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<document id="EDC5A1F490CD0BF1BF3564C755C22BF8" ID-DOI="10.11646/phytotaxa.372.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13728728" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725682108736" checkinUser="felipe" docAuthor="Do, Hai Van, Nguyen, Mai Thi, Nguyen, Huong Thi Thanh, Choudhary, Ritesh Kumar &amp; Deng, Yunfei" docDate="2018" docId="C10F30128070590CFF6857739E5F21D8" docLanguage="en" docName="phytotaxa.372.2.3.pdf" docOrigin="Phytotaxa 372 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.372.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Phlogacanthus paniculatus J. B. Imlay 1939" docType="treatment" docVersion="2" lastPageNumber="168" masterDocId="3D36486A8071590DFFE054479D1D2753" masterDocTitle="An updated taxonomy of the genus Phlogacanthus (Acanthaceae) in Vietnam" masterLastPageNumber="175" masterPageNumber="167" pageNumber="168" updateTime="1725693357352" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="D9D806C5306D43BFA3D1F40925FE65A1">An updated taxonomy of the genus Phlogacanthus (Acanthaceae) in Vietnam</mods:title>
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<mods:namePart id="87FCC645B3C7E58F3E39B8938D839FC2">Do, Hai Van</mods:namePart>
<mods:affiliation id="CACFEC57243EB02395E272DB39ED1134">Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="91835F6AD1F681E6755D5154CEEDAEA0">Nguyen, Mai Thi</mods:namePart>
<mods:affiliation id="868DA8400C5ABAA2927865D5A70E47AB">Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="2BBD2D0202520F44ACC0E5FEB16AAB6B">Nguyen, Huong Thi Thanh</mods:namePart>
<mods:affiliation id="3B0477E466572BD0F9F5DA7FE1CFA697">Department of Botany, Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="F3E2D81E5B9694600CEA0CF4BE6FFC3D">Choudhary, Ritesh Kumar</mods:namePart>
<mods:affiliation id="DA0361869B874A0EFE1CF0654D049454">Biodiversity &amp; Palaeobiology Group, Agharkar Research Institute, G. G. Agarkar Road, Pune- 411004, India</mods:affiliation>
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<mods:namePart id="D9CDC92E27B6949A2E7B403C3855A866">Deng, Yunfei</mods:namePart>
<mods:affiliation id="C89E95865CC071628FF56D0A18A09F3C">Key Laboratory of Plant Resources Conservation and Sustainable Utilization, South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, China.</mods:affiliation>
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<mods:title id="180B341DFDC4A63198AA2FBA4736B803">Phytotaxa</mods:title>
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<mods:date id="363C191A21023086923FDC528030D221">2018</mods:date>
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<treatmentCitationGroup id="69B6A62A8070590CFF6857739ECC241C" box="[136,977,820,847]" pageId="1" pageNumber="168">
<treatmentCitation id="C807A7158070590CFF6857739E2E241C" author="Imlay, J. B." box="[136,819,820,847]" page=")" pageId="1" pageNumber="168" year="1939">
<taxonomicName id="8EA6FA878070590CFF6857739E2E241C" ID-CoL="4GHF9" authority="(T. Anderson) Imlay (1939: 128)" authorityName="J. B. Imlay" authorityPageNumber="128" authorityYear="1939" baseAuthorityName="T. Anderson" baseAuthorityYear="1867" box="[136,819,820,847]" class="Magnoliopsida" family="Acanthaceae" genus="Phlogacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="paniculatus">
<emphasis id="7BD25D168070590CFF6857739CA2241C" bold="true" box="[136,447,820,847]" italics="true" pageId="1" pageNumber="168">Phlogacanthus paniculatus</emphasis>
(T. Anderson)
<bibRefCitation id="2D37FCF58070590CFD8557719E2E241C" author="Imlay, J. B." box="[613,819,822,847]" pageId="1" pageNumber="168" pagination="109 - 150" refId="ref4223" refString="Imlay, J. B. (1939) Contributions to the flora of Siam, Additamentum LI, new and renamed Siamense Acanthaceae. Bulletin of Miscellaneous Information Kew 3: 109 - 150. https: // doi. org / 10.2307 / 4111677" type="journal article" year="1939">Imlay (1939: 128)</bibRefCitation>
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.
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(
<figureCitation id="D19D9D818070590CFCA557719E89241C" box="[837,916,822,847]" captionStart="FIGURE 1" captionStartId="2.[136,229,1569,1590]" captionTargetBox="[168,1426,186,1524]" captionTargetId="figure-15@2.[158,1430,171,1541]" captionTargetPageId="2" captionText="FIGURE 1. Phlogacanthus paniculatus (T. Anderson) J.B. Imlay: A. Flowering and fruiting twig; B. Calyx and bracteoles; C. Corolla (open); D. Ovary and style; E. Fruit (Open) (Illustration drawn by Mrs Le Kim Chi, from D.V. Hai 14, HN)." figureDoi="http://doi.org/10.5281/zenodo.13728730" httpUri="https://zenodo.org/record/13728730/files/figure.png" pageId="1" pageNumber="168">Figs. 1</figureCitation>
&amp;
<figureCitation id="D19D9D818070590CFC5A57719ED4241C" box="[954,969,822,847]" captionStart="FIGURE 2" captionStartId="3.[136,229,1530,1551]" captionTargetBox="[151,1435,190,1505]" captionTargetId="figure-14@3.[151,1436,190,1505]" captionTargetPageId="3" captionText="FIGURE 2. Phlogacanthus paniculatus (T. Anderson) J.B. Imlay: A. Flowering and fruiting twig; B. Flower (front view); C. Calyx and flower; D. Calyx and fruit (Photos by D.V. Hai)." figureDoi="http://doi.org/10.5281/zenodo.13728732" httpUri="https://zenodo.org/record/13728732/files/figure.png" pageId="1" pageNumber="168">2</figureCitation>
)
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<treatmentCitationGroup id="69B6A62A8070590CFF68571B9F962422" box="[136,651,860,881]" pageId="1" pageNumber="168">
=
<treatmentCitation id="C807A7158070590CFF7C571B9F9A2422" author="Anderson, T." box="[156,647,860,881]" page=")" pageId="1" pageNumber="168" year="1867">
<taxonomicName id="8EA6FA878070590CFF7C571B9F9A2422" ID-CoL="33P7W" authority="T. Anderson (1867: 458)" authorityName="T. Anderson" authorityPageNumber="458" authorityYear="1867" box="[156,647,860,881]" class="Magnoliopsida" family="Acanthaceae" genus="Cystacanthus" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="168" phylum="Tracheophyta" rank="species" species="paniculatus">
<emphasis id="7BD25D168070590CFF7C571B9C8E2422" box="[156,403,860,881]" italics="true" pageId="1" pageNumber="168">Cystacanthus paniculatus</emphasis>
T.
<bibRefCitation id="2D37FCF58070590CFE50571B9F9A2422" author="Anderson, T." box="[432,647,860,881]" pageId="1" pageNumber="168" pagination="425 - 526" refId="ref3754" refString="Anderson, T. (1867) An enumeration of the Indian species of Acanthaceae. Journal of the Linnean Society, Botany 9: 425 - 526." type="journal article" year="1867">Anderson (1867: 458)</bibRefCitation>
</taxonomicName>
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.
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<emphasis id="7BD25D168070590CFF6857C79DC724C6" bold="true" box="[136,218,896,917]" pageId="1" pageNumber="168">
<typeStatus id="961D3FA68070590CFF6857C79DA024C6" box="[136,189,896,917]" pageId="1" pageNumber="168">Type</typeStatus>
:—
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. Moulmein,
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.Lobb s. n
</emphasis>
. (
<collectionCode id="2FB719C18070590CFDB157C79F9824C6" box="[593,645,896,917]" country="India" httpUri="http://biocol.org/urn:lsid:biocol.org:col:12692" lsid="urn:lsid:biocol.org:col:12692" name="Botanical Survey of India" pageId="1" pageNumber="168" type="Herbarium">CAL</collectionCode>
,
<collectionCode id="2FB719C18070590CFD6E57C79FB824C6" box="[654,677,896,917]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="168" type="Herbarium">K</collectionCode>
!).
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<paragraph id="491981048070590CFF68578198BB2238" blockId="1.[136,1452,966,1675]" pageId="1" pageNumber="168">
Shrubs to
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tall. Stems 4-angled, stout, glabrous; young branches winged. Petiole
<quantity id="8E5E2CE18070590CFBAE578199A6248C" box="[1102,1211,966,991]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="1" pageNumber="168" unit="cm" value="1.35" valueMax="1.7" valueMin="1.0">11.7 cm</quantity>
; leaf blade oblong to oblong-lanceolate, 612 ×
<quantity id="8E5E2CE18070590CFE5957AD9F262350" box="[441,571,1002,1027]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.5" metricValueMin="1.5" pageId="1" pageNumber="168" unit="cm" value="3.0" valueMax="4.5" valueMin="1.5">1.54.5 cm</quantity>
, abaxially pubescent along veins, adaxially glabrous, secondary veins 68 on each side of midvein, base attenuate and decurrent onto petiole, margin undulate to crenulate, apex acuminate. Thyrses terminal, to
<quantity id="8E5E2CE18070590CFEF750759C7E2318" box="[279,355,1074,1099]" metricMagnitude="-1" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="168" unit="cm" value="13.0">13 cm</quantity>
, many-flowered; rachis densely villous with gland-tipped trichomes; bracts oblonglanceolate, shorter than calyx, gland-tipped pubescent, caducous; bracteoles similar to bracts. Pedicel
<quantity id="8E5E2CE18070590CFB4B50119813233C" box="[1195,1294,1110,1135]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="1" pageNumber="168" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
, gland-tipped pubescent. Calyx
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, outside gland-tipped pubescent, inside slightly pubescent; lobes lanceolate, 713 ×
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, equal, apex acute. Corolla white or light purple,
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; bent to 90º and abruptly inflated at middle, ventricose, outside gland-tipped pubescent, inside glabrous with purple stripes; limb 2-lipped; lower lip 3-lobed with lobes orbicular, subequal, patent; upper lip 2-lobed with lobes orbicular, equal. Stamens two, included; filaments barbate; anther thecae oblong, ca.
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, connective barbate; staminodes very small. Ovary pubescent, with 4 ovules per locule; style
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, glabrous or pubescent; stigma slightly 2-cleft or entire. Capsule greyish brown, clavate, ca.
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, pubescent, 8-seeded. Seeds ovate in outline, ca. 3.5 ×
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; densely shortly hairy; hairs standing out when wetted.
</paragraph>
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<emphasis id="7BD25D168070590CFF5D51319CC222DC" bold="true" box="[189,479,1398,1423]" pageId="1" pageNumber="168">Flowering and Fruiting:</emphasis>
June to October.
</paragraph>
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<emphasis id="7BD25D168070590CFF5D51DD9CA722E0" bold="true" box="[189,442,1434,1459]" pageId="1" pageNumber="168">Ecology and habitat:</emphasis>
The species usually grows in the understorey of primary broad-leaved evergreen closed forest, in humid areas.
</paragraph>
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<emphasis id="7BD25D168070590CFF5D51A59C4422A8" bold="true" box="[189,345,1506,1531]" pageId="1" pageNumber="168">Distribution:</emphasis>
<collectingCountry id="31B1C1948070590CFE8551A59CD922A8" box="[357,452,1506,1531]" name="Vietnam" pageId="1" pageNumber="168">Vietnam</collectingCountry>
(
<collectingRegion id="8B624FE68070590CFE3751A59FC522A8" box="[471,728,1506,1531]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="168">Quang Binh Province</collectingRegion>
,
<collectingRegion id="8B624FE68070590CFD0951A59EC622A8" box="[745,987,1506,1531]" country="Vietnam" name="Quang Ninh" pageId="1" pageNumber="168">Quang Ninh District</collectingRegion>
, Ba Ren farm forest);
<collectingCountry id="31B1C1948070590CFB0D51A5982D22A8" box="[1261,1328,1506,1531]" name="China" pageId="1" pageNumber="168">China</collectingCountry>
(
<collectingRegion id="8B624FE68070590CFAA451A598BD22A8" box="[1348,1440,1506,1531]" country="China" name="Yunnan" pageId="1" pageNumber="168">Yunnan</collectingRegion>
),
<collectingCountry id="31B1C1948070590CFF6852419DEA214C" box="[136,247,1542,1567]" name="Myanmar" pageId="1" pageNumber="168">Myanmar</collectingCountry>
.
</paragraph>
<paragraph id="491981048070590CFF5D526D9E5F21D8" blockId="1.[136,1452,966,1675]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFF5D526D9CDC2110" bold="true" box="[189,449,1578,1603]" pageId="1" pageNumber="168">Specimens examined:</emphasis>
<materialsCitation id="F9CE8B598070590CFE29526D9DF42134" collectingDate="1981-10-24" collectionCode="HN" collectorName="D. D. Huyen" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
<emphasis id="7BD25D168070590CFE29526D9F4A2110" bold="true" box="[457,599,1578,1603]" pageId="1" pageNumber="168">
<collectingCountry id="31B1C1948070590CFE29526D9F4F2110" box="[457,594,1578,1603]" name="Vietnam" pageId="1" pageNumber="168">VIETNAM</collectingCountry>
.
</emphasis>
<collectingRegion id="8B624FE68070590CFD80526D9E452110" box="[608,856,1578,1603]" country="Vietnam" name="Quang Binh" pageId="1" pageNumber="168">Quang Binh Province</collectingRegion>
,
<collectingRegion id="8B624FE68070590CFC85526D994D2110" box="[869,1104,1578,1603]" country="Vietnam" name="Quang Ninh" pageId="1" pageNumber="168">Quang Ninh District</collectingRegion>
,
<date id="3D18A7C48070590CFBBD526D98032110" box="[1117,1310,1578,1603]" pageId="1" pageNumber="168" value="1981-10-24">
<collectingDate id="2D5C5E2C8070590CFBBD526D98032110" box="[1117,1310,1578,1603]" pageId="1" pageNumber="168" value="1981-10-24">24 October 1981</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFACC526D9DB92134" italics="true" pageId="1" pageNumber="168">
<collectorName id="E453E4D28070590CFACC526D98B62110" box="[1324,1451,1578,1603]" pageId="1" pageNumber="168">D.D.Huyen</collectorName>
88
</emphasis>
(
<collectionCode id="2FB719C18070590CFF5452099DFF2134" box="[180,226,1614,1639]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598070590CFF1652099E072134" box="[246,794,1614,1639]" collectingDate="2008-08-12" collectionCode="V, HN" collectorName="Hai" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
same locality,
<date id="3D18A7C48070590CFE7752099F512134" box="[407,588,1614,1639]" pageId="1" pageNumber="168" value="2008-08-12">
<collectingDate id="2D5C5E2C8070590CFE7752099F512134" box="[407,588,1614,1639]" pageId="1" pageNumber="168" value="2008-08-12">12 August 2008</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFDB752099FC92134" box="[599,724,1614,1639]" italics="true" pageId="1" pageNumber="168">
D.
<collectionCode id="2FB719C18070590CFD9052099F9F2134" box="[624,642,1614,1639]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="168" type="Museum">V</collectionCode>
.
<collectorName id="E453E4D28070590CFD6552099FAC2134" box="[645,689,1614,1639]" pageId="1" pageNumber="168">Hai</collectorName>
14
</emphasis>
(
<collectionCode id="2FB719C18070590CFD0452099E0F2134" box="[740,786,1614,1639]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598070590CFCC65209987B2134" box="[806,1382,1614,1639]" collectingDate="1981-08-22" collectionCode="T, K, HN" collectorName="Lien" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
same locality,
<date id="3D18A7C48070590CFC27520999662134" box="[967,1147,1614,1639]" pageId="1" pageNumber="168" value="1981-08-22">
<collectingDate id="2D5C5E2C8070590CFC27520999662134" box="[967,1147,1614,1639]" pageId="1" pageNumber="168" value="1981-08-22">22 August 1981</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFB685209983D2134" box="[1160,1312,1614,1639]" italics="true" pageId="1" pageNumber="168">
<collectionCode id="2FB719C18070590CFB68520999872134" box="[1160,1178,1614,1639]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="168">T</collectionCode>
.
<collectionCode id="2FB719C18070590CFB7D520999AE2134" box="[1181,1203,1614,1639]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="1" pageNumber="168" type="Herbarium">K</collectionCode>
.
<collectorName id="E453E4D28070590CFB5D520999F22134" box="[1213,1263,1614,1639]" pageId="1" pageNumber="168">Lien</collectorName>
385
</emphasis>
(
<collectionCode id="2FB719C18070590CFAD0520998432134" box="[1328,1374,1614,1639]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="F9CE8B598070590CFA9252099F8921D8" collectingDate="2003-06-07" collectionCode="N, Q" collectorName="Binh &amp; D. D. Cuong" country="Vietnam" location="Quang Binh Province" pageId="1" pageNumber="168" specimenCount="1" stateProvince="Quang Binh">
same locality,
<date id="3D18A7C48070590CFF0852359C6E21D8" box="[232,371,1650,1675]" pageId="1" pageNumber="168" value="2003-06-07">
<collectingDate id="2D5C5E2C8070590CFF0852359C6E21D8" box="[232,371,1650,1675]" pageId="1" pageNumber="168" value="2003-06-07">7 June 2003</collectingDate>
</date>
,
<emphasis id="7BD25D168070590CFE9E52359CF321D8" box="[382,494,1650,1675]" italics="true" pageId="1" pageNumber="168">
<collectionCode id="2FB719C18070590CFE9E52359C8A21D8" box="[382,407,1650,1675]" country="China" lsid="urn:lsid:biocol.org:col:13092" name="Nanjing University" pageId="1" pageNumber="168" type="Herbarium">N</collectionCode>
.
<collectionCode id="2FB719C18070590CFE7A52359CB221D8" box="[410,431,1650,1675]" country="Ecuador" lsid="urn:lsid:biocol.org:col:14273" name="Universidad Central" pageId="1" pageNumber="168" type="Herbarium">Q</collectionCode>
.
<collectorName id="E453E4D28070590CFE5952359CF321D8" box="[441,494,1650,1675]" pageId="1" pageNumber="168">Binh</collectorName>
</emphasis>
&amp;
<collectorName id="E453E4D28070590CFDF252359F8921D8" box="[530,660,1650,1675]" pageId="1" pageNumber="168">
<emphasis id="7BD25D168070590CFDF252359F8921D8" box="[530,660,1650,1675]" italics="true" pageId="1" pageNumber="168">D.D.Cuong</emphasis>
</collectorName>
</materialsCitation>
<materialsCitation id="F9CE8B598070590CFD7A52359E2321D8" box="[666,830,1650,1675]" collectionCode="HN" country="Vietnam" location="VN" pageId="1" pageNumber="168" specimenCount="1">
<emphasis id="7BD25D168070590CFD7A52359FE421D8" box="[666,761,1650,1675]" italics="true" pageId="1" pageNumber="168">
<collectingCountry id="31B1C1948070590CFD7A52359FA321D8" box="[666,702,1650,1675]" name="Vietnam" pageId="1" pageNumber="168">VN</collectingCountry>
1112
</emphasis>
(
<collectionCode id="2FB719C18070590CFCE852359E2B21D8" box="[776,822,1650,1675]" country="Viet Nam" lsid="urn:lsid:biocol.org:col:13708" name="National Center for Natural Sciences and Technology" pageId="1" pageNumber="168" type="Herbarium">HN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>