532 lines
56 KiB
XML
532 lines
56 KiB
XML
<document id="91772AB854AD2B93EE5CA2A3B74CA94D" ID-CLB-Dataset="298085" ID-DOI="10.3897/zookeys.1204.120164" ID-GBIF-Dataset="31abc999-2e6e-474c-a4f3-149976ea0813" ID-ZooBank="02D0B0C3-219A-46F6-A433-D1D77CE5F312" ID-publisher-id="120164" URI-arpha="700BDC66-D10E-588B-806C-D4467FDA22C6" URI-zoobank="http://zoobank.org/02D0B0C3-219A-46F6-A433-D1D77CE5F312" XM.bibliography_approvedBy="admin" XM.materialsCitations_approvedBy="admin" XM.treatmentCitations_approvedBy="admin" article-type="research-article" checkinTime="1717518109694" checkinUser="pensoft" docAuthor="Fan, Wenlong, Zhang, Chao & Zhang, Feng" docDate="2024" docId="B6E20436911855ED8C022D2A376881F0" docLanguage="en" docName="ZooKeys 1204: 155-190" docOrigin="ZooKeys 1204" docSource="https://zookeys.pensoft.net/article/120164/download/xml/" docStyle="DocumentStyle:PensoftTaxPub.0000.journal_article.generic" docStyleName="PensoftTaxPub.0000.journal_article.generic" docTitle="Karschia (Karschia) namling Fan & Zhang & Zhang 2024, sp. nov." docType="treatment" docUuid="D73D6B1F-73C4-4D8A-8802-AF0C93B46AE6" docUuidSource="ZooBank" docVersion="6" dtd-version="3.0" lastPageNumber="190" masterDocId="700BDC66D10E588B806CD4467FDA22C6" masterDocTitle="Review of the genus Karschia Walter, 1889 from Xizang, China (Solifugae, Karschiidae)" masterLastPageNumber="190" masterPageNumber="155" pageNumber="155" updateTime="1717541794717" updateUser="ExternalLinkService">
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<mods:title id="94D9F62734D6CBFE2A768AD929CA7662">Review of the genus Karschia Walter, 1889 from Xizang, China (Solifugae, Karschiidae)</mods:title>
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<mods:name id="A1B8060AECE3A84C09D9388E48396187" type="personal">
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<mods:namePart id="030D840FA5B7D710FB9F76DD752080C2">Fan, Wenlong</mods:namePart>
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<mods:affiliation id="351A3BC784FE394550C5A94A553355B5">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="12E56E269D74C3069E6B15193B1B7E8C">Zhang, Chao</mods:namePart>
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<mods:affiliation id="9ED05D98F9A96DDB426363756B33EE6D">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:namePart id="0628BD22A810ABC1DE1809B9D37DF0A6">Zhang, Feng</mods:namePart>
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<mods:affiliation id="BBD4473BAE949E9BDF2E80EEC5031F29">Key Laboratory of Zoological Systematics and Application, College of Life Sciences, Hebei University, Baoding, Hebei 071002, China</mods:affiliation>
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<mods:date id="44915BE93F73D964015486E8471BF8D9">2024</mods:date>
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<mods:number id="E1EBD257DDAFDF650DCD696717E97CAA">2024-06-04</mods:number>
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<mods:number id="15317FFB5F7567ADC96FB7DEB24D190B">1204</mods:number>
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<treatment id="B6E20436911855ED8C022D2A376881F0" ID-DOI="http://doi.org/10.5281/zenodo.11476738" ID-GBIF-Taxon="232137232" ID-Zenodo-Dep="11476738" ID-arpha="B6E20436-9118-55ED-8C02-2D2A376881F0" ID-zoobank="https://zoobank.org/D73D6B1F-73C4-4D8A-8802-AF0C93B46AE6" LSID="urn:lsid:zoobank.org:act:D73D6B1F-73C4-4D8A-8802-AF0C93B46AE6" LSID-ZBK="urn:lsid:zoobank.org:act:D73D6B1F-73C4-4D8A-8802-AF0C93B46AE6" httpUri="http://treatment.plazi.org/id/B6E20436911855ED8C022D2A376881F0">
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<subSubSection id="B39C5B6FC2CAD44D3AF16D191357631E" type="nomenclature">
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<paragraph id="1891E41272ECD5EDDF4D3078D7686D89">
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<taxonomicName id="626D5BE3BDA53FE9B4F93871FB831689" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="namling" status="sp. nov." subGenus="Karschia">
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<emphasis id="E1ED405A6A35FE41C5871CE02C652DD0" italics="true">Karschia</emphasis>
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(
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<emphasis id="FB5D239689D3D8517D83741F9B7E1106" italics="true">Karschia</emphasis>
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)
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<emphasis id="76FB2E2614602EB3526A12D7B2083E88" italics="true">namling</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="7716FA3FD245A659BF24E8BB3E9893D4" rank="species">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="72D6B6F182DADC634354198C5E4E99D1" type="description">
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<paragraph id="B4592E99A6EF81E30A58DB499AE693AD">
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<figureCitation id="322BB3E0D1F364F12435CEE06DFEBDB0" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Map plotting known locality records." figureDoi="10.3897/zookeys.1204.120164.figure1" httpUri="https://binary.pensoft.net/fig/1062103">Figs 1</figureCitation>
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,
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<figureCitation id="B5769E315E83F4979BB0042E2C15CB9D" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat where K. dingye sp. nov. (A, B), K. lhasa sp. nov. (C), K. shigatse sp. nov. (E – G) and K. namling sp. nov. (H) have been found. A, B Xizang, Dingye County, Gyangkar Town C Xizang, Lhasa City, Maizhokunggar County D Xizang, Lhasa City, Drepung Monastery E, F Xizang, Nyalam County, Mainqu Town G Xizang, Gyirong County, Zheba Town H Xizang, Namling County, Nubma Town." figureDoi="10.3897/zookeys.1204.120164.figure2" httpUri="https://binary.pensoft.net/fig/1062104">2 H</figureCitation>
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,
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<figureCitation id="A3C2ADA389C03D5A415043108B04232D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus A – D K. shigatse sp. nov., habitus, male (A, B) and female (C, D) E – H K. namling sp. nov., habitus, male (E, F) and female (G, H). Scale bars: 10 mm." figureDoi="10.3897/zookeys.1204.120164.figure5" httpUri="https://binary.pensoft.net/fig/1062107">5 E – H</figureCitation>
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,
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<figureCitation id="72C97D03E1D6C1C5C00B3B2A2D4BA287" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Propeltidium A. K. zhui sp. nov., holotype male B. K. zhui sp. nov., paratype female C. K. shigatse sp. nov., holotype male D. K. shigatse sp. nov., paratype female E. K. namling sp. nov., holotype male F K. namling sp. nov., paratype female. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1204.120164.figure7" httpUri="https://binary.pensoft.net/fig/1062109">7 E, F</figureCitation>
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||
,
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<figureCitation id="FD82EEECAB10A32AAA84A3DE316DCF5B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure10" httpUri="https://binary.pensoft.net/fig/1062112">10 E – H</figureCitation>
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||
,
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<figureCitation id="DD95E0D43507CAF6A3C54A8BA5DE7B3F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Flagellum A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure11" httpUri="https://binary.pensoft.net/fig/1062113">11 F</figureCitation>
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||
,
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||
<figureCitation id="3EBE8F7C0543093356BBF5463D8096A5" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female fixed finger ventral views A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFSM, retrofondal submedial tooth / teeth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure12" httpUri="https://binary.pensoft.net/fig/1062114">12 F</figureCitation>
|
||
,
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||
<figureCitation id="4011A21475933FA56B7C1CCF4ADC326A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RF, retrofondal teeth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PF, profondal teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth; MSP, movable finger, subproximal tooth / teeth; MP, movable finger, proximal tooth; MSM, movable finger, submedial tooth / teeth; MM, movable finger, medial tooth; MST, movable finger, subterminal teeth; pvd, proventral distal setae; fcp (modified pvd), flagellar complex plumose setae; pdp, prodo rsal proximal setae; fcs, flagellar complex subspiniform to spiniform setae; pvsd, proventral subdistal setae; pvd, proventral distal setae. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure15" httpUri="https://binary.pensoft.net/fig/1062117">15 E – H</figureCitation>
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,
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<figureCitation id="3307A1BFB9630309B49648101BC57C75" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male pedipalp A, B K. tibetana C, D K. dingye sp. nov. E, F K. lhasa sp. nov. G, H K. zhui sp. nov. I, J K. shigatse sp. nov. K, L K. namling sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure16" httpUri="https://binary.pensoft.net/fig/1062118">16 K, L</figureCitation>
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,
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<figureCitation id="8E56890D72578EF789C98C139D9EC7D9" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Genital operculum of female A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure17" httpUri="https://binary.pensoft.net/fig/1062119">17 F</figureCitation>
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,
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<figureCitation id="AF6A713487C0343E395F2DBA7E76FA9A" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ctenidia on sternite IV of female A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure18" httpUri="https://binary.pensoft.net/fig/1062120">18 E</figureCitation>
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,
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<figureCitation id="1C8B8671F3661D8CEFD0FE8BC06F697B" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ctenidia on sternite III (A, B, E, F, I, J) and sternite IV (C, D, G, H, K, L) of male A, C K. tibetana B, D K. dingye sp. nov. E, G K. lhasa sp. nov. F, H K. zhui sp. nov. I, K. K. shigatse sp. nov. J, L K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure19" httpUri="https://binary.pensoft.net/fig/1062121">19 J, L</figureCitation>
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||
,
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<tableCitation id="8D4841E0CBA2383F3954765B3779F14C" captionStart="Table 1" captionStartId="T1" captionText="Table 1. Voucher specimen information." httpUri="http://table.plazi.org/id/01146FFD08B653C3C7E9907E7C5B0943" tableUuid="01146FFD08B653C3C7E9907E7C5B0943">Tables 1</tableCitation>
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,
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<tableCitation id="29E7DE3E048691C97A4504182F96B814" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Genetic distance among the six species." httpUri="http://table.plazi.org/id/AAF6FF40179CED3530634E534F91C65C" tableUuid="AAF6FF40179CED3530634E534F91C65C">2</tableCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="SECID0ENODI" type="type material">
|
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<paragraph id="18DAEA2CF5AAAD29AA3397BDB0C975DA">
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<heading id="6124E703F3EC8EA310550D375F07174A" reason="title">Type material.</heading>
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</paragraph>
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<paragraph id="4A5504B44B9A98566E1DB7D1E2F39749">
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<materialsCitation id="017AE79808ACBE866C8DE814A9F24E1E" ID-GBIF-Occurrence="4873240314" collectingDate="2023-08-30" collectionCode="MHBU" collectorName="Yanmeng Hou & Zhiyong Yang" country="China" county="Namling County" elevation="4016" latitude="29.5172" location="Nubma Town" longLatPrecision="6" longitude="89.6237" municipality="Nubma Town" specimenCode="Sol-XZ 2023073001" specimenCount="1" specimenCount-male="1" stateProvince="Xizang" typeStatus="holotype">
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<emphasis id="9891ABE523BE45C21250F120A8EE417B" bold="true" italics="true">
|
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<typeStatus id="37E479027BEC1006F5196A3B3ADC4319" type="holotype">Holotype</typeStatus>
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</emphasis>
|
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<specimenCount id="1A025D4DA5C51463E11E4F5CDA98A0F7" count="1" type="male">♂</specimenCount>
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(
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<abbrev id="ABBRID0EYODI" xlink_title="Museum of Hebei University">
|
||
<collectionCode id="E3F1C4D963C34BEBD0CD2A6AD8CAD6D6">MHBU</collectionCode>
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</abbrev>
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-
|
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<specimenCode id="3659A795C716C360FB877B38AF7BF66A">Sol-XZ 2023073001</specimenCode>
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),
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<collectingCountry id="436F78795780F731CE6E046C2FA8751C" name="China">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="4E1BCD20B28897D2B8DF8D124997139D" country="China" name="Xizang">Xizang</collectingRegion>
|
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,
|
||
<collectingCounty id="3325E0E2347C4392B537ED16B503A12B">Namling County</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="B4BA2BC4712E27C40046E8A7ED7BB2DE" country="China" county="Namling County" latitude="29.5172" longLatPrecision="6" longitude="89.6237" name="Nubma Town" stateProvince="Xizang">Nubma Town</collectingMunicipality>
|
||
,
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E6ODI" specific-use="{"type":"Point","coordinates":[89.623700,29.517200]}">
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<geoCoordinate id="C66D6F05FE9129FD99CDDF7FE1E0AF98" degrees="29.5172" direction="north" orientation="latitude" precision="5" value="29.5172">29.5172 ° N</geoCoordinate>
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,
|
||
<geoCoordinate id="B4CA2CC0ED2F7DCA8FEC5652515404A0" degrees="89.6237" direction="east" orientation="longitude" precision="5" value="89.6237">89.6237 ° E</geoCoordinate>
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</named-content>
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,
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<quantity id="0AFD1EB6BCFC13FE32D97428CC0ECA4D" metricMagnitude="3" metricUnit="m" metricValue="4.01627" unit="m" value="4016.27">
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<elevation id="A7EEA74CDFE042AEA2FB07BDED36F85F" metricMagnitude="3" metricUnit="m" metricValue="4.01627" unit="m" value="4016.27">4016.27 m</elevation>
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||
</quantity>
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||
elev.,
|
||
<date id="03485705ABC9B87316AECC9105E529E2" value="2023-08-30">
|
||
<collectingDate id="FA531B610757842C57ED74064953EA06" value="2023-08-30">30. VIII. 2023</collectingDate>
|
||
</date>
|
||
, leg.
|
||
<collectorName id="BE5741515871FBD012CB55496CF3C78D">Yanmeng Hou</collectorName>
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||
,
|
||
<collectorName id="12ED0EEE87008C95F993CFD4ABA801F1">Zhiyong Yang</collectorName>
|
||
.
|
||
</materialsCitation>
|
||
<materialsCitation id="971E14B07BD703F3EC4DCA4A8B58940E" ID-GBIF-Occurrence="4873240304" collectingDate="2023-08-30" collectionCode="MHBU" collectorName="Yanmeng Hou & Zhiyong Yang" country="China" county="Namling County" elevation="4016" latitude="29.5172" location="Nubma Town" longLatPrecision="6" longitude="89.6237" municipality="Nubma Town" specimenCode="Sol-XZ 2023073002, Sol-XZ 2023073003" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Xizang" typeStatus="paratype">
|
||
<emphasis id="3550D4F4A84E43B1259337BD4F690F57" bold="true" italics="true">
|
||
<typeStatus id="7548F9016851014DF3CAC095CBF0CB90" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<specimenCount id="D221FF7B271D961B2889062F9096D41B" count="1" type="male">1 ♂</specimenCount>
|
||
(
|
||
<abbrev id="ABBRID0EHPDI" xlink_title="Museum of Hebei University">
|
||
<collectionCode id="728933A9DC07669D12DCEC516006E97E">MHBU</collectionCode>
|
||
</abbrev>
|
||
-
|
||
<specimenCode id="C1D8B50888CFD147153648D0209333AC">Sol-XZ 2023073002</specimenCode>
|
||
),
|
||
<specimenCount id="0229B255F4C5C185EE7E2FEF91505489" count="4" type="female">4 ♀♀</specimenCount>
|
||
(
|
||
<abbrev id="ABBRID0ELPDI" xlink_title="Museum of Hebei University">
|
||
<collectionCode id="286546BAEF89CB6664712CAC76AE6FE3">MHBU</collectionCode>
|
||
</abbrev>
|
||
-
|
||
<specimenCode id="53CB60FDFF05DA21DCA487FCFFEE3202">Sol-XZ 2023073003</specimenCode>
|
||
–07), same data as holotype
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EPPDI" type="etymology">
|
||
<paragraph id="BFDD9572F6B9A0206FE72E6009334C52">
|
||
<heading id="CF71F18464D887FD7C07705DB3C0B1DB" reason="title">Etymology.</heading>
|
||
</paragraph>
|
||
<paragraph id="88C1520441B6BB95A797682D0F7BA8B7">Noun in apposition taken from Namling County, where this species was collected.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EUPDI" type="diagnosis">
|
||
<paragraph id="882DADEF8A15F1C60D8ED0629BE9837F">
|
||
<heading id="2FA3A73C0B4AC1C4E5EE9EDF8A179555" reason="title">Diagnosis.</heading>
|
||
</paragraph>
|
||
<paragraph id="DEC490D032F426724349C7D4336AB2A9">
|
||
<taxonomicName id="B7AEB8F93F40D7786ADAFB25CD29A6EF" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="namling" status="sp. nov.">
|
||
<emphasis id="738A1CAF1180BE4D1A0255BC3C1572C1" italics="true">K. namling</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="B10ADA102C93195EDB6B5D0F3FE23F4F" rank="species">sp. nov.</taxonomicNameLabel>
|
||
differs from
|
||
<taxonomicName id="E6A05E08B3DC7D385509A6395D0A0461" authorityName="Lawrence" authorityYear="1954" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="nubigena">
|
||
<emphasis id="20318BB5B7222DFBB61D077CD0C65D12" italics="true">K. nubigena</emphasis>
|
||
</taxonomicName>
|
||
by have fringed flagellum (Fig.
|
||
<figureCitation id="390AC7F1855B869C08D7DC17898A995F" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Flagellum A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure11" httpUri="https://binary.pensoft.net/fig/1062113">11 F</figureCitation>
|
||
), differs from
|
||
<taxonomicName id="A1B17ED87A2DC14345BE1967BD2E5EE2" authorityName="Hirst" authorityYear="1907" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="tibetana">
|
||
<emphasis id="50D45B27FFCEB2290957CB364F5EA0CC" italics="true">K. tibetana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="44F31E25BC8B0BD9BF137F2C24DCFFE6" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="dingye" status="sp. nov.">
|
||
<emphasis id="C9959477D0F2F0961A5FF33F30B9460E" italics="true">K. dingye</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="8644BDF489AA4BE4D00CC018CA28587A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
<taxonomicName id="31A7A720958D50A078D2E8D396E0DADB" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="lhasa" status="sp. nov.">
|
||
<emphasis id="516C1B6E3164F5E02B4E92F01F35AB61" italics="true">K. lhasa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94D87517A3436EFF346CCD87B6C9F5E5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="DA9A2716B80D12FA3A42CD57FABDF8C2" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="zhui" status="sp. nov.">
|
||
<emphasis id="1594EFC94E78ADB6D829A7CEBC460532" italics="true">K. zhui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="BA0DEB2827507089CBA31F8DEA5BF667" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by flagellum with lateral apophysis (Fig.
|
||
<figureCitation id="C4E0726E45BA82CC0626B4AC26399DF9" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Flagellum A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure11" httpUri="https://binary.pensoft.net/fig/1062113">11 F</figureCitation>
|
||
), and from
|
||
<taxonomicName id="BFCCDCAE58458B0452EC5AB4DB502478" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="shigatse" status="sp. nov.">
|
||
<emphasis id="1FB670CE068D7545473977AB470C4988" italics="true">Karschia shigatse</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="554313AE47E0A1F1B5695F5F73118C4B" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by less wide cheliceral fixed finger mucron, flagellum with small lateral apophysis (Fig.
|
||
<figureCitation id="2F7A5327128D6B08977FB78D7538B646" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Flagellum A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure11" httpUri="https://binary.pensoft.net/fig/1062113">11 F</figureCitation>
|
||
). Female differs from other species by its genital operculum triangular (Fig.
|
||
<figureCitation id="97209352969FC10A24AE5DE4D7086547" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Genital operculum of female A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure17" httpUri="https://binary.pensoft.net/fig/1062119">17 F</figureCitation>
|
||
) and ctenidia on sternite IV very short (Fig.
|
||
<figureCitation id="EDBD5E4203F2DB2702E6F55EDCE02716" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ctenidia on sternite IV of female A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure18" httpUri="https://binary.pensoft.net/fig/1062120">18 F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E2SDI" type="description">
|
||
<paragraph id="F9E414301B43EE1936F800A26FF3729F">
|
||
<heading id="D264D50046D37914697AB2FEC833F782" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="45C6A2085F5576CE27B068449B3AEF0E">
|
||
<emphasis id="CEA7CFBAC17674269175633858C6B0B9" bold="true">Male.</emphasis>
|
||
<typeStatus id="8CD16582C406AA5F620191D16AE80BB1" type="holotype">Holotype</typeStatus>
|
||
(
|
||
<abbrev id="ABBRID0EDTDI" xlink_title="Museum of Hebei University">
|
||
<collectionCode id="FEEAC8BB9FD2B3AE5DD0703A262708CD">MHBU</collectionCode>
|
||
</abbrev>
|
||
-
|
||
<specimenCode id="1E4D736E8B0741590C90CFFEAA4E90D4">Sol-XZ 2023073001</specimenCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="2FE224BB110AC99B1E9DE4EFB8275E58">
|
||
<emphasis id="E79347D20ACFC101F59C0B1D93FAEABA" bold="true">
|
||
<emphasis id="D17B3FD96C420F256FCC12B69800E0A2" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Total body length 16.61,
|
||
<abbrev id="ABBRID0EOTDI" xlink_title="chelicera length">CL</abbrev>
|
||
4.68,
|
||
<abbrev id="ABBRID0ESTDI" xlink_title="chelicera height">CH</abbrev>
|
||
1.58,
|
||
<abbrev id="ABBRID0EWTDI" xlink_title="Propeltidium length">PL</abbrev>
|
||
2.35, PW 3.04,
|
||
<abbrev id="ABBRID0E1TDI" xlink_title="sum of the lengths of pedipalp, leg I, and leg IV divided by the sum of the lengths of chelicera and propeltidium, indicating the length of appendages in relation to body size">A / CP</abbrev>
|
||
8.66,
|
||
<abbrev id="ABBRID0E5TDI" xlink_title="chelicera length/height, large CL/CH ratios suggest a narrow cheliceral morphology, while a more robust morphology is represented by a smaller ratio">CL / CH</abbrev>
|
||
2.97. Pedipalp 22.21 (6.51, 6.56, 4.60, 1.50), Leg I 14.99 (3.56, 3.93, 2.59, 1.36, 0.12), Leg II 11.46 (2.52, 2.65, 1.46, 1.17, 0.19), Leg III 16.72 (3.45, 4.39, 2.86, 0.92, 0.92), Leg IV 23.73 (4.81, 6.47, 3.54, 1.42, 1.43).
|
||
</paragraph>
|
||
<paragraph id="1A3E00653DA73FCCBBD8977829B91288">
|
||
<emphasis id="577183AC07D21B65CB64D00C630DF00D" bold="true">
|
||
<emphasis id="105D4340E2FE525B1AEFD3B44C9886A7" bold="true" italics="true">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
In 95 % ethanol-preserved specimens (Fig.
|
||
<figureCitation id="8D3685F4CE3FC9D565D0E5BD6FA1090F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus A – D K. shigatse sp. nov., habitus, male (A, B) and female (C, D) E – H K. namling sp. nov., habitus, male (E, F) and female (G, H). Scale bars: 10 mm." figureDoi="10.3897/zookeys.1204.120164.figure5" httpUri="https://binary.pensoft.net/fig/1062107">5 E, F</figureCitation>
|
||
). The general background pale yellowish. Opisthosoma slightly darker, with black tergites and yellow around the black sternites. Propeltidium pale tan and tinged with pale brown. Ocular tubercle black. Mesopeltidium and metapeltidium with special black stripes. Chelicerae with manus predominantly brown yellowish, with some black areas, and a retrolateral view of chelicerae with three black longitudinal stripes. Pedipalps and legs pale yellow, legs III and legs IV tinged with pale brown on distal regions of femora and proximal parts of tibiae. Proximal regions of the pedipalpal femur, tibia, metatarsus, and tarsus were tinged with brown. Malleoli white.
|
||
</paragraph>
|
||
<paragraph id="6DA10E03C478D299BCBECFDBF866C6FD">
|
||
<emphasis id="9689749E16EE31AE72623FEBFD7470DE" bold="true">
|
||
<emphasis id="1F09CC1914565BAEBA786A7CDCBB951B" bold="true" italics="true">Propeltidium</emphasis>
|
||
.
|
||
</emphasis>
|
||
Wider than long with a dense pubescence of thin, short, anteriorly directed setae. Anterior, posterior, and lateral edges with several long, curved spiniform setae that stand perpendicular to the surface of the propeltidium. Ocular tubercle with four middle distal spiniform setae, one middle spiniform setae, and one proximal spiniform setae. (Fig.
|
||
<figureCitation id="E1301EFE6F40F0B0497F7A973D420428" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Propeltidium A. K. zhui sp. nov., holotype male B. K. zhui sp. nov., paratype female C. K. shigatse sp. nov., holotype male D. K. shigatse sp. nov., paratype female E. K. namling sp. nov., holotype male F K. namling sp. nov., paratype female. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1204.120164.figure7" httpUri="https://binary.pensoft.net/fig/1062109">7 E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B53370A12D9B0991819E0FC937A2EDDC">
|
||
<emphasis id="6C68734D1BB08D5F130CE4B639D05BB4" bold="true">
|
||
<emphasis id="C494C487CFFB5C2A81DDD3AE0D74D9AA" bold="true" italics="true">Chelicerae</emphasis>
|
||
.
|
||
</emphasis>
|
||
Fixed finger primary teeth graded as
|
||
<abbrev id="ABBRID0EKVDI" xlink_title="fixed finger, distal tooth">FD</abbrev>
|
||
<
|
||
<abbrev id="ABBRID0EOVDI" xlink_title="fixed finger, medial tooth">FM</abbrev>
|
||
≈
|
||
<abbrev id="ABBRID0ESVDI" xlink_title="fixed finger, proximal tooth">FP</abbrev>
|
||
. Profondal teeth series with three tiny teeth; retrofondal teeth series with six teeth. Dental formulation of fixed finger:
|
||
<abbrev id="ABBRID0EWVDI" xlink_title="fixed finger, distal tooth">FD</abbrev>
|
||
- (2) -
|
||
<abbrev id="ABBRID0E1VDI" xlink_title="fixed finger, medial tooth">FM</abbrev>
|
||
- (2) -
|
||
<abbrev id="ABBRID0E5VDI" xlink_title="fixed finger, proximal tooth">FP</abbrev>
|
||
- (6 RF) (3 PF). Fixed finger mucron without dorsal crest. Movable finger
|
||
<abbrev id="ABBRID0ECWDI" xlink_title="movable finger, proximal tooth">MP</abbrev>
|
||
tooth about the same size as
|
||
<abbrev id="ABBRID0EGWDI" xlink_title="movable finger, medial tooth">MM</abbrev>
|
||
. Dental formulation of movable finger:
|
||
<abbrev id="ABBRID0EKWDI" xlink_title="movable finger, medial tooth">MM</abbrev>
|
||
- (2) -
|
||
<abbrev id="ABBRID0EOWDI" xlink_title="movable finger, proximal tooth">MP</abbrev>
|
||
, with two tiny
|
||
<abbrev id="ABBRID0ESWDI" xlink_title="movable finger, submedial tooth/teeth">MSM</abbrev>
|
||
and three
|
||
<abbrev id="ABBRID0EWWDI" xlink_title="movable finger, subproximal tooth/teeth">MSP</abbrev>
|
||
(Figs
|
||
<figureCitation id="E7731A8DD91E529A28A111084335E59D" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure10" httpUri="https://binary.pensoft.net/fig/1062112">10 E</figureCitation>
|
||
,
|
||
<figureCitation id="3EBCC4E4AF5722461F11D42FE69D6B2F" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RF, retrofondal teeth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PF, profondal teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth; MSP, movable finger, subproximal tooth / teeth; MP, movable finger, proximal tooth; MSM, movable finger, submedial tooth / teeth; MM, movable finger, medial tooth; MST, movable finger, subterminal teeth; pvd, proventral distal setae; fcp (modified pvd), flagellar complex plumose setae; pdp, prodo rsal proximal setae; fcs, flagellar complex subspiniform to spiniform setae; pvsd, proventral subdistal setae; pvd, proventral distal setae. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure15" httpUri="https://binary.pensoft.net/fig/1062117">15 E</figureCitation>
|
||
). Flagellum coiled, fringed and sessile, with a small lateral apophysis. The flagellar complex includes two short
|
||
<abbrev id="ABBRID0EDXDI" xlink_title="(modified pvd) , flagellar complex plumose setae">
|
||
<emphasis id="C15900E585A8007B9AC9622D784B1923" italics="true">fcp</emphasis>
|
||
</abbrev>
|
||
and two short, thick
|
||
<abbrev id="ABBRID0EIXDI" xlink_title="flagellar complex subspiniform to spiniform setae">
|
||
<emphasis id="EAE834C5C665AA7D8B92AD146B88D224" italics="true">fcs</emphasis>
|
||
</abbrev>
|
||
(Figs
|
||
<figureCitation id="87F70CD69CAB689811706EDFDE63B2D5" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure10" httpUri="https://binary.pensoft.net/fig/1062112">10 F</figureCitation>
|
||
,
|
||
<figureCitation id="3A327DDA7C0D6B53491D5E4469A42E81" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Flagellum A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure11" httpUri="https://binary.pensoft.net/fig/1062113">11 F</figureCitation>
|
||
,
|
||
<figureCitation id="33F654C27465CD5E02A52DF8BA54DF32" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RF, retrofondal teeth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PF, profondal teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth; MSP, movable finger, subproximal tooth / teeth; MP, movable finger, proximal tooth; MSM, movable finger, submedial tooth / teeth; MM, movable finger, medial tooth; MST, movable finger, subterminal teeth; pvd, proventral distal setae; fcp (modified pvd), flagellar complex plumose setae; pdp, prodo rsal proximal setae; fcs, flagellar complex subspiniform to spiniform setae; pvsd, proventral subdistal setae; pvd, proventral distal setae. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure15" httpUri="https://binary.pensoft.net/fig/1062117">15 F</figureCitation>
|
||
). Retrolateral and dorsal surfaces of the manus with large, bifurcated tip setae and short, simple tip bristle-like setae; retrolateral and dorsal surfaces of the fixed finger with simple tip setae of different sizes. Retrolateral setose area reaching the
|
||
<abbrev id="ABBRID0EYXDI" xlink_title="fixed finger, submedial tooth/teeth">FSM</abbrev>
|
||
teeth; prolateral surface with an array of setal types (Figs
|
||
<figureCitation id="3069A429BB71B904C8E8C43C1A6E7BC6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure10" httpUri="https://binary.pensoft.net/fig/1062112">10 E, F</figureCitation>
|
||
,
|
||
<figureCitation id="61D4EDCFF57989A8A7AEBF1B52DE778B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RF, retrofondal teeth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PF, profondal teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth; MSP, movable finger, subproximal tooth / teeth; MP, movable finger, proximal tooth; MSM, movable finger, submedial tooth / teeth; MM, movable finger, medial tooth; MST, movable finger, subterminal teeth; pvd, proventral distal setae; fcp (modified pvd), flagellar complex plumose setae; pdp, prodo rsal proximal setae; fcs, flagellar complex subspiniform to spiniform setae; pvsd, proventral subdistal setae; pvd, proventral distal setae. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure15" httpUri="https://binary.pensoft.net/fig/1062117">15 E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D2ECBC250F6B58750C3C86474C880D23">
|
||
<emphasis id="13E723D26E751FE02BADD1CCD232A627" bold="true">
|
||
<emphasis id="75B3547AD5E7973BA31DCD10261C2469" bold="true" italics="true">Opisthosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
The entire surface covered with almost adpressed setae and numerous long, curved, bifurcate setae. Sternite III with numbers short and cylindrical ctenidia (Fig.
|
||
<figureCitation id="30C74619FC1B5501804FA36D27919817" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ctenidia on sternite III (A, B, E, F, I, J) and sternite IV (C, D, G, H, K, L) of male A, C K. tibetana B, D K. dingye sp. nov. E, G K. lhasa sp. nov. F, H K. zhui sp. nov. I, K. K. shigatse sp. nov. J, L K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure19" httpUri="https://binary.pensoft.net/fig/1062121">19 J</figureCitation>
|
||
). Sternite IV with 14 long peg-like ctenidia, the length of which almost 1 / 3 the width of the succeeding sternite (Fig.
|
||
<figureCitation id="520EE80A4CEC5C8B9707B7FC2BB70440" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Ctenidia on sternite III (A, B, E, F, I, J) and sternite IV (C, D, G, H, K, L) of male A, C K. tibetana B, D K. dingye sp. nov. E, G K. lhasa sp. nov. F, H K. zhui sp. nov. I, K. K. shigatse sp. nov. J, L K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure19" httpUri="https://binary.pensoft.net/fig/1062121">19 L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="196A19164A1757DEF39E7E9A720D2273" ID-DOI="10.3897/zookeys.1204.120164.figure19" ID-arpha="C98C5AC0-B6E8-5129-B08A-4435A9006F79" httpUri="https://binary.pensoft.net/fig/1062121" startId="F19">
|
||
<paragraph id="9931F28DEEDBEF9F22491A027D181F75">
|
||
<label id="778218F9FC5598C968D11D704C9B4B65">Figure 19.</label>
|
||
</paragraph>
|
||
<paragraph id="EF0CD8640B38BB32686D129683EE5CE3">
|
||
Ctenidia on sternite III (A, B, E, F, I, J) and sternite IV (C, D, G, H, K, L) of male
|
||
<emphasis id="F579983819625544CF1CD5B963261C92" bold="true">A, C</emphasis>
|
||
<taxonomicName id="4FC01CD0938E4314CFF735F8BBDB3748" authorityName="Hirst" authorityYear="1907" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="tibetana">
|
||
<emphasis id="8A5DE44574DD79FF01C02F3C4EB2BE72" italics="true">K. tibetana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="9DBE8647D7D7D78A1E707A381F22710A" bold="true">B, D</emphasis>
|
||
<taxonomicName id="5EA42DB94D64DD9014BAAEC2BEBF33FD" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="dingye" status="sp. nov.">
|
||
<emphasis id="A69B05FF4D7B71F72A3396D358D25F73" italics="true">K. dingye</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="185BF5264699AB492BB835D7B61A4F79" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="DFF3599F0E0357A60D058C1CBB1FC265" bold="true">E, G</emphasis>
|
||
<taxonomicName id="08BF40CA603FDC7FCBB3AF5E058218E9" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="lhasa" status="sp. nov.">
|
||
<emphasis id="AAEFCAA51777D3A7165157554D814FE7" italics="true">K. lhasa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FCD553AF5312E68B19245A0F95D7722A" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="DCF5B44D36FA60B3A69CE6B5BFA7DC49" bold="true">F, H</emphasis>
|
||
<taxonomicName id="B6E57B10F015F4514C1DEEE6B185C073" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="zhui" status="sp. nov.">
|
||
<emphasis id="BEC642809C32225387752A530B2639F0" italics="true">K. zhui</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5D68F79DAF7328AE6D55786A455965C" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="24B61F6087E96F4863A1D21F86EC41B7" bold="true">I, K.</emphasis>
|
||
<taxonomicName id="137DC4A368EAEF3C8EAA64ACF1BD96AC" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="shigatse" status="sp. nov.">
|
||
<emphasis id="07257482A49A7025E384811E2B3D190E" italics="true">K. shigatse</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="81B5608A46A1461030890E83833BB787" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="77305202885D0D39DB844426AADE66D0" bold="true">J, L</emphasis>
|
||
<taxonomicName id="EC6D91BFCE527A78E39E030A0DEB4CBC" authority="Fan & Zhang & Zhang, 2024" authorityName="Fan & Zhang & Zhang" authorityYear="2024" class="Arachnida" family="Karschiidae" genus="Karschia" kingdom="Animalia" order="Solifugae" phylum="Arthropoda" rank="species" species="namling" status="sp. nov.">
|
||
<emphasis id="A005052EA251984327585ABE709E6F6A" italics="true">K. namling</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9929841F9FED090EA998D19422952376" rank="species">sp. nov.</taxonomicNameLabel>
|
||
Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="2B094E73A703F20D9A2CF989A547E7CE">
|
||
<emphasis id="0DA487CE37F51B39FE8103D693A4A073" bold="true">
|
||
<emphasis id="17816A0687E1DBBAB84E8F54E72E2FBC" bold="true" italics="true">Pedipalps</emphasis>
|
||
.
|
||
</emphasis>
|
||
Totally covered with short setae and long, thick setae. Tarsus with six sturdy ventral spines; metatarsus with eight ventral spines not arranged in pairs and with thick papillae (Fig.
|
||
<figureCitation id="4C0B1E77BE55709FBB9EF19DAD5423A4" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Male pedipalp A, B K. tibetana C, D K. dingye sp. nov. E, F K. lhasa sp. nov. G, H K. zhui sp. nov. I, J K. shigatse sp. nov. K, L K. namling sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure16" httpUri="https://binary.pensoft.net/fig/1062118">16 K, L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CC01BC9891B92177172F67069DEE0819">
|
||
<emphasis id="839CEED5C76902ED3DEC5A5AE7076971" bold="true">
|
||
<emphasis id="D31E18C0666758939BF2FC78C94BDBC8" bold="true" italics="true">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
Totally covered with long, thick setae and short setae. Leg I with no spines and two small claws. Tibias II, III, and IV with a pair of distal spines ventrally. Tibias II and III with a single dorsal spine; metatarsi II and III with a series of three dorsal spines, a pair of distal spines ventrally, and some paired short, thick, spine-shaped bristles over their entire ventral surface. Metatarsus IV also with these paired bristles over its entire ventral surface and two distal spines ventrally.
|
||
</paragraph>
|
||
<paragraph id="6274979AE1405FD92480FBDF5E96648E">
|
||
<emphasis id="D1A7553534B00EED930970B26B1DFDAF" bold="true">
|
||
Female.
|
||
<typeStatus id="112B7239BCB80DE868F23B4534EDAC0C" type="paratype">Paratype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
(
|
||
<abbrev id="ABBRID0EE3DI" xlink_title="Museum of Hebei University">
|
||
<collectionCode id="8BD0D45ECE86C7C91501BC28F6A36B57">MHBU</collectionCode>
|
||
</abbrev>
|
||
-
|
||
<specimenCode id="38629F3DE346557CA3FA6BF7523AFA3C">Sol-XZ 2023073003</specimenCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="DA3E9826DE17E5480AD192CE11DC482E">
|
||
<emphasis id="F98DD07F783762EB6997674623947375" bold="true">
|
||
<emphasis id="F6BC301AFCC55322D595472596C2CCB8" bold="true" italics="true">Measurements</emphasis>
|
||
.
|
||
</emphasis>
|
||
Total body length 21.28,
|
||
<abbrev id="ABBRID0EP3DI" xlink_title="chelicera length">CL</abbrev>
|
||
6.77,
|
||
<abbrev id="ABBRID0ET3DI" xlink_title="chelicera height">CH</abbrev>
|
||
2.53,
|
||
<abbrev id="ABBRID0EX3DI" xlink_title="Propeltidium length">PL</abbrev>
|
||
2.76, PW 4.39,
|
||
<abbrev id="ABBRID0E23DI" xlink_title="sum of the lengths of pedipalp, leg I, and leg IV divided by the sum of the lengths of chelicera and propeltidium, indicating the length of appendages in relation to body size">A / CP</abbrev>
|
||
4.6,
|
||
<abbrev id="ABBRID0E63DI" xlink_title="chelicera length/height, large CL/CH ratios suggest a narrow cheliceral morphology, while a more robust morphology is represented by a smaller ratio">CL / CH</abbrev>
|
||
2.67. Pedipalp 16.16 (4.70, 4.40, 3.63, 1.25), Leg I 9.872 (2.18, 2.89, 1.66, 0.98, 0.19), Leg II 10.49 (1.93, 2.21, 1.80, 0.84, 0.66), Leg III 11.56 (2.04, 2.96, 1.44, 0.43, 0.50), Leg IV 17.90 (4.06, 4.74, 2.06, 0.49, 0.99).
|
||
</paragraph>
|
||
<paragraph id="135AD91A09D92D884BB5BBB81D4BD31C">
|
||
<emphasis id="352D51002BDA9855317FA0039D269BA2" bold="true">
|
||
<emphasis id="987F75BD920C17866CBE311DD6C964BE" bold="true" italics="true">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
In 75 % ethanol-preserved specimens (Fig.
|
||
<figureCitation id="4A8E93C9F5B34FB787CBAA21432F9667" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Habitus A – D K. shigatse sp. nov., habitus, male (A, B) and female (C, D) E – H K. namling sp. nov., habitus, male (E, F) and female (G, H). Scale bars: 10 mm." figureDoi="10.3897/zookeys.1204.120164.figure5" httpUri="https://binary.pensoft.net/fig/1062107">5 G, H</figureCitation>
|
||
). Coloration as in the males.
|
||
</paragraph>
|
||
<paragraph id="3C8AF86C6AC20BA937D5BBEF932EB38D">
|
||
<emphasis id="390655F9D3FCCCE8ACD35B53BC4EE2EC" bold="true">
|
||
<emphasis id="AEFB2610260ED3026AFB9F95D981B43A" bold="true" italics="true">Propeltidium</emphasis>
|
||
.
|
||
</emphasis>
|
||
Much wider than long with a dense pubescence of thin, short, anteriorly directed setae. Anterior, posterior, and lateral edges with several long, curved spiniform setae that perpendicular to the surface of the propeltidium. Ocular tubercle with four middle distal spiniform setae and three middle spiniform setae arranged in a triangle shape (Fig.
|
||
<figureCitation id="8896F2C6556779D56E0942113D1F9817" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Propeltidium A. K. zhui sp. nov., holotype male B. K. zhui sp. nov., paratype female C. K. shigatse sp. nov., holotype male D. K. shigatse sp. nov., paratype female E. K. namling sp. nov., holotype male F K. namling sp. nov., paratype female. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1204.120164.figure7" httpUri="https://binary.pensoft.net/fig/1062109">7 F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="48E46C50A56CCD580C0CAB34B5C58E21">
|
||
<emphasis id="223F661D5280F3B9C00D89F356B900D3" bold="true">
|
||
<emphasis id="57963CA125341AA55E5A20C92573C533" bold="true" italics="true">Chelicerae</emphasis>
|
||
.
|
||
</emphasis>
|
||
Dental formulation of fixed finger:
|
||
<abbrev id="ABBRID0EF5DI" xlink_title="fixed finger, distal tooth">FD</abbrev>
|
||
- (2) -
|
||
<abbrev id="ABBRID0EJ5DI" xlink_title="fixed finger, medial tooth">FM</abbrev>
|
||
- (2) -
|
||
<abbrev id="ABBRID0EN5DI" xlink_title="fixed finger, proximal tooth">FP</abbrev>
|
||
- (6 RF) (5 PF). Dental formulation of movable finger:
|
||
<abbrev id="ABBRID0ER5DI" xlink_title="movable finger, medial tooth">MM</abbrev>
|
||
- (2) -
|
||
<abbrev id="ABBRID0EV5DI" xlink_title="movable finger, proximal tooth">MP</abbrev>
|
||
, with four
|
||
<abbrev id="ABBRID0EZ5DI" xlink_title="movable finger, subterminal tooth/teeth">MST</abbrev>
|
||
and four
|
||
<abbrev id="ABBRID0E45DI" xlink_title="movable finger, subproximal tooth/teeth">MSP</abbrev>
|
||
. Fondal teeth graded as II, III, IV, V, I, tiny VI retrolaterally; I, II, III, IV, tiny V prolaterally (Figs
|
||
<figureCitation id="89A444F8EF8AB93F016BE17EE693DD14" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure10" httpUri="https://binary.pensoft.net/fig/1062112">10 G, H</figureCitation>
|
||
,
|
||
<figureCitation id="ACA5FCA2F2F8F6A898DDCB9AEFBB87BE" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Female fixed finger ventral views A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFSM, retrofondal submedial tooth / teeth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure12" httpUri="https://binary.pensoft.net/fig/1062114">12 F</figureCitation>
|
||
,
|
||
<figureCitation id="E32A12E3954B58B242DA4CDD9F1319EC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Retrolateral (left) and prolateral (right) cheliceral views A, B K. shigatse sp. nov., holotype male C, D K. shigatse sp. nov., paratype female E, F K. namling sp. nov., holotype male G, H K. namling sp. nov., paratype female. Abbreviations: FD, fixed finger, distal tooth; FSD, fixed finger, subdistal tooth / teeth; FM, fixed finger, medial tooth; FSM, fixed finger, submedial tooth / teeth; FP, fixed finger, proximal tooth; RF, retrofondal teeth; RFA, retrofondal apical tooth / teeth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth / teeth; PF, profondal teeth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSP, profondal subproximal tooth; MSP, movable finger, subproximal tooth / teeth; MP, movable finger, proximal tooth; MSM, movable finger, submedial tooth / teeth; MM, movable finger, medial tooth; MST, movable finger, subterminal teeth; pvd, proventral distal setae; fcp (modified pvd), flagellar complex plumose setae; pdp, prodo rsal proximal setae; fcs, flagellar complex subspiniform to spiniform setae; pvsd, proventral subdistal setae; pvd, proventral distal setae. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure15" httpUri="https://binary.pensoft.net/fig/1062117">15 G, H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="F7F7FFF5B214AC22170DD0F76ADB099A">
|
||
<emphasis id="86B50E02D416C8AF6300AF1840B0894A" bold="true">
|
||
<emphasis id="61526A5AFE27CF42F494C9F93E0A1928" bold="true" italics="true">Opisthosoma</emphasis>
|
||
.
|
||
</emphasis>
|
||
The entire surface covered with almost adpressed setae and numerous long, curved, bifurcate setae. Genital operculum triangular in shape with no clear demarcation between the plates., and the rear edge of the genital sternite not chitinized (Fig.
|
||
<figureCitation id="9ADFA29AD27707C702E9A4A910C968F6" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Genital operculum of female A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 1.0 mm." figureDoi="10.3897/zookeys.1204.120164.figure17" httpUri="https://binary.pensoft.net/fig/1062119">17 F</figureCitation>
|
||
). Sternite IV with 14 short needle-like ctenidia extending 1 / 3 the length of the succeeding sternite (Fig.
|
||
<figureCitation id="844DB0AF0A038F2FDEBFD17E661647F3" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Ctenidia on sternite IV of female A K. tibetana B K. dingye sp. nov. C K. lhasa sp. nov. D K. zhui sp. nov. E K. shigatse sp. nov. F K. namling sp. nov. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1204.120164.figure18" httpUri="https://binary.pensoft.net/fig/1062120">18 E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="14B8B7314959212AFD1C0CB09C786E03">
|
||
<emphasis id="9865CC2608CE7C4667097E7F7540371D" bold="true">
|
||
<emphasis id="E765DB4BCEDC6415940B4779803629A7" bold="true" italics="true">Pedipalps</emphasis>
|
||
.
|
||
</emphasis>
|
||
Totally covered with short setae and long, thick setae.
|
||
</paragraph>
|
||
<paragraph id="BDA4CA6E950F4D831353FF4C53192A84">
|
||
<emphasis id="0A4D7FE39C1EF2EA07ACD640AC0D5B50" bold="true">
|
||
<emphasis id="65E550CF7A0DB6593040AE3A5D81E222" bold="true" italics="true">Legs</emphasis>
|
||
.
|
||
</emphasis>
|
||
As in the males.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0ELAAK" type="variability">
|
||
<paragraph id="3C1800F429A008B1A2AA8A3BA784F920">
|
||
<heading id="AB651BAE8F9489BABAB121574F38A592" reason="title">Variability.</heading>
|
||
</paragraph>
|
||
<paragraph id="48B2FBC24C4627D55F9F5A738A980E6D">
|
||
Female. Total length 20.13–23.67. Body coloration pale yellow to yellow. The number of cheliceral fixed finger fondal teeth 10–12 (profondal teeth 4–6; retrofondal teeth 6–7).
|
||
<abbrev id="ABBRID0ERAAK" xlink_title="movable finger, subterminal tooth/teeth">MST</abbrev>
|
||
3–5,
|
||
<abbrev id="ABBRID0EVAAK" xlink_title="movable finger, subproximal tooth/teeth">MSP</abbrev>
|
||
3–4. The number of ctenidia on sternite IV 14–16.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EZAAK" type="distribution">
|
||
<paragraph id="BD54F06491BAEE1A8AEE05A9435FA98B">
|
||
<heading id="5F40A1BC1B75A193F7F829A3D97CD571" reason="title">Distribution and habitat.</heading>
|
||
</paragraph>
|
||
<paragraph id="D8E96D3AA4818901DAC2DABEB0D6D8B8">
|
||
<collectingCountry id="B382E1CABCA77EE0A4D081F0D38CD35D" name="China">China</collectingCountry>
|
||
(
|
||
<collectingRegion id="AC5EA03C30840BC4F6389D3F1EF7CF0C" country="China" name="Xizang">Xizang</collectingRegion>
|
||
). Habitat: grassland (Fig.
|
||
<figureCitation id="53231304932EFA1725953C4D307D86D2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitat where K. dingye sp. nov. (A, B), K. lhasa sp. nov. (C), K. shigatse sp. nov. (E – G) and K. namling sp. nov. (H) have been found. A, B Xizang, Dingye County, Gyangkar Town C Xizang, Lhasa City, Maizhokunggar County D Xizang, Lhasa City, Drepung Monastery E, F Xizang, Nyalam County, Mainqu Town G Xizang, Gyirong County, Zheba Town H Xizang, Namling County, Nubma Town." figureDoi="10.3897/zookeys.1204.120164.figure2" httpUri="https://binary.pensoft.net/fig/1062104">2 H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EDBAK" type="remarks">
|
||
<paragraph id="FC798DD8088FCA9C387443C7C14489A2">
|
||
<heading id="FA7FBA2C508A320F4BA14EB06DEC17C4" reason="title">Remark.</heading>
|
||
</paragraph>
|
||
<paragraph id="721CBEA046FE6F838C0BA7C5A2D53EDB">
|
||
Based on the comparison of genetic distances, with a genetic distance of 0 % (Table
|
||
<tableCitation id="5BAAC7337472A26165012D03035E3883" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Genetic distance among the six species." httpUri="http://table.plazi.org/id/AAF6FF40179CED3530634E534F91C65C" tableUuid="AAF6FF40179CED3530634E534F91C65C">2</tableCitation>
|
||
) between male and female collected from same locations, we believe that they are same species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |