8AFFE830BAFA86FFEF3D42" blockId="41.[132,1457,1342,1359]" box="[132,1457,1342,1359]" pageId="41" pageNumber="258">
|
|
FIG. 34. —
|
|
<taxonomicName id="4C08A9AEFF8AFFE830E5FA86FBB83D42" authority="Zittel, 1887" authorityName="Zittel" authorityYear="1887" box="[219,486,1342,1359]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="cayluxi">
|
|
<emphasis id="B97C0E3FFF8AFFE830E5FA86FBD73D42" box="[219,393,1342,1359]" italics="true" pageId="41" pageNumber="258">Palaeovaranus cayluxi</emphasis>
|
|
<bibRefCitation id="EF99AFDCFF8AFFE831B5FA86FBB83D42" author="ZITTEL K. A." box="[395,486,1342,1359]" pageId="41" pageNumber="258" refId="ref40223" refString="ZITTEL K. A. 1887 - 1890. - Handbuch der Palaontologie. Palaeozoologie. III. Pisces, Amphibia, Reptilia, Aves. Druck und Verlag von R. Oldenbourg, Munchen, Leipzig, 900 p. https: // www. biodiversitylibrary. org / page / 40393265" type="book" year="1887">Zittel,1887</bibRefCitation>
|
|
</taxonomicName>
|
|
-1890.Photographs of parietal NHMW 2019/0048/0001 in dorsal (
|
|
<emphasis id="B97C0E3FFF8AFFE833D5FA86F9A93D42" bold="true" box="[1003,1015,1342,1359]" pageId="41" pageNumber="258">A</emphasis>
|
|
), ventral (
|
|
<emphasis id="B97C0E3FFF8AFFE8347DFA86FE0E3D42" bold="true" box="[1091,1104,1342,1359]" pageId="41" pageNumber="258">B</emphasis>
|
|
), and right lateral (
|
|
<emphasis id="B97C0E3FFF8AFFE834E1FA86FEB23D42" bold="true" box="[1247,1260,1342,1359]" pageId="41" pageNumber="258">C</emphasis>
|
|
) views.Scale bar:5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="8BB7D22DFF8AFFE830BAFA2BFB843F01" blockId="41.[130,775,1427,1804]" pageId="41" pageNumber="258">
|
|
<bibRefCitation id="EF99AFDCFF8AFFE830BAFA2BFB503DA3" author="AUGE M." box="[132,270,1427,1454]" pageId="41" pageNumber="258" refId="ref32060" refString="AUGE M. 2005. - Evolution des lezards du Paleogene en Europe. Museum national d'Histoire naturelle, Paris: 1 - 369 (Memoires du Museum national d'Histoire naturelle; 192)." type="journal volume" year="2005">Augé (2005)</bibRefCitation>
|
|
described the frontal MNHN.F.QU17175 as belonging to “
|
|
<taxonomicName id="4C08A9AEFF8AFFE83121FA0CFBB43DC0" authorityName="Filhol" authorityYear="1873" box="[287,490,1459,1485]" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="cayluxi">
|
|
<emphasis id="B97C0E3FFF8AFFE83121FA0CFBB43DC0" box="[287,490,1459,1485]" italics="true" pageId="41" pageNumber="258">Necrosaurus cayluxi</emphasis>
|
|
</taxonomicName>
|
|
” and that of the specimen UM PRA 8 as “
|
|
<taxonomicName id="4C08A9AEFF8AFFE8310EFA6CF8773DE0" authorityName="" (Auge" authorityYear="2005" box="[304,553,1492,1517]" class="Reptilia" family="Necrosauridae" genus="Necrosaurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Squamata" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="eucarinatus">
|
|
<emphasis id="B97C0E3FFF8AFFE8310EFA6CF8773DE0" box="[304,553,1492,1517]" italics="true" pageId="41" pageNumber="258">Necrosaurus eucarinatus</emphasis>
|
|
</taxonomicName>
|
|
”. In contrast to
|
|
<taxonomicName id="4C08A9AEFF8AFFE832E3FA6BFB1F3E00" authorityName="Zittel" authorityYear="1887" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="cayluxi">
|
|
<emphasis id="B97C0E3FFF8AFFE832E3FA6BFB1F3E00" italics="true" pageId="41" pageNumber="258">Palaeovaranus cayluxi</emphasis>
|
|
</taxonomicName>
|
|
, however, both these frontals have the same
|
|
<typeStatus id="54B36C8FFF8AFFE830BCF9ABFAEE3E20" box="[130,176,1555,1581]" pageId="41" pageNumber="258">type</typeStatus>
|
|
of mounds as those present in the dorsal surface of the parietals of
|
|
<taxonomicName id="4C08A9AEFF8AFFE83139F98AF8713E41" authority="Georgalis & Čerňanský & Klembara, 2021" authorityName="Georgalis & Čerňanský & Klembara" authorityYear="2021" box="[263,559,1586,1612]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
|
|
<emphasis id="B97C0E3FFF8AFFE83139F98AF8713E41" box="[263,559,1586,1612]" italics="true" pageId="41" pageNumber="258">Palaeovaranus lismonimenos</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A24FB344FF8AFFE83208F98BF82D3E40" box="[566,627,1587,1613]" pageId="41" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
|
|
(
|
|
<figureCitation id="1333CEA8FF8AFFE832BAF98AF88C3E40" box="[644,722,1586,1613]" captionStart="FIG" captionStartId="48.[132,142,1821,1838]" captionTargetBox="[263,1345,214,1779]" captionTargetId="figure-93@48.[219,1369,201,1796]" captionTargetPageId="48" captionText="FIG. 43. — Palaeovaranus lismonimenos n. sp. frontals: A, B, photographs of frontal UM PRA 8 in dorsal (A) and ventral (B) views; C, D, photographs of frontal MNHN.F.QU17175 in dorsal (C) and ventral (D) views.Both these specimens originally appeared asdrawings inAugé (2005: figs196 and 189 respectively).Scale bars:5 mm." figureDoi="http://doi.org/10.5281/zenodo.4720911" httpUri="https://zenodo.org/record/4720911/files/figure.png" pageId="41" pageNumber="258">Fig. 43</figureCitation>
|
|
). To the contrary, the dorsal surfaces of all three known parietals of
|
|
<taxonomicName id="4C08A9AEFF8AFFE830A1F9CAFB213E81" authorityName="Zittel" authorityYear="1887" box="[159,383,1650,1676]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="cayluxi">
|
|
<emphasis id="B97C0E3FFF8AFFE830A1F9CAFB213E81" box="[159,383,1650,1676]" italics="true" pageId="41" pageNumber="258">Palaeovaranus cayluxi</emphasis>
|
|
</taxonomicName>
|
|
have no such
|
|
<typeStatus id="54B36C8FFF8AFFE8322FF9CBF8613E80" box="[529,575,1651,1677]" pageId="41" pageNumber="258">type</typeStatus>
|
|
of ornamentation, as that exhibited in
|
|
<taxonomicName id="4C08A9AEFF8AFFE83162F92AF8DF3EA1" authority="Georgalis & Čerňanský & Klembara, 2021" authorityName="Georgalis & Čerňanský & Klembara" authorityYear="2021" box="[348,641,1682,1708]" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
|
|
<emphasis id="B97C0E3FFF8AFFE83162F92AF8DF3EA1" box="[348,641,1682,1708]" italics="true" pageId="41" pageNumber="258">Palaeovaranus lismonimenos</emphasis>
|
|
</taxonomicName>
|
|
<taxonomicNameLabel id="A24FB344FF8AFFE832B7F92BF89A3EA0" box="[649,708,1683,1709]" pageId="41" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
|
|
Thus, it is highly probable that these two frontals belong to
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<taxonomicName id="4C08A9AEFF8AFFE83284F90AFB363EE1" authority="Georgalis & Čerňanský & Klembara, 2021" authorityName="Georgalis & Čerňanský & Klembara" authorityYear="2021" class="Reptilia" genus="Palaeovaranus" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="41" pageNumber="258" phylum="Chordata" rank="species" species="lismonimenos" status="sp. nov.">
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<emphasis id="B97C0E3FFF8AFFE83284F90AFB363EE1" italics="true" pageId="41" pageNumber="258">Palaeovaranus lismonimenos</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24FB344FF8AFFE8314FF96AFBF03EE1" box="[369,430,1746,1772]" pageId="41" pageNumber="258" rank="species">n. sp.</taxonomicNameLabel>
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and we accordingly, tentatively assign them to our new species.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |