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<document id="9523A326B204C4A4E34E594E732EA5DB" ID-CLB-Dataset="23975" ID-DOI="10.1080/00222933.2019.1620893" ID-GBIF-Dataset="42ae9e66-896e-456b-85a1-156825ee9a3e" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1582027338249" checkinUser="valdenar" docAuthor="Osvaldo Villarreal, Ludson Neves de Ázara &amp; Kury, Adriano Brilhante" docDate="2019" docId="921387C61265FFEDDD8CFAFEFD94FB20" docLanguage="en" docName="JNATHIST.53.1516.965989.pdf.imf" docOrigin="Journal of Natural History 53 (15)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Protimesius orcus Osvaldo Villarreal &amp; Ludson Neves de Ázara &amp; Kury 2019, sp. nov." docType="treatment" docVersion="16" lastPageNumber="978" masterDocId="6E2AFFBE1269FFE3DC7EFFFDFF8AFFA3" masterDocTitle="Revalidation of Obidosus Roewerı 1913 and description of two new cave-dwelling species of Protimesius Roewerı 1913 from Brazil (Opiliones: Stygnidae)" masterLastPageNumber="989" masterPageNumber="965" pageNumber="976" updateTime="1698784517371" updateUser="ExternalLinkService">
<mods:mods id="1D2F9C0B4C2BBE545339C12171FBDA54" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C688FBEBD246CC1A421D718245A26B27">Revalidation of Obidosus Roewerı 1913 and description of two new cave-dwelling species of Protimesius Roewerı 1913 from Brazil (Opiliones: Stygnidae)</mods:title>
</mods:titleInfo>
<mods:name id="20F55B04CEFE8B067B78121A855235C4" type="personal">
<mods:role id="2B3890BFD399B77094224325B4F0A2F2">
<mods:roleTerm id="E71DA399149FF912716C011357D3C9A4">Author</mods:roleTerm>
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<mods:namePart id="8801D79C799CB9E9BAA363930F361937">Osvaldo Villarreal</mods:namePart>
</mods:name>
<mods:name id="68820EF1B57B68FB94E8BDF4786C2D00" type="personal">
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<mods:namePart id="5C6C02383F72EE3A7BAB645C4573605B">Ludson Neves de Ázara</mods:namePart>
</mods:name>
<mods:name id="FB0A10B7329781F1313741ECCC707C5D" type="personal">
<mods:role id="798B6312F695381BD716580085FA096E">
<mods:roleTerm id="8B216E6863F5C07607693C6BE6E7D552">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B7A91F516687FF3A878223A7F87D1E0">Kury, Adriano Brilhante</mods:namePart>
</mods:name>
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<mods:title id="7667A11C65E2411887A0A37B93180D4D">Journal of Natural History</mods:title>
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<mods:part id="B234A6FB7AAF9AF5557800CFB3C151E2">
<mods:date id="B926085FDB1D866D48DAF2E8F072DC90">2019</mods:date>
<mods:detail id="12518323265415FDD78D39601F02884A" type="pubDate">
<mods:number id="F88CAB930D35D3A97346AB12600B0164">2019-06-03</mods:number>
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<mods:detail id="ED06D882B6B245A9ABB2790FFADA7096" type="volume">
<mods:number id="30C78B7B0BB800B2B8D3C3A0256B8405">53</mods:number>
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<mods:detail id="B936C006AD77E640E9684B98BBCBB8DC" type="issue">
<mods:number id="DCFD1D7243D0D7896E545E49494ED626">15</mods:number>
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<mods:classification id="19219299D667B15BDEB7AB71DAA1FFD4">journal article</mods:classification>
<mods:identifier id="C3ECEA7717ACDEC602D46B70A8D21E7B" type="CLB-Dataset">23975</mods:identifier>
<mods:identifier id="33D23E8FDF51650454D256520D2C7B30" type="DOI">10.1080/00222933.2019.1620893</mods:identifier>
<mods:identifier id="F60A0505230FAF88A04889CA5AC896B5" type="GBIF-Dataset">42ae9e66-896e-456b-85a1-156825ee9a3e</mods:identifier>
<mods:identifier id="FC4E473B411A6A66EC96F1991B65FAF4" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="3C30A85EF2F646BE23A5233F8833EAE7" type="Zenodo-Dep">3673222</mods:identifier>
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<treatment id="921387C61265FFEDDD8CFAFEFD94FB20" ID-DOI="http://doi.org/10.5281/zenodo.3679637" ID-GBIF-Taxon="162161830" ID-Zenodo-Dep="3679637" LSID="urn:lsid:plazi:treatment:921387C61265FFEDDD8CFAFEFD94FB20" httpUri="http://treatment.plazi.org/id/921387C61265FFEDDD8CFAFEFD94FB20" lastPageId="14" lastPageNumber="978" pageId="12" pageNumber="976">
<subSubSection id="52A0655B1265FFEFDD8CFAFEFCB8FABE" box="[498,818,1283,1309]" pageId="12" pageNumber="976" type="nomenclature">
<paragraph id="1A0536D01265FFEFDD8CFAFEFCB8FABE" blockId="12.[304,1012,1283,1344]" box="[498,818,1283,1309]" pageId="12" pageNumber="976">
<heading id="414D81BC1265FFEFDD8CFAFEFCB8FABE" box="[498,818,1283,1309]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="976" reason="2">
<taxonomicName id="DDBA4D531265FFEFDD8CFAFEFD46FABE" ID-CoL="7YHMX" authority="Osvaldo Villarreal &amp; Ludson Neves de Ázara &amp; Kury, 2019" authorityName="Osvaldo Villarreal &amp; Ludson Neves de Ázara &amp; Kury" authorityYear="2019" box="[498,716,1283,1309]" class="Arachnida" family="Stygnidae" genus="Protimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="12" pageNumber="976" phylum="Arthropoda" rank="species" species="orcus" status="sp. nov.">
<emphasis id="28CEEAC21265FFEFDD8CFAFEFD46FABE" bold="true" box="[498,716,1283,1309]" italics="true" pageId="12" pageNumber="976">Protimesius orcus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33FD57B91265FFEFDEABFAF9FCB8FABE" box="[725,818,1284,1309]" pageId="12" pageNumber="976" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="52A0655B1265FFEFDD4EFADAFC7EFAE3" box="[304,1012,1319,1344]" pageId="12" pageNumber="976" type="description">
<paragraph id="1A0536D01265FFEFDD4EFADAFC7EFAE3" blockId="12.[304,1012,1283,1344]" box="[304,1012,1319,1344]" pageId="12" pageNumber="976">
<heading id="414D81BC1265FFEFDD4EFADAFC7EFAE3" box="[304,1012,1319,1344]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="976" reason="5">
(
<figureCitation id="82812A551265FFEFDD49FADAFE13FAE3" box="[311,409,1319,1344]" captionStart="Figure 2" captionStartId="5.[160,225,848,871]" captionTargetBox="[166,1153,142,816]" captionTargetId="figure@5.[163,1153,142,817]" captionTargetPageId="5" captionText="Figure 2. Protimesius orcus sp. nov.ı holotype (MNRJ 19ı403)ı (ac) habitus: (a) dorsal viewı (b) ventral viewı (c) lateral view; (dıe) patellaı tibia and tarsus of right pedipalp: ectal view (d)ı mesal view (e); (fıg) distal portion of penis (MNRJ 4278): dorsal view (f)ı lateral view (g). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673228" httpUri="https://zenodo.org/record/3673228/files/figure.png" pageId="12" pageNumber="976">Figure 2</figureCitation>
, 3 (b,d), 4 (g
<emphasis id="28CEEAC21265FFEFDE4AFADAFDC8FAE3" box="[564,578,1319,1344]" italics="true" pageId="12" pageNumber="976"></emphasis>
l), 5 (c,d), 6 (a,c), 7 (d
<emphasis id="28CEEAC21265FFEFDF40FADAFCC6FAE3" box="[830,844,1319,1344]" italics="true" pageId="12" pageNumber="976"></emphasis>
f), 8 (d
<emphasis id="28CEEAC21265FFEFDFE4FADAFC22FAE3" box="[922,936,1319,1344]" italics="true" pageId="12" pageNumber="976"></emphasis>
g), 10)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="52A0655B1265FFEFDCDEFA70FC1CF93A" pageId="12" pageNumber="976" type="materials_examined">
<paragraph id="1A0536D01265FFEFDCDEFA70FE92FA05" blockId="12.[160,1157,1421,1689]" box="[160,280,1421,1446]" pageId="12" pageNumber="976">
<emphasis id="28CEEAC21265FFEFDCDEFA70FE92FA05" bold="true" box="[160,280,1421,1446]" italics="true" pageId="12" pageNumber="976">Type data</emphasis>
</paragraph>
<paragraph id="1A0536D01265FFEFDCDEFA4CFC1CF93A" blockId="12.[160,1157,1421,1689]" pageId="12" pageNumber="976">
<materialsCitation id="AAD23C8D1265FFEFDCDEFA4CFC10FA4F" ID-GBIF-Occurrence="2565952393" collectingDate="2014-01" collectionCode="MNRJ" collectorName="Brazili Cearai Santa Quiteriai Caverna" isTailEnumeration="true" location="Pellegatti-Franco" pageId="12" pageNumber="976" specimenCode="MNRJ 9060" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
³
<typeStatus id="C50188721265FFEFDCBBFA4CFEA3FA6A" box="[197,297,1457,1481]" pageId="12" pageNumber="976" type="holotype">holotype</typeStatus>
(
<collectionCode id="7CABAE151265FFEFDD41FA4CFE0AFA6A" box="[319,384,1457,1481]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="976">MNRJ</collectionCode>
19403), Brazil, Ceará, Santa Quitéria, Caverna P-08 [
<geoCoordinate id="7F8E50171265FFEFD870FA4CFB0BFA6A" box="[1038,1153,1457,1481]" pageId="12" pageNumber="976">4.56796°</geoCoordinate>
,
<geoCoordinate id="7F8E50171265FFEFDCDEFA2EFEBBFA4F" box="[160,305,1491,1516]" pageId="12" pageNumber="976">39.763976°</geoCoordinate>
], 13.ii.
<date id="6E0410101265FFEFDD11FA29FE39FA4F" box="[367,435,1492,1516]" pageId="12" pageNumber="976" value="2014-01">
<collectingDate id="7E40E9F81265FFEFDD11FA29FE39FA4F" box="[367,435,1492,1516]" pageId="12" pageNumber="976" value="2014-01">2014,</collectingDate>
</date>
F.
<location id="1F65600B1265FFEFDDA7FA29FD1CFA4F" LSID="urn:lsid:plazi:treatment:921387C61265FFEDDD8CFAFEFD94FB20:1F65600B1265FFEFDDA7FA29FD1CFA4F" box="[473,662,1492,1516]" name="Pellegatti-Franco" pageId="12" pageNumber="976">Pellegatti-Franco</location>
and D.R. Pedroso leg.
</materialsCitation>
<materialsCitation id="AAD23C8D1265FFEFDFDAFA29FEBBF992" ID-GBIF-Occurrence="2565952427" collectionCode="MNRJ" collectorName="Brazili Cearai Santa Quiteriai Caverna" isTailEnumeration="true" location="Pellegatti-Franco" pageId="12" pageNumber="976" specimenCode="MNRJ 9060" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<typeStatus id="C50188721265FFEFDFDAFA29FB9EFA4F" box="[932,1044,1492,1516]" pageId="12" pageNumber="976" type="paratype">Paratypes</typeStatus>
: 3 ³
<specimenCount id="0CBCFD591265FFEFD829FA29FB09FA48" box="[1111,1155,1491,1516]" pageId="12" pageNumber="976" type="female">1 ♀</specimenCount>
(
<specimenCode id="4A1C9EAB1265FFEFDCDBFA0BFEA0F9AD" box="[165,298,1526,1550]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="976">MNRJ 9060</specimenCode>
), same locality as the holotype, 15
<emphasis id="28CEEAC21265FFEFDECCFA0BFD35F9AD" box="[690,703,1526,1550]" italics="true" pageId="12" pageNumber="976"></emphasis>
21.vii.
<date id="6E0410101265FFEFDE87FA0BFCB7F9AD" box="[761,829,1526,1550]" pageId="12" pageNumber="976" value="2014-01">2014,</date>
F.
<location id="1F65600B1265FFEFDF21FA0BFB96F9AD" LSID="urn:lsid:plazi:treatment:921387C61265FFEDDD8CFAFEFD94FB20:1F65600B1265FFEFDF21FA0BFB96F9AD" box="[863,1052,1526,1550]" name="Pellegatti-Franco" pageId="12" pageNumber="976">Pellegatti-Franco</location>
and D.R. Pedroso leg.;
</materialsCitation>
<materialsCitation id="AAD23C8D1265FFEFDD47F9E4FD0BF9D5" ID-GBIF-Occurrence="2565952423" collectionCode="MNRJ" collectorName="Brazili Cearai Santa Quiteriai Caverna" isTailEnumeration="true" location="Pellegatti-Franco" pageId="12" pageNumber="976" specimenCode="MNRJ 9060" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
4 ³
<specimenCount id="0CBCFD591265FFEFDD15F9E4FE1FF992" box="[363,405,1561,1585]" pageId="12" pageNumber="976" type="female">5 ♀</specimenCount>
(
<specimenCode id="4A1C9EAB1265FFEFDDDDF9E4FDA2F992" box="[419,552,1561,1585]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="976">MNRJ 4279</specimenCode>
), same data as the holotype; 1 ³ (
<specimenCode id="4A1C9EAB1265FFEFDFD2F9E4FBBBF992" box="[940,1073,1561,1585]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="976">MNRJ 4278</specimenCode>
),
<collectorName id="B74F53061265FFEFD83CF9E4FDA7F9F7" pageId="12" pageNumber="976">Brazil, Ceará, Santa Quitéria, Caverna</collectorName>
E-16 [4.560115°, 39.757793°], 03
<emphasis id="28CEEAC21265FFEFD879F9C6FB9EF9F0" box="[1031,1044,1595,1619]" italics="true" pageId="12" pageNumber="976"></emphasis>
13.ii.
<date id="6E0410101265FFEFD83EF9C1FB0FF9F7" box="[1088,1157,1596,1620]" pageId="12" pageNumber="976" value="2014-01">2014,</date>
F.
<location id="1F65600B1265FFEFDCC3F9A3FEF0F9D5" LSID="urn:lsid:plazi:treatment:921387C61265FFEDDD8CFAFEFD94FB20:1F65600B1265FFEFDCC3F9A3FEF0F9D5" box="[189,378,1630,1654]" name="Pellegatti-Franco" pageId="12" pageNumber="976">Pellegatti-Franco</location>
and D.R. Pedroso leg.
</materialsCitation>
<materialsCitation id="AAD23C8D1265FFEFDEFEF9A3FC1CF93A" ID-GBIF-Occurrence="2565952397" collectionCode="MNRJ" collectorName="Brazili Cearai Santa Quiteriai Caverna" isTailEnumeration="true" location="Pellegatti-Franco" pageId="12" pageNumber="976" specimenCode="MNRJ 9060" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
; 5 ³
<specimenCount id="0CBCFD591265FFEFDEBBF9A3FD78F9D5" box="[709,754,1630,1654]" pageId="12" pageNumber="976" type="female">1 ♀</specimenCount>
(
<specimenCode id="4A1C9EAB1265FFEFDF7CF9A3FC03F9D5" box="[770,905,1630,1654]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="12" pageNumber="976">MNRJ 9061</specimenCode>
), same locality as the holotype, 15
<emphasis id="28CEEAC21265FFEFDD50F97CFEB1F93A" box="[302,315,1665,1689]" italics="true" pageId="12" pageNumber="976"></emphasis>
21.vii.
<date id="6E0410101265FFEFDD0BF97CFE33F93A" box="[373,441,1665,1689]" pageId="12" pageNumber="976" value="2014-01">2014,</date>
F.
<location id="1F65600B1265FFEFDDA5F97CFD12F93A" LSID="urn:lsid:plazi:treatment:921387C61265FFEDDD8CFAFEFD94FB20:1F65600B1265FFEFDDA5F97CFD12F93A" box="[475,664,1665,1689]" name="Pellegatti-Franco" pageId="12" pageNumber="976">Pellegatti-Franco</location>
and D.R. Pedroso leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="52A0655B1264FFEEDCDEFF73FEEDFEAC" pageId="13" pageNumber="977" type="etymology">
<paragraph id="1A0536D01264FFEEDCDEFF73FE97FF04" blockId="13.[160,1155,142,271]" box="[160,285,142,167]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFF73FE97FF04" bold="true" box="[160,285,142,167]" italics="true" pageId="13" pageNumber="977">Etymology</emphasis>
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFF4FFEEDFEAC" blockId="13.[160,1155,142,271]" pageId="13" pageNumber="977">
From Latin
<emphasis id="28CEEAC21264FFEEDD5EFF4FFED4FF69" box="[288,350,178,202]" italics="true" pageId="13" pageNumber="977">Orcus</emphasis>
, a god of the underworld, punisher of broken oaths in Italic and Roman mythologies. As with the Greek Hades, the name of the god was also used for the underworld itself.
</paragraph>
</subSubSection>
<subSubSection id="52A0655B1264FFEEDCDEFEC1FD9FFE7C" pageId="13" pageNumber="977" type="diagnosis">
<paragraph id="1A0536D01264FFEEDCDEFEC1FE9EFEF6" blockId="13.[160,1155,316,479]" box="[160,276,316,341]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFEC1FE9EFEF6" bold="true" box="[160,276,316,341]" italics="true" pageId="13" pageNumber="977">Diagnosis</emphasis>
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFEA2FD9FFE7C" blockId="13.[160,1155,316,479]" pageId="13" pageNumber="977">
Dorsal scutum
<typeStatus id="C50188721264FFEEDD37FEA2FEF1FED4" box="[329,379,351,375]" pageId="13" pageNumber="977">type</typeStatus>
<emphasis id="28CEEAC21264FFEEDDFCFEA2FE44FED4" box="[386,462,351,375]" italics="true" pageId="13" pageNumber="977">epsilon</emphasis>
, with rectangular edges (
<figureCitation id="82812A551264FFEEDE90FEA2FCC1FED4" box="[750,843,351,375]" captionStart="Figure 3" captionStartId="6.[160,225,1398,1421]" captionTargetBox="[258,1057,142,1366]" captionTargetId="figure@6.[258,1058,142,1367]" captionTargetPageId="6" captionText="Figure 3. Protimesius lucifer sp. nov.ı holotype (MNRJ 2560) (aıc) and P. orcus sp. nov. holotype (MNRJ 19ı403) (bıd). Habitus: (aıb) dorsal viewı (cıd) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673230" httpUri="https://zenodo.org/record/3673230/files/figure.png" pageId="13" pageNumber="977">Figure 3</figureCitation>
(b); 2(a)); mesotergal area III unarmed (
<figureCitation id="82812A551264FFEEDD69FE7FFEFCFE39" box="[279,374,386,410]" captionStart="Figure 3" captionStartId="6.[160,225,1398,1421]" captionTargetBox="[258,1057,142,1366]" captionTargetId="figure@6.[258,1058,142,1367]" captionTargetPageId="6" captionText="Figure 3. Protimesius lucifer sp. nov.ı holotype (MNRJ 2560) (aıc) and P. orcus sp. nov. holotype (MNRJ 19ı403) (bıd). Habitus: (aıb) dorsal viewı (cıd) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673230" httpUri="https://zenodo.org/record/3673230/files/figure.png" pageId="13" pageNumber="977">Figure 3</figureCitation>
(b,d); 2(a,c)); pedipalpal tibia smooth (
<figureCitation id="82812A551264FFEEDF53FE7FFC01FE39" box="[813,907,386,410]" captionStart="Figure 5" captionStartId="8.[160,225,855,878]" captionTargetBox="[164,1152,143,822]" captionTargetId="figure@8.[163,1153,142,823]" captionTargetPageId="8" captionText="Figure 5. Protimesius lucifer sp. nov. male paratype (MNRJ 9251)ı (aıb) tibia and tarsus of right pedipalp: (a) ectal viewı (b) mesal view. Protimesius orcus sp. nov. male paratype (MNRJ 4279)ı (cıd) tibia and tarsus of right pedipalp: (c) ectal viewı (d) mesal view. Scale bars (aıb) = 1 mmı (c) = 200 µmı (d) = 500 µm." figureDoi="http://doi.org/10.5281/zenodo.3673234" httpUri="https://zenodo.org/record/3673234/files/figure.png" pageId="13" pageNumber="977">Figure 5</figureCitation>
(c,d)) (vs coarse granulation in
<taxonomicName id="DDBA4D531264FFEEDD6FFE59FEFDFE1F" authorityName="Roeweri" authorityYear="1913" box="[273,375,420,444]" class="Arachnida" family="Stygnidae" genus="Protimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="977" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="28CEEAC21264FFEEDD6FFE59FEFDFE1F" box="[273,375,420,444]" italics="true" pageId="13" pageNumber="977">P. gracilis</emphasis>
</taxonomicName>
or scarce and low dorsal granulation in
<taxonomicName id="DDBA4D531264FFEEDF49FE59FC19FE1F" authority="Osvaldo Villarreal &amp; Ludson Neves de Ázara &amp; Kury, 2019" authorityName="Osvaldo Villarreal &amp; Ludson Neves de Ázara &amp; Kury" authorityYear="2019" box="[823,915,420,444]" class="Arachnida" family="Stygnidae" genus="Protimesius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="13" pageNumber="977" phylum="Arthropoda" rank="species" species="lucifer" status="sp. nov.">
<emphasis id="28CEEAC21264FFEEDF49FE59FC19FE1F" box="[823,915,420,444]" italics="true" pageId="13" pageNumber="977">P. lucifer</emphasis>
</taxonomicName>
<taxonomicNameLabel id="33FD57B91264FFEEDFE7FE59FC78FE1F" box="[921,1010,420,444]" pageId="13" pageNumber="977" rank="species">sp. nov.</taxonomicNameLabel>
); colouration uniformly orange (
<figureCitation id="82812A551264FFEEDD0AFE3AFE58FE7C" box="[372,466,455,479]" captionStart="Figure 2" captionStartId="5.[160,225,848,871]" captionTargetBox="[166,1153,142,816]" captionTargetId="figure@5.[163,1153,142,817]" captionTargetPageId="5" captionText="Figure 2. Protimesius orcus sp. nov.ı holotype (MNRJ 19ı403)ı (ac) habitus: (a) dorsal viewı (b) ventral viewı (c) lateral view; (dıe) patellaı tibia and tarsus of right pedipalp: ectal view (d)ı mesal view (e); (fıg) distal portion of penis (MNRJ 4278): dorsal view (f)ı lateral view (g). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673228" httpUri="https://zenodo.org/record/3673228/files/figure.png" pageId="13" pageNumber="977">Figure 2</figureCitation>
(a
<emphasis id="28CEEAC21264FFEEDD98FE3AFE79FE7C" box="[486,499,455,479]" italics="true" pageId="13" pageNumber="977"></emphasis>
c)).
</paragraph>
</subSubSection>
<subSubSection id="52A0655B1264FFEDDCDEFDF1FDE4FBB6" lastPageId="14" lastPageNumber="978" pageId="13" pageNumber="977" type="description">
<paragraph id="1A0536D01264FFEEDCDEFDF1FEACFD86" blockId="13.[160,1156,524,652]" box="[160,294,524,549]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFDF1FEACFD86" bold="true" box="[160,294,524,549]" italics="true" pageId="13" pageNumber="977">Description</emphasis>
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFDD2FC12FD2F" blockId="13.[160,1156,524,652]" pageId="13" pageNumber="977">
Measurements of male
<typeStatus id="C50188721264FFEEDDD6FDD2FD86FDE4" box="[424,524,559,583]" pageId="13" pageNumber="977" type="holotype">holotype</typeStatus>
(MNRJ 19403). DSL = 3.4; DSW = 3.0; ID = 1.8; pedipalp: coxa = 1.0, trochanter = 0.6, femur = 3.3, patella = 2.9, tibia = 2.1, tarsus = 1.5, total = 10.0; leg IV: femur = 4.7, tibia = 3.4, metatarsus = 5.6, total = 17.9.
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFD44FD02FC01" blockId="13.[160,1156,696,930]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFD44FE77FD71" bold="true" box="[160,509,696,722]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFD44FF77FD71" bold="true" box="[160,253,697,722]" italics="true" pageId="13" pageNumber="977">Dorsum</emphasis>
(
<figureCitation id="82812A551264FFEEDD6EFD45FEFCFD71" box="[272,374,696,722]" captionStart="Figure 3" captionStartId="6.[160,225,1398,1421]" captionTargetBox="[258,1057,142,1366]" captionTargetId="figure@6.[258,1058,142,1367]" captionTargetPageId="6" captionText="Figure 3. Protimesius lucifer sp. nov.ı holotype (MNRJ 2560) (aıc) and P. orcus sp. nov. holotype (MNRJ 19ı403) (bıd). Habitus: (aıb) dorsal viewı (cıd) lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673230" httpUri="https://zenodo.org/record/3673230/files/figure.png" pageId="13" pageNumber="977">Figure 3</figureCitation>
(bıd); (6a)).
</emphasis>
Anterior margin of prosoma smooth, with low anterior eminence. Interocular region and eye mound smooth. Area I divided, smooth. Lateral margin almost smooth, only with a longitudinal row of minute granules. Area I divided, with one small granule on each side; II with one small granule on each side; III with a pair of small paramedian tubercle and with a row of three small granules on each side. Areas I
<emphasis id="28CEEAC21264FFEEDC8EFC9AFF77FCDC" box="[240,253,871,895]" italics="true" pageId="13" pageNumber="977"></emphasis>
III straight. Posterior margin slightly convex with row of small granules. Free tergites I
<emphasis id="28CEEAC21264FFEEDD7AFC74FE9BFC02" box="[260,273,905,929]" italics="true" pageId="13" pageNumber="977"></emphasis>
III with row of medium granules.
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFC33FE39FBA9" blockId="13.[160,1155,973,1034]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFC33FE1CFC44" bold="true" box="[160,406,973,999]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFC33FF67FC44" bold="true" box="[160,237,974,999]" italics="true" pageId="13" pageNumber="977">Venter</emphasis>
(
<figureCitation id="82812A551264FFEEDC81FC30FEEFFC44" box="[255,357,973,999]" captionStart="Figure 2" captionStartId="5.[160,225,848,871]" captionTargetBox="[166,1153,142,816]" captionTargetId="figure@5.[163,1153,142,817]" captionTargetPageId="5" captionText="Figure 2. Protimesius orcus sp. nov.ı holotype (MNRJ 19ı403)ı (ac) habitus: (a) dorsal viewı (b) ventral viewı (c) lateral view; (dıe) patellaı tibia and tarsus of right pedipalp: ectal view (d)ı mesal view (e); (fıg) distal portion of penis (MNRJ 4278): dorsal view (f)ı lateral view (g). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673228" httpUri="https://zenodo.org/record/3673228/files/figure.png" pageId="13" pageNumber="977">Figure 2</figureCitation>
(b)).
</emphasis>
Genital operculum with three small granules. Free sternites with a row of small granules.
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFBCBFBF7FBD1" blockId="13.[160,1155,1077,1138]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFBCBFE57FBEC" bold="true" box="[160,477,1077,1103]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFBCBFE90FBEC" bold="true" box="[160,282,1078,1103]" italics="true" pageId="13" pageNumber="977">Chelicerae</emphasis>
(
<figureCitation id="82812A551264FFEEDD53FBC8FE1EFBEC" box="[301,404,1077,1103]" captionStart="Figure 2" captionStartId="5.[160,225,848,871]" captionTargetBox="[166,1153,142,816]" captionTargetId="figure@5.[163,1153,142,817]" captionTargetPageId="5" captionText="Figure 2. Protimesius orcus sp. nov.ı holotype (MNRJ 19ı403)ı (ac) habitus: (a) dorsal viewı (b) ventral viewı (c) lateral view; (dıe) patellaı tibia and tarsus of right pedipalp: ectal view (d)ı mesal view (e); (fıg) distal portion of penis (MNRJ 4278): dorsal view (f)ı lateral view (g). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673228" httpUri="https://zenodo.org/record/3673228/files/figure.png" pageId="13" pageNumber="977">Figure 2</figureCitation>
(ac)).
</emphasis>
Swollen. Segments I
<emphasis id="28CEEAC21264FFEEDEAAFBCAFD6BFBEC" box="[724,737,1079,1103]" italics="true" pageId="13" pageNumber="977"></emphasis>
II smooth. Mobile finger with a high basal tooth and with three small distal teeth; fixed finger with three small distal teeth.
</paragraph>
<paragraph id="1A0536D01264FFEEDCDEFB63FBCCFAE1" blockId="13.[160,1156,1181,1346]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFB63FDB7FB14" bold="true" box="[160,573,1181,1207]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFB63FE98FB14" bold="true" box="[160,274,1182,1207]" italics="true" pageId="13" pageNumber="977">Pedipalps</emphasis>
(
<figureCitation id="82812A551264FFEEDD56FB60FE1BFB14" box="[296,401,1181,1207]" captionStart="Figure 2" captionStartId="5.[160,225,848,871]" captionTargetBox="[166,1153,142,816]" captionTargetId="figure@5.[163,1153,142,817]" captionTargetPageId="5" captionText="Figure 2. Protimesius orcus sp. nov.ı holotype (MNRJ 19ı403)ı (ac) habitus: (a) dorsal viewı (b) ventral viewı (c) lateral view; (dıe) patellaı tibia and tarsus of right pedipalp: ectal view (d)ı mesal view (e); (fıg) distal portion of penis (MNRJ 4278): dorsal view (f)ı lateral view (g). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673228" httpUri="https://zenodo.org/record/3673228/files/figure.png" pageId="13" pageNumber="977">Figure 2</figureCitation>
(de); 5(cd)).
</emphasis>
Coxa with three basal ventral medium tubercles. Trochanter with one basal and one medial ventral medium tubercle. Femur with a ventral distal small tubercle. Patella generally smooth. Tibia dorsally smooth, ventrally with a row of ectal medium tubercles; tibia mesal IIiIi, ectal IIiIi. Tarsus dorsally smooth, but with a ventral row of tubercles in the ectal and mesal portion, tarsus mesal IiIii, ectal iIiii.
</paragraph>
<paragraph id="1A0536D01264FFEDDCDEFA93FC2EFED5" blockId="13.[160,1156,1389,1727]" lastBlockId="14.[160,1156,142,374]" lastPageId="14" lastPageNumber="978" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFA93FE57FA24" bold="true" box="[160,477,1389,1415]" pageId="13" pageNumber="977">
<emphasis id="28CEEAC21264FFEEDCDEFA93FF5FFA24" bold="true" box="[160,213,1390,1415]" italics="true" pageId="13" pageNumber="977">Legs</emphasis>
(
<figureCitation id="82812A551264FFEEDC9BFA90FEC3FA24" box="[229,329,1389,1415]" captionStart="Figure 4" captionStartId="7.[160,225,870,893]" captionTargetBox="[165,1153,142,838]" captionTargetId="figure@7.[163,1153,142,838]" captionTargetPageId="7" captionText="Figure 4. Protimesius lucifer sp. nov.ı (af) and P. orcus sp. nov.ı (gl). (aıg) Femur IVı dorsal view; (bıh) femur IVı ventral view; (cıi) patella and tibia IVı dorsal view; (dıj) patella and tibia IVı ventral view; (eık) male basitarsus Iı lateral view; (fıl) female basitarsus Iı lateral view. Vouchers: (ae) holotype (MNRJ 2560); (f) female paratype (CNHUFPI 2177); (gk) holotype (MNRJ 19ı403); (l) female paratype (MNRJ 9060). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673232" httpUri="https://zenodo.org/record/3673232/files/figure.png" pageId="13" pageNumber="977">Figure 4</figureCitation>
(gl); 6(aıc)).
</emphasis>
Coxa I with a row of five medium tubercles; II with a row of small tubercles; III
<emphasis id="28CEEAC21264FFEEDD13FA6FFEF0FA09" box="[365,378,1426,1450]" italics="true" pageId="13" pageNumber="977"></emphasis>
IV with scattered small granules. Trochanter I
<emphasis id="28CEEAC21264FFEEDFFCFA6FFC05FA09" box="[898,911,1426,1450]" italics="true" pageId="13" pageNumber="977"></emphasis>
II with one basal and two apical ventral medium tubercles, one retrolateral medium tubercle; III with five ventral medium tubercles and one retrolateral medium tubercle; IV with a basal ventral row of three medium tubercles, two subdistal ventral medium tubercle, one retrolateral and one prolateral tubercle (this larger), besides five medium dorsal tubercles. Femur I
<emphasis id="28CEEAC21264FFEED808F9E1FB09F997" box="[1142,1155,1564,1588]" italics="true" pageId="13" pageNumber="977"></emphasis>
III with retroventral and proventral rows with small tubercles, others rows with minute granules; III with retroventral and proventral rows with medium tubercles, other rows with small granules, one retrodorsal and one prodorsal apical medium large tubercles; IV with retroventral and proventral rows with high tubercles, others rows with medium tubercles, one retrodorsal and one prodorsal apical large tubercle. Patella I
<emphasis id="28CEEAC21267FFEDDF95FF73FC72FF05" box="[1003,1016,142,166]" italics="true" pageId="14" pageNumber="978"></emphasis>
II smooth, III with scattered small granules and ending with two medium dorsal large tubercles; IV two ventral rows of medium tubercles and ending with two high dorsal large tubercles. Tibia I
<emphasis id="28CEEAC21267FFEDDC96FF0BFF7FFEAD" box="[232,245,246,270]" italics="true" pageId="14" pageNumber="978"></emphasis>
II smooth; III with retroventral and proventral rows of small granules; IV with retroventral and proventral rows of medium tubercles increasing apically. Metatarsus I
<emphasis id="28CEEAC21267FFEDD808FEE4FB09FE92" box="[1142,1155,281,305]" italics="true" pageId="14" pageNumber="978"></emphasis>
IV smooth. Distitarsus not inflated. Tarsal process reduced, with a terminal long set. Leg III
<emphasis id="28CEEAC21267FFEDDCCDFEA3FF4AFED5" box="[179,192,350,374]" italics="true" pageId="14" pageNumber="978"></emphasis>
IV without tarsal scopula. Tarsal segmentation: 8(3)/19
<emphasis id="28CEEAC21267FFEDDF54FEA3FCBDFED5" box="[810,823,350,374]" italics="true" pageId="14" pageNumber="978"></emphasis>
20(3)/6/7.
</paragraph>
<paragraph id="1A0536D01267FFEDDCDEFE54FEAAFD17" blockId="14.[160,1156,424,692]" pageId="14" pageNumber="978">
<emphasis id="28CEEAC21267FFEDDCDEFE54FE76FE61" bold="true" box="[160,508,424,450]" pageId="14" pageNumber="978">
<emphasis id="28CEEAC21267FFEDDCDEFE54FF54FE61" bold="true" box="[160,222,425,450]" italics="true" pageId="14" pageNumber="978">Penis</emphasis>
(
<figureCitation id="82812A551267FFEDDC8BFE55FED5FE61" box="[245,351,424,450]" captionStart="Figure 2" captionStartId="5.[160,225,848,871]" captionTargetBox="[166,1153,142,816]" captionTargetId="figure@5.[163,1153,142,817]" captionTargetPageId="5" captionText="Figure 2. Protimesius orcus sp. nov.ı holotype (MNRJ 19ı403)ı (ac) habitus: (a) dorsal viewı (b) ventral viewı (c) lateral view; (dıe) patellaı tibia and tarsus of right pedipalp: ectal view (d)ı mesal view (e); (fıg) distal portion of penis (MNRJ 4278): dorsal view (f)ı lateral view (g). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3673228" httpUri="https://zenodo.org/record/3673228/files/figure.png" pageId="14" pageNumber="978">Figure 2</figureCitation>
(fıg); 7(df)).
</emphasis>
LP trapezoidal, with lateral sides concave, and distal border with triangular ventral tongue; LP clearly differentiated from the truncus/malleus by a ventral crack. Stylus curved, with large triangular dorsal process. MS A1
<emphasis id="28CEEAC21267FFEDD87AFE12FB9BFDA4" box="[1028,1041,495,519]" italics="true" pageId="14" pageNumber="978"></emphasis>
2 straight, positioned dorsolaterally, slightly below half of the LP; MS B absent; MS C1
<emphasis id="28CEEAC21267FFEDDF9BFDECFC78FD8A" box="[997,1010,529,553]" italics="true" pageId="14" pageNumber="978"></emphasis>
3 being born close to the dorsomedial region, large and apically curved; MS D1 absent, MS D2 straight and large, dorsomedially located, at level of MS A. Microsetae campus not restricted to lateral regions of the LP, invading slightly the medial zone, with a small gap between both fields.
</paragraph>
<paragraph id="1A0536D01267FFEDDCDEFD1BFE0AFCE6" blockId="14.[160,1157,742,1045]" pageId="14" pageNumber="978">
<emphasis id="28CEEAC21267FFEDDCDEFD1BFEB9FD5C" bold="true" box="[160,307,742,767]" italics="true" pageId="14" pageNumber="978">Colouration.</emphasis>
Body and appendages in alcohol Vivid Yellow (82) and Vivid Orange Yellow (66) in vivo, chelicera reticulated, legs I
<emphasis id="28CEEAC21267FFEDDEA0FCF7FD61FC81" box="[734,747,778,802]" italics="true" pageId="14" pageNumber="978"></emphasis>
III in alcohol and in vivo Brilliant Orange Yellow (76).
</paragraph>
<paragraph id="1A0536D01267FFEDDCBEFCB3FDE4FBB6" blockId="14.[160,1157,742,1045]" pageId="14" pageNumber="978">
<emphasis id="28CEEAC21267FFEDDCBEFCB3FDADFCC4" bold="true" box="[192,551,846,871]" italics="true" pageId="14" pageNumber="978">
Female
<typeStatus id="C50188721267FFEDDD60FCB3FE0DFCC4" box="[286,391,846,871]" pageId="14" pageNumber="978" type="paratype">paratype</typeStatus>
(MNRJ 9060)
</emphasis>
: Dorsal scutum length = 2.9; dorsal scutum width = 2.6; interocular distance = 1.3; pedipalp: coxa = 0.8, trochanter = 0.4, femur = 4.1, patella = 3.1, tibia = 1.0, tarsus = 1.6, total = 9.9; leg IV: femur = 5.5, tibia = 3.7, metatarsus = 6.6, total = 20.3. Similar to male, except for: chelicera not swollen, legs finely granular. Pedipalpal tibia mesal IIiiIi, ectal IIiIi; tarsus mesal IiIii, ectal iIiii (this variation could be intraspecific and no sexual dimorphism).
</paragraph>
</subSubSection>
<subSubSection id="52A0655B1267FFEDDCDEFBBAFD94FB20" box="[160,542,1095,1155]" pageId="14" pageNumber="978" type="distribution">
<paragraph id="1A0536D01267FFEDDCDEFBBAFEA6FBC3" blockId="14.[160,542,1095,1155]" box="[160,300,1095,1120]" pageId="14" pageNumber="978">
<emphasis id="28CEEAC21267FFEDDCDEFBBAFEA6FBC3" bold="true" box="[160,300,1095,1120]" italics="true" pageId="14" pageNumber="978">Distribution</emphasis>
</paragraph>
<paragraph id="1A0536D01267FFEDDCDEFB96FD94FB20" blockId="14.[160,542,1095,1155]" box="[160,542,1131,1155]" pageId="14" pageNumber="978">
Brazil,
<collectingRegion id="D87EF8321267FFEDDC94FB96FEE1FB20" box="[234,363,1131,1155]" country="Brazil" name="Ceara" pageId="14" pageNumber="978">Ceará state</collectingRegion>
: Santa Quitéria.
</paragraph>
</subSubSection>
</treatment>
</document>