235 lines
35 KiB
XML
235 lines
35 KiB
XML
<document id="87986C993EE49A1E699FE393BC751616" ID-DOI="10.11646/zootaxa.5422.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="10805617" ID-ZooBank="AE955C5E-803E-44CB-A3B2-9C2616D9F185" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710175333099" checkinUser="plazi" docAuthor="Taylor, John F., Loch, James D. & Repetski, John E." docDate="2024" docId="926387DBFF99CA1BFF388398FDF115E0" docLanguage="en" docName="zootaxa.5422.1.1.pdf" docOrigin="Zootaxa 5422 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5422.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.18:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="18" docTitle="Lotagnostus rushtoni Taylor & Loch & Repetski 2024, n. sp." docType="treatment" docVersion="1" lastPageNumber="50" masterDocId="6E5AFFA3FFB7CA2AFFAF8670FF911415" masterDocTitle="Taxonomy and stratigraphic distribution of Lotagnostus (Agnostida: Agnostidae) and associated trilobites and conodonts in the Upper Cambrian (Furongian) of Laurentia" masterLastPageNumber="66" masterPageNumber="1" pageNumber="47" updateTime="1710175939639" updateUser="ExternalLinkService">
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<mods:titleInfo id="FBBEF9FB20CDB576235BB7D6342A6609">
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<mods:title id="2B0594A5AC610BA6A93171832D7F0384">Taxonomy and stratigraphic distribution of Lotagnostus (Agnostida: Agnostidae) and associated trilobites and conodonts in the Upper Cambrian (Furongian) of Laurentia</mods:title>
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<mods:name id="21FD720FE4CEF51D894820A9DC6E86F3" type="personal">
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<mods:role id="E24274B70974FD59813AEB3034B1D04C">
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<mods:roleTerm id="5E605620CBEF6E39E23FB3B29E4E83D2">Author</mods:roleTerm>
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<mods:namePart id="FD1BE1D0AA1EE00B337AD5A0E184FE22">Taylor, John F.</mods:namePart>
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<mods:nameIdentifier id="7A10A77915BD31E44C5521B3C31EA2B4" type="ORCID">0000-0003-0162-0087</mods:nameIdentifier>
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<mods:affiliation id="C9A503D8FA31A458E6CE19DE18D1778F">Department of Geography, Geology, Environment and Planning, Indiana University of Pennsylvania, Indiana, Pennsylvania 15705. jftaylor @ iup. edu; https: // orcid. org / 0000 - 0003 - 0162 - 0087</mods:affiliation>
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<mods:nameIdentifier id="DC18F940F1592C0D03CD5945A19DD03C" type="email">jftaylor@iup.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="DA17D1CC006E94C1DDF269EEBA624147" type="personal">
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<mods:role id="BB4CEEA70648B8C9917107211B1C34D6">
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<mods:roleTerm id="E5A478FC1C9CE41011EAF0C225764019">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="DA58E26B0E02751C81510102F1A9C8A8">Loch, James D.</mods:namePart>
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<mods:nameIdentifier id="D3F74416AAD7F1BB7D552B4432D2A015" type="ORCID">0000-0003-4468-3512</mods:nameIdentifier>
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<mods:affiliation id="9EEFB4AD374B103482E268144040FB32">Department of Physical Sciences, University of Central Missouri, Warrensburg, Missouri, 64093 loch @ ucmo. edu; https: // orcid. org / 0000 - 0003 - 4468 - 3512</mods:affiliation>
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<mods:nameIdentifier id="E45A9F04652DA93F199D57760D5491E9" type="email">loch@ucmo.edu</mods:nameIdentifier>
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</mods:name>
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<mods:name id="951CFC1C4C4C2217138490049E00C39E" type="personal">
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<mods:role id="217714A79C76A161CF0F269E9203536D">
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<mods:roleTerm id="AFBD1775D4724CC7146DF793B55DAECE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="92C36358A17AE072D8D69F9220ADB865">Repetski, John E.</mods:namePart>
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<mods:nameIdentifier id="D722DC56AB122CDAECF3DFCC5862C012" type="ORCID">0000-0003-0162-0087</mods:nameIdentifier>
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<mods:affiliation id="DC90054DF74F6A7D5DDC5E0E0603A87A">Department of Geography, Geology, Environment and Planning, Indiana University of Pennsylvania, Indiana, Pennsylvania 15705. jftaylor @ iup. edu; https: // orcid. org / 0000 - 0003 - 0162 - 0087 & U. S. Geological Survey, MS 926 A, National Center, Reston, Virginia 20192. jrepetski @ usgs. gov; https: // orcid. org / 0000 - 0002 - 2298 - 7120 & Department of Geography, Geology, Environment and Planning, Indiana University of Pennsylvania, Indiana, Pennsylvania 15705. jftaylor @ iup. edu; https: // orcid. org / 0000 - 0003 - 0162 - 0087</mods:affiliation>
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<mods:nameIdentifier id="96C5E061C3CFDE861A3E4EC55AEC39A0" type="email">jftaylor@iup.edu</mods:nameIdentifier>
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<mods:title id="F11BD3DEE6683B0578BF5C2EE6C29960">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="C7F9D54DD9260594DCCEFE312133CD21">
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<mods:date id="B66835AFED5F8C1C6E139EFF155F5A2D">2024</mods:date>
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<mods:number id="2EA2D60309CFE492ED85F7E71CBFEF48">2024-03-11</mods:number>
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<mods:detail id="CB24E615F0AE8855220176249A1D5176" type="volume">
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<mods:number id="2B5C1B6182AB543D4763B2EB2C297F7F">5422</mods:number>
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<mods:number id="F9CA64270AAE4E58F12A1CD848853CE7">1</mods:number>
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<mods:start id="C68233F60121C9BFAEF0E3108CC0CF3B">1</mods:start>
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<mods:location id="A854E37FA85C1D95323446F278244DE7">
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<mods:url id="EC46CF61AE56141176DA9266319669D3">http://dx.doi.org/10.11646/zootaxa.5422.1.1</mods:url>
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</mods:location>
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<mods:classification id="63590F9A4B169F644F1931CBD3C01ED4">journal article</mods:classification>
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<mods:identifier id="869345C58E595194A1F513C6071475E3" type="DOI">10.11646/zootaxa.5422.1.1</mods:identifier>
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<mods:identifier id="351144B6897047B628F6766AF7CF4ED8" type="ISSN">1175-5334</mods:identifier>
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<mods:identifier id="3307EC3E647FFDD84ECE87F656AB1A57" type="Zenodo-Dep">10805617</mods:identifier>
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<mods:identifier id="3AB6A39D46C7E217F75ACD387A2138E8" type="ZooBank">AE955C5E-803E-44CB-A3B2-9C2616D9F185</mods:identifier>
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</mods:mods>
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<treatment id="926387DBFF99CA1BFF388398FDF115E0" LSID="urn:lsid:plazi:treatment:926387DBFF99CA1BFF388398FDF115E0" httpUri="http://treatment.plazi.org/id/926387DBFF99CA1BFF388398FDF115E0" lastPageId="49" lastPageNumber="50" pageId="46" pageNumber="47">
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<subSubSection id="52D06546FF99CA04FF388398FE721217" box="[151,483,1511,1538]" pageId="46" pageNumber="47" type="nomenclature">
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<paragraph id="1A7536CDFF99CA04FF388398FE721217" blockId="46.[151,483,1511,1574]" box="[151,483,1511,1538]" pageId="46" pageNumber="47">
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<heading id="413D81A1FF99CA04FF388398FE721217" bold="true" box="[151,483,1511,1538]" fontSize="11" level="2" pageId="46" pageNumber="47" reason="4">
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<emphasis id="28BEEADFFF99CA04FF388398FE721217" bold="true" box="[151,483,1511,1538]" pageId="46" pageNumber="47">
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<taxonomicName id="DDCA4D4EFF99CA04FF388398FE0B1217" authority="Taylor & Loch & Repetski, 2024" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[151,410,1511,1538]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="rushtoni" status="sp. nov.">
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<emphasis id="28BEEADFFF99CA04FF388398FE0B1217" bold="true" box="[151,410,1511,1538]" italics="true" pageId="46" pageNumber="47">Lotagnostus rushtoni</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="338D57A4FF99CA04FE0E8398FE721217" box="[417,483,1512,1538]" pageId="46" pageNumber="47" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="52D06546FF99CA04FF38807CFE941233" box="[151,261,1548,1574]" pageId="46" pageNumber="47" type="description">
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<paragraph id="1A7536CDFF99CA04FF38807CFE941233" blockId="46.[151,483,1511,1574]" box="[151,261,1548,1574]" pageId="46" pageNumber="47">
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(
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<figureCitation id="82F12A48FF99CA04FF30807CFF6C1233" box="[159,253,1548,1574]" captionStart="PLATE 12" captionStartId="48.[152,238,1775,1799]" captionTargetBox="[151,1435,151,1758]" captionTargetId="figure-29@48.[151,1435,151,1758]" captionTargetPageId="48" captionText="PLATE 12. Lotagnostus rushtoni n. sp. Each white scale bar represents 2mm. 1-3. Cephalon, CM 41383, from 5/22/08D, dorsal, anterior, and left lateral views. 4-6. Cephalon, USMN 775739, from D3383-CO, dorsal, anterior, and right lateral views. 7. Cephalon, USMN 775740, from D3383-CO, dorsal view. 8. Cephalon, USMN 775741, from D3383-CO, dorsal view. 9. Cephalon, USMN 775742, from D3383-CO, dorsal view. 10. Cephalon, CM 41384, from 5/22/08D, dorsal view. 11- 14. Pygidium, holotype, CM 41385, from 5/22/08D, dorsal, posterior, right lateral, and posterior oblique views. 15. Pygidium, USMN 775743, from D3383-CO, dorsal view. 16. Pygidium, USMN 775744, from D3383-CO, dorsal view. 17-19. Pygidium, USMN 775745, from D3383-CO, dorsal, left lateral, and posterior views. 20. Pygidium, CM 41386, from 5/22/08D, dorsal view. 21. Pygidium, USMN 775746, from 3D3381-CO, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.10805662" httpUri="https://zenodo.org/record/10805662/files/figure.png" pageId="46" pageNumber="47">Plate 12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="52D06546FF99CA04FF388024FEC11353" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="1A7536CDFF99CA04FF388024FEC11353" blockId="46.[151,1437,1620,2043]" pageId="46" pageNumber="47">
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<emphasis id="28BEEADFFF99CA04FF388024FE83127B" bold="true" box="[151,274,1620,1646]" pageId="46" pageNumber="47">Diagnosis.</emphasis>
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Non-scrobiculate
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<taxonomicName id="DDCA4D4EFF99CA04FE4C8025FDFD127B" authorityName="Whitehouse" authorityYear="1936" box="[483,620,1621,1646]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">
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<emphasis id="28BEEADFFF99CA04FE4C8025FDFD127B" box="[483,620,1621,1646]" italics="true" pageId="46" pageNumber="47">Lotagnostus</emphasis>
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</taxonomicName>
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with narrow, strongly curved anterior margin; glabella long and ogival; lateral lobes of M3 defined only by inflation with no bounding longitudinal furrows; slightly elongate glabellar node sits high on elliptical elevated platform on anterior half of M2; large, inflated basal lobes with convex lateral margins extend well behind glabellar culmination; pygidium with transaxial F1; axial furrows narrow and moderately impressed; broad (tr.) posteroaxis with intranotular area undefined or extremely faint; acrolobe unconstricted; post-axial pleural field narrow (sag.); border furrow wide, deep, constant in width, with sharp boundaries with pleural field and border.
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</paragraph>
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</subSubSection>
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<subSubSection id="52D06546FF99CA04FF688120FEB9139B" pageId="46" pageNumber="47" type="etymology">
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<paragraph id="1A7536CDFF99CA04FF688120FEB9139B" blockId="46.[151,1437,1620,2043]" pageId="46" pageNumber="47">
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<emphasis id="28BEEADFFF99CA04FF688120FEDF137F" bold="true" box="[199,334,1872,1898]" pageId="46" pageNumber="47">Etymology.</emphasis>
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Named after A.W.A. Rushton for his meticulous work on Cambrian taxa, among them
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<emphasis id="28BEEADFFF99CA04FF388105FEB9139B" box="[151,296,1909,1934]" italics="true" pageId="46" pageNumber="47">
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<taxonomicName id="DDCA4D4EFF99CA04FF388105FEB5139B" authorityName="Whitehouse" authorityYear="1936" box="[151,292,1909,1934]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Lotagnostus</taxonomicName>
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.
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="52D06546FF99CA05FF6881E8FB0D14C1" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
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<paragraph id="1A7536CDFF99CA05FF6881E8FB0D14C1" blockId="46.[151,1437,1620,2043]" lastBlockId="47.[151,1437,150,2036]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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<emphasis id="28BEEADFFF99CA04FF6881E8FE6613A7" bold="true" box="[199,503,1944,1970]" pageId="46" pageNumber="47">Material and occurrence.</emphasis>
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<materialsCitation id="AAA23C90FF99CA05FE5081E8FB0D14C1" collectionCode="CM, USNM" lastPageId="47" lastPageNumber="48" location="Ninemile Canyon" municipality="Windfall Formation" pageId="46" pageNumber="47" specimenCode="CM 41385, CM 41383-41384, CM41386, USNM 775743, 775746, USNM 775739-775742, USNM 775744-775745" specimenCount="1" stateProvince="Nevada" typeStatus="holotype">
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<typeStatus id="C571886FFF99CA04FE5081E8FDF613A7" box="[511,615,1944,1970]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
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<specimenCode id="4A6C9EB6FF99CA04FDC081E8FD7913A6" box="[623,744,1944,1971]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="46" pageNumber="47">CM 41385</specimenCode>
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is a pygidium from collection 5/22/08D: assigned specimens include 2 cephala
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<specimenCode id="4A6C9EB6FF99CA04FEF281CCFDB913C3" box="[349,552,1980,2006]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="46" pageNumber="47">CM 41383–41384</specimenCode>
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and 1 pygidium
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<specimenCode id="4A6C9EB6FF99CA04FD4F81CCFCC013C3" box="[736,849,1980,2006]" collectionCode="CM" country="China" name="Chongqing Museum" pageId="46" pageNumber="47">CM41386</specimenCode>
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from collection 5/22/08D; 2 pygidia
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<specimenCode id="4A6C9EB6FF99CA04FB4481CCFA0913C2" box="[1259,1432,1980,2007]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="46" pageNumber="47" type="Museum">USNM 775743</specimenCode>
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,
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<specimenCode id="4A6C9EB6FF99CA04FF388190FF7A13EE" box="[151,235,2016,2043]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="46" pageNumber="47" type="Museum">775746</specimenCode>
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from collection D3381-CO; and 4 cephala
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<specimenCode id="4A6C9EB6FF99CA04FD798190FC7513EE" box="[726,996,2016,2043]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="46" pageNumber="47" type="Museum">USNM 775739–775742</specimenCode>
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and 2 pygidia
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<specimenCode id="4A6C9EB6FF99CA04FB218190FA0D13EE" box="[1166,1436,2016,2043]" collectionCode="USNM" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="46" pageNumber="47" type="Museum">USNM 775744–775745</specimenCode>
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from collection D3383-CO.
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<taxonomicName id="DDCA4D4EFF98CA05FE7686E7FC8B14A4" authority="Fauna" authorityName="Fauna" box="[473,794,151,177]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="rushtoni">
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<emphasis id="28BEEADFFF98CA05FE7686E7FD5914A4" box="[473,712,151,177]" italics="true" pageId="47" pageNumber="48">Lotagnostus rushtoni</emphasis>
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Fauna
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</taxonomicName>
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: collection 5/22/08C (32-40), 5/22/08D (36-34), D3381- CO (0-3) and D3383-CO (9-18) from the
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<collectingMunicipality id="FA11ACB7FF98CA05FDC586CAFCD914C1" box="[618,840,186,212]" pageId="47" pageNumber="48">Windfall Formation</collectingMunicipality>
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at
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<location id="1F156016FF98CA05FCC586CAFBA514C0" LSID="urn:lsid:plazi:treatment:926387DBFF99CA1BFF388398FDF115E0:1F156016FF98CA05FCC586CAFBA514C0" box="[874,1076,186,213]" municipality="Windfall Formation" name="Ninemile Canyon" pageId="47" pageNumber="48" stateProvince="Nevada">Ninemile Canyon</location>
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,
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<collectingRegion id="D80EF82FFF98CA05FB9086CAFB0914C1" box="[1087,1176,186,212]" country="United States of America" name="Nevada" pageId="47" pageNumber="48">Nevada</collectingRegion>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="52D06546FF98CA05FF6886AEFC501308" pageId="47" pageNumber="48" type="description">
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<paragraph id="1A7536CDFF98CA05FF6886AEFCB417D6" blockId="47.[151,1437,150,2036]" pageId="47" pageNumber="48">
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<emphasis id="28BEEADFFF98CA05FF6886AEFEC814ED" bold="true" box="[199,345,222,248]" pageId="47" pageNumber="48">Description.</emphasis>
|
||
Cephalon semiovate in outline with maximum cephalic width equal to cephalic length (97– 105%); slightly convex (sag.) to moderately convex (tr.). Glabella long (sag.), 76% (71–81%) of cephalic length; narrow, accounting for approximately a third of cephalic width at M3; slightly convex (tr.), rising only slightly above genae in anterior and lateral views. Axial furrows moderately impressed; strongly convergent (30º angle to midline) from posterior margin to anterior end of basal lobes, parallel to very slightly convergent to poorly defined anterolateral corners of glabella, strongly convergent (45º) to front of glabella. Basal lobes long, 32% (26–35%) of glabellar length; basal furrow moderately impressed, but slightly narrower than axial furrows. Very short (sag.), gently curved, posteriorly convex occipital band. F1 a very shallow (tr.) indentation in side of glabella near anterior tip of basal lobe. Glabellar culmination very bluntly pointed with very faint terminal node. M2 equal in width (tr.) or slightly narrower than M3, with rounded anterolateral corners; prominent, slightly elongate glabellar node centered on inflated, elliptical area on posterior half of M2; F2 moderately impressed abaxially, faintly impressed medially in small specimens, incomplete across axis in larger specimens, directly slightly anteromedially. M3 divided transversely into a shorter (sag.), anteriorly tapering, medial lobe flanked by slightly inflated, elongate, subelliptical lateral lobes; boundaries between medial and lateral lobes defined by change in slope rather than furrows. Transglabellar F3 moderately impressed, slightly to moderately concave anteriorly in larger specimens. Ogival anteroglabella constitutes 33% (31–36%) of glabellar length (sag.). Width of genal field, equal to glabellar width (tr.) at F2, steadily decreases anteriorly to where preglabellar field constitutes only 16% (12–18%) of cephalic length at midline; slope angle of genal field increases abaxially from near-horizontal adjacent to axial furrow to vertical at border furrow, causing genal field to overhang border at posterolateral corners. Genal and preglabellar fields nonscrobiculate. Border narrow, convex, narrowing slightly posteriorly. Border furrow well impressed, uniform in depth and width, slightly narrower than border.
|
||
</paragraph>
|
||
<paragraph id="1A7536CDFF98CA05FF6885BCFC501308" blockId="47.[151,1437,150,2036]" pageId="47" pageNumber="48">Pygidium semiovate in outline, pygidial length exclusive of articulating half-ring 91% (85–94%) of maximum width (tr.) at F2; moderately convex (sag., tr.). Axis long, 82% (78–85%) of pygidial length; slightly convex (sag., tr.), standing in slight relief above adjacent pleural fields; post-axial pleural field continues gentle posterior slope of posteroaxis in lateral view. Anteroaxis accounts for less than half (42–46%) of length (sag.) of axis. Axial furrows moderately impressed; slightly convergent from anterior margin to F2, bowed outward along M1, less so along M2; bowed outward along sides of ogival posteroaxis. Articulating half ring short (sag.), crescentic; width (tr.) approximately 70% of M1; articulating furrow deep and broad (sag.), narrowing laterally in front of lateral lobes of M1. M1 wider (tr.) than M2 and posteroaxis, trisected by longitudinal furrows into slightly shorter (sag.) and narrower (tr.) medial lobe with gently abaxially convex sides, and inflated lateral lobes with broadly rounded anterolateral corners. F1 transaxial, bowed forward slightly, moderately impressed, shallowing over midline in some specimens. M2 trisected, longitudinal furrows slightly shallower than on M1; medial lobe narrower (tr.) than medial lobe of M1, longer (sag.) and narrower than lateral lobes of M2; large, strongly asymmetrical axial node on posterior half of M2 extends postero-dorsally to overhang F2. Transaxial F2 moderately impressed, transverse to gently anteriorly convex; wider (exsag.) and curved forward slightly at intersection with axial furrows on some specimens, producing rounded posterolateral corners on lateral lobes of M2; shallower and deflected sightly forward over midline at base of axial node. Posteroaxis slightly wider (tr.) than M2; maximum width slightly greater than length (sag.); some specimens display very faint intranotular axis circumscribed by slight change in convexity and flat, terminating in flat, upturned tip. Pleural fields narrow, widest (tr.) at level of slightly constricted M2, narrowing posteriorly to produce short (sag.) post-axial field; moderately convex (tr., sag.); moderately declined post-axially as seen in lateral view; slightly declined from axial furrows increasing to moderately sloping near border furrow. Non-scrobiculate. Border flatly convex, moderate in width, 8% (7–10%) of pygidial length exclusive of articulating half-ring; widest at midline, tapering toward anterolateral corners. Border furrow broad, well impressed with very steep boundaries with pleural field and border; narrows slightly anterior to short posterolateral spines that lie just anterior to end of axis. Shoulder furrow well impressed, narrow (exsag).</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="52D06546FF98CA1BFF688156FDF115E0" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="discussion">
|
||
<paragraph id="1A7536CDFF98CA1BFF688156FD7E14C0" blockId="47.[151,1437,150,2036]" lastBlockId="49.[151,1437,151,501]" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48">
|
||
<emphasis id="28BEEADFFF98CA05FF688156FEDD1355" bold="true" box="[199,332,1830,1856]" pageId="47" pageNumber="48">Discussion.</emphasis>
|
||
<taxonomicName id="DDCA4D4EFF98CA05FEFB8157FDD21355" box="[340,579,1830,1856]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF98CA05FEFB8157FDD21355" box="[340,579,1830,1856]" italics="true" pageId="47" pageNumber="48">Lotagnostus rushtoni</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from
|
||
<emphasis id="28BEEADFFF98CA05FCED8157FC471355" box="[834,982,1831,1856]" italics="true" pageId="47" pageNumber="48">
|
||
<taxonomicName id="DDCA4D4EFF98CA05FCED8157FC431355" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[834,978,1831,1856]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="morrisoni">L. morrisoni</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
with which it co-occurs in the Windfall Formation, by the gentler convexity and abaxial slope of the proximal genal and pleural fields, longer and more pointed anteroglabella, transaxial pygidial F1, relatively longer and narrower posteroaxis (
|
||
<figureCitation id="82F12A48FF98CA05FBD1811EFA93139D" box="[1150,1282,1902,1928]" captionStart="FIGURE 10" captionStartId="21.[151,250,1735,1759]" captionTargetBox="[155,1434,390,1704]" captionTargetId="figure-107@21.[151,1435,381,1712]" captionTargetPageId="21" captionText="FIGURE 10. Morphometric comparisons of some Lotagnostus species. (1) Dimensions measured on cephala. Abbreviations: AAL, Axial Anteroglabella Length; APL, Axial Posteroglabella Length; EBL, Exsaggital, Basal lobe Length; PFL, Preglabellar Field Length. (2) Scatter plot of EBL/APL ratio versus length of the cephalon for 3 species of Lotagnostus. L. “trisectus” refers to species identified by that name from Sweden in previous studies. (3) Scatter plot of PFL/AAL ratio versus length of the cephalon for 4 species of Lotagnostus. (4) Graphic comparison of posteroaxis length/length of axis as a function of pygidial length (excluding articulating half ring) for 3 species of Lotagnostus." figureDoi="http://doi.org/10.5281/zenodo.10805642" httpUri="https://zenodo.org/record/10805642/files/figure.png" pageId="47" pageNumber="48">Figure 10.4</figureCitation>
|
||
), and absence of notular furrows. All
|
||
<taxonomicName id="DDCA4D4EFF98CA05FE3A81E2FDF413BE" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[405,613,1937,1963]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="clarki">
|
||
<emphasis id="28BEEADFFF98CA05FE3A81E2FDF413BE" box="[405,613,1937,1963]" italics="true" pageId="47" pageNumber="48">Lotagnostus clarki</emphasis>
|
||
</taxonomicName>
|
||
cephala differ from those of
|
||
<taxonomicName id="DDCA4D4EFF98CA05FC0B81E2FB8F13BE" box="[932,1054,1937,1963]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF98CA05FC0B81E2FB8F13BE" box="[932,1054,1937,1963]" italics="true" pageId="47" pageNumber="48">L. rushtoni</emphasis>
|
||
</taxonomicName>
|
||
in exhibiting a longer preglabellar field, less pointed anteroglabella, and a smaller glabellar node not elevated by inflation of the surrounding area. Larger
|
||
<taxonomicName id="DDCA4D4EFF98CA05FF4281AAFEC113E6" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[237,336,2009,2035]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="clarki">
|
||
<emphasis id="28BEEADFFF98CA05FF4281AAFEC113E6" box="[237,336,2009,2035]" italics="true" pageId="47" pageNumber="48">L. clarki</emphasis>
|
||
</taxonomicName>
|
||
cephala are weakly scrobiculate, whereas those of
|
||
<taxonomicName id="DDCA4D4EFF98CA05FC1D81AAFBA013E6" box="[946,1073,2009,2035]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF98CA05FC1D81AAFBA013E6" box="[946,1073,2009,2035]" italics="true" pageId="47" pageNumber="48">L. rushtoni</emphasis>
|
||
</taxonomicName>
|
||
are smooth. The transaxial F1 distinguishes all
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFEFD86E7FE5C14A4" box="[338,461,151,177]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF86CA1BFEFD86E7FE5C14A4" box="[338,461,151,177]" italics="true" pageId="49" pageNumber="50">L. rushtoni</emphasis>
|
||
</taxonomicName>
|
||
pygidia from those of
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFD6586E7FCB814A4" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[714,809,151,177]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="clarki">
|
||
<emphasis id="28BEEADFFF86CA1BFD6586E7FCB814A4" box="[714,809,151,177]" italics="true" pageId="49" pageNumber="50">L. clarki</emphasis>
|
||
</taxonomicName>
|
||
and larger sclerites of the former display a deeper, more steep-sided border furrow and wider posterior border.
|
||
</paragraph>
|
||
<caption id="4EB56645FF87CA1AFF37809FFCDD13EC" ID-DOI="http://doi.org/10.5281/zenodo.10805662" ID-Zenodo-Dep="10805662" httpUri="https://zenodo.org/record/10805662/files/figure.png" pageId="48" pageNumber="49" startId="48.[152,238,1775,1799]" targetBox="[151,1435,151,1758]" targetPageId="48" targetType="plate">
|
||
<paragraph id="1A7536CDFF87CA1AFF37809FFCDD13EC" blockId="48.[151,1437,1775,2041]" pageId="48" pageNumber="49">
|
||
<emphasis id="28BEEADFFF87CA1AFF37809FFE861312" bold="true" box="[152,279,1775,1799]" pageId="48" pageNumber="49">PLATE 12.</emphasis>
|
||
<taxonomicName id="DDCA4D4EFF87CA1AFE8F8080FE681312" authority="Taylor & Loch & Repetski, 2024" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[288,505,1775,1799]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="rushtoni" status="sp. nov.">
|
||
<emphasis id="28BEEADFFF87CA1AFE8F8080FE681312" box="[288,505,1775,1799]" italics="true" pageId="48" pageNumber="49">Lotagnostus rushtoni</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="338D57A4FF87CA1AFDAC809FFDAD1313" box="[515,572,1775,1798]" pageId="48" pageNumber="49" rank="species">n. sp.</taxonomicNameLabel>
|
||
Each white scale bar represents 2mm. 1-3. Cephalon, CM 41383, from 5/22/08D, dorsal, anterior, and left lateral views. 4-6. Cephalon, USMN 775739, from D3383-CO, dorsal, anterior, and right lateral views. 7. Cephalon, USMN 775740, from D3383-CO, dorsal view. 8. Cephalon, USMN 775741, from D3383-CO, dorsal view. 9. Cephalon, USMN 775742, from D3383-CO, dorsal view. 10. Cephalon, CM 41384, from 5/22/08D, dorsal view. 11- 14. Pygidium, holotype, CM 41385, from 5/22/08D, dorsal, posterior, right lateral, and posterior oblique views. 15. Pygidium, USMN 775743, from D3383-CO, dorsal view. 16. Pygidium, USMN 775744, from D3383-CO, dorsal view. 17-19. Pygidium, USMN 775745, from D3383-CO, dorsal, left lateral, and posterior views. 20. Pygidium, CM 41386, from 5/22/08D, dorsal view. 21. Pygidium, USMN 775746, from 3D3381-CO, dorsal view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="1A7536CDFF86CA1BFF6886AFFDF115E0" blockId="49.[151,1437,151,501]" pageId="49" pageNumber="50">
|
||
The transaxial pygidial F1 sets
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFD8986AFFC8214EC" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[550,787,223,249]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF86CA1BFD8986AFFC8214EC" box="[550,787,223,249]" italics="true" pageId="49" pageNumber="50">Lotagnostus rushtoni</emphasis>
|
||
</taxonomicName>
|
||
apart from all other non-scrobiculate species of the genus except
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFF468773FEDE1508" baseAuthorityName="Troedsson" baseAuthorityYear="1937" box="[233,335,259,285]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hedini">
|
||
<emphasis id="28BEEADFFF86CA1BFF468773FEDE1508" box="[233,335,259,285]" italics="true" pageId="49" pageNumber="50">L. hedini</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<emphasis id="28BEEADFFF86CA1BFE258773FE301509" box="[394,417,259,284]" italics="true" pageId="49" pageNumber="50">L.</emphasis>
|
||
aff.
|
||
<emphasis id="28BEEADFFF86CA1BFE778773FDCC1508" box="[472,605,259,285]" italics="true" pageId="49" pageNumber="50">
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFE778773FDC81508" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[472,601,259,285]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rushtoni">L. rushtoni</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The latter species is discussed in more detail below. The cephalon of
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFA298773FF651554" authorityName="Taylor & Loch & Repetski" authorityYear="2024" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF86CA1BFA298773FF651554" italics="true" pageId="49" pageNumber="50">L. rushtoni</emphasis>
|
||
</taxonomicName>
|
||
differs from that of
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFE4C8757FDD91554" baseAuthorityName="Troedsson" baseAuthorityYear="1937" box="[483,584,295,321]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hedini">
|
||
<emphasis id="28BEEADFFF86CA1BFE4C8757FDD91554" box="[483,584,295,321]" italics="true" pageId="49" pageNumber="50">L. hedini</emphasis>
|
||
</taxonomicName>
|
||
in possessing longer and more inflated basal lobes that extend posteriorly beyond the glabellar culmination; the posterior ends of the basal lobes of
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFC4B873BFBD81570" baseAuthorityName="Troedsson" baseAuthorityYear="1937" box="[996,1097,331,357]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hedini">
|
||
<emphasis id="28BEEADFFF86CA1BFC4B873BFBD81570" box="[996,1097,331,357]" italics="true" pageId="49" pageNumber="50">L. hedini</emphasis>
|
||
</taxonomicName>
|
||
lie in line with, or slightly in front of the glabellar culmination.
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFD8B871FFD30159C" authorityName="Taylor & Loch & Repetski" authorityYear="2024" box="[548,673,367,393]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="rushtoni">
|
||
<emphasis id="28BEEADFFF86CA1BFD8B871FFD30159C" box="[548,673,367,393]" italics="true" pageId="49" pageNumber="50">L. rushtoni</emphasis>
|
||
</taxonomicName>
|
||
also has a shorter (sag.) and more tapered glabella. The pygidium differs from that of
|
||
<taxonomicName id="DDCA4D4EFF86CA1BFEC087E3FE4315B8" baseAuthorityName="Troedsson" baseAuthorityYear="1937" box="[367,466,403,429]" class="Trilobita" family="Agnostidae" genus="Lotagnostus" kingdom="Animalia" order="Agnostida" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="hedini">
|
||
<emphasis id="28BEEADFFF86CA1BFEC087E3FE4315B8" box="[367,466,403,429]" italics="true" pageId="49" pageNumber="50">L. hedini</emphasis>
|
||
</taxonomicName>
|
||
in having a proportionally shorter (
|
||
<figureCitation id="82F12A48FF86CA1BFCF687E3FC4D15B8" box="[857,988,403,429]" captionStart="FIGURE 10" captionStartId="21.[151,250,1735,1759]" captionTargetBox="[155,1434,390,1704]" captionTargetId="figure-107@21.[151,1435,381,1712]" captionTargetPageId="21" captionText="FIGURE 10. Morphometric comparisons of some Lotagnostus species. (1) Dimensions measured on cephala. Abbreviations: AAL, Axial Anteroglabella Length; APL, Axial Posteroglabella Length; EBL, Exsaggital, Basal lobe Length; PFL, Preglabellar Field Length. (2) Scatter plot of EBL/APL ratio versus length of the cephalon for 3 species of Lotagnostus. L. “trisectus” refers to species identified by that name from Sweden in previous studies. (3) Scatter plot of PFL/AAL ratio versus length of the cephalon for 4 species of Lotagnostus. (4) Graphic comparison of posteroaxis length/length of axis as a function of pygidial length (excluding articulating half ring) for 3 species of Lotagnostus." figureDoi="http://doi.org/10.5281/zenodo.10805642" httpUri="https://zenodo.org/record/10805642/files/figure.png" pageId="49" pageNumber="50">Figure 10.4</figureCitation>
|
||
) and more broadly rounded posteroaxis, an unconstricted acrolobe, and a deep, symmetrical, trough-like border furrow that is less variable in width, and is narrower than the border posterior to F2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |