170 lines
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170 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.403.7145" ID-GBIF-Dataset="a6fda06f-575a-4ae4-8d40-ae927a01f959" ID-PMC="PMC4023238" ID-Pensoft-Pub="1313-2970-403-15" ID-PubMed="24843265" ID-ZBK="328E01EFBF324352AD7DBE989A3D716B" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-403-15" ModsDocOrigin="ZooKeys 403" ModsDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" checkinTime="1451245966222" checkinUser="pensoft" docAuthor="Yoshida, Takahiro & Hirowatari, Toshiya" docDate="2014" docId="497581866A8D1AB91E01FDEF89691858" docLanguage="en" docName="ZooKeys 403: 15-45" docOrigin="ZooKeys 403" docSource="http://dx.doi.org/10.3897/zookeys.403.7145" docTitle="Psammoecus boreas Yoshida & Hirowatari, 2014, sp. n." docType="treatment" docUuid="1E58DE65-FC6D-4E40-B55E-A952B311A454" docUuidSource="ZooBank" docVersion="4" lastPageNumber="31" masterDocId="F8057B36FF80C436875CFFBFFFE6FF8F" masterDocTitle="A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)" masterLastPageNumber="45" masterPageNumber="15" pageNumber="30" updateTime="1668158244412" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of Japanese species of the genus Psammoecus Latreille (Coleoptera, Silvanidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yoshida, Takahiro</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hirowatari, Toshiya</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2014</mods:date>
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<mods:detail type="volume">
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<mods:number>403</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>15</mods:start>
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<mods:end>45</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.403.7145</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.403.7145</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-403-15</mods:identifier>
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<mods:identifier type="ZBK">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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<mods:identifier type="ZooBank">328E01EFBF324352AD7DBE989A3D716B</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152052619" LSID="urn:lsid:zoobank.org:act:1E58DE65-FC6D-4E40-B55E-A952B311A454" httpUri="http://treatment.plazi.org/id/497581866A8D1AB91E01FDEF89691858" lastPageId="16" lastPageNumber="31" pageId="15" pageNumber="30">
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<subSubSection pageId="15" pageNumber="30" type="nomenclature">
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<paragraph pageId="15" pageNumber="30">
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<taxonomicName LSID="http://zoobank.org/1E58DE65-FC6D-4E40-B55E-A952B311A454" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus boreas" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="boreas">
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<pageBreakToken pageId="15" pageNumber="30" start="start">Psammoecus</pageBreakToken>
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boreas
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</taxonomicName>
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<taxonomicNameLabel pageId="15" pageNumber="30">sp. n.</taxonomicNameLabel>
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Figs 2A, 8and 14
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<normalizedToken originalValue="D–F">D-F</normalizedToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="30" type="reference_group">
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<paragraph pageId="15" pageNumber="30">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus</taxonomicName>
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:
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<bibRefCitation author="Nakane, T" editor="Nakane, T" journalOrPublisher="Hokuryukan, Tokyo" pageId="27" pageNumber="42" pagination="195 - 196" title="Iconographia Insectorum Japonicorum Colore naturali edita" year="1963">Nakane 1963</bibRefCitation>
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: 195, fig. 16 in pl. 98.
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</paragraph>
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<paragraph pageId="15" pageNumber="30">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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sp. 3,
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano 2009</bibRefCitation>
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: 63, 66, 67, fig. 8. -
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano 2010</bibRefCitation>
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: 12, 16.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="30" type="diagnosis">
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<paragraph pageId="15" pageNumber="30">Diagnosis.</paragraph>
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<paragraph pageId="15" pageNumber="30">
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This species is similar to
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
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and other species closely similar to
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus trimaculatus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="trimaculatus">Psammoecus trimaculatus</taxonomicName>
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.
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<bibRefCitation author="Nakane, T" editor="Nakane, T" journalOrPublisher="Hokuryukan, Tokyo" pageId="27" pageNumber="42" pagination="195 - 196" title="Iconographia Insectorum Japonicorum Colore naturali edita" year="1963">Nakane (1963)</bibRefCitation>
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provided a figure of this species as
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus triguttatus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="triguttatus">Psammoecus triguttatus</taxonomicName>
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. It differs from the aforementioned species by the shorter lateral teeth of the pronotum. It is also very similar to
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus harmandi" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="harmandi">Psammoecus harmandi</taxonomicName>
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Grouvelle, 1912 both in external characters and male genital structure as illustrated by Pal (1980), but can be distinguished from it by the longer antennae and the comparatively oblong 10th antennomere.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="16" lastPageNumber="31" pageId="15" pageNumber="30" type="description">
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<paragraph pageId="15" pageNumber="30">Description.</paragraph>
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<paragraph pageId="15" pageNumber="30">Body length. 2.74-3.27 mm (n=19).</paragraph>
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<paragraph pageId="15" pageNumber="30">Coloration (Fig. 2A). Head and pronotum yellowish-brown. Elytra yellowish-brown with dark brown maculae at half, oval ones at center of each elytron and dark oblique bands toward posterior portion, sometimes connected at elytral suture, forming a V-shaped band. Elytra of lighter color specimens with reduced maculae, oval ones and bands separated. Antennae yellowish-brown basally, posterior ends of 8th to 10th antennomeres darkened, or all antennomeres yellowish-brown in lighter colored specimens.</paragraph>
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<caption pageId="15" pageNumber="30">
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<paragraph pageId="15" pageNumber="30">
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Figures 2. Habitus of
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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spp. A
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus boreas" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="boreas">Psammoecus boreas</taxonomicName>
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sp. n., holotype B
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus omotoensis" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="omotoensis">Psammoecus omotoensis</taxonomicName>
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sp. n., holotype C
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus simoni" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="simoni">Psammoecus simoni</taxonomicName>
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Grouvelle, 1892 D
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus fasciatus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="fasciatus">Psammoecus fasciatus</taxonomicName>
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Reitter, 1874, lectotype E
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus hiranoi" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="hiranoi">Psammoecus hiranoi</taxonomicName>
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Yoshida & Hirowatari, 2013 F
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus quadrimaculatus" order="Coleoptera" pageId="15" pageNumber="30" phylum="Arthropoda" rank="species" species="quadrimaculatus">Psammoecus quadrimaculatus</taxonomicName>
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Reitter, 1874, holotype. Scale: 1.0 mm.
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</paragraph>
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</caption>
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<paragraph pageId="15" pageNumber="30">Head (Fig. 8A, B, C). Broad, HW/HL 1.35-1.65; IE/HL 0.96-1.13. Temples narrowed around each base. Eyes small, moderately rounded. Dorsal surface with moderately dense punctuation, ventral surface punctuated sparsely. Antennae 1.54-1.71 mm, thin, very long; covered with considerable long semi-erect pubescence on each antennomere; approximate ratio of holotype as follows: 2.6: 1.0: 1.1: 1.2: 1.2: 1.3: 1.2: 1.0: 1.0: 1.1: 1.7 (Fig. 8A).</paragraph>
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<paragraph pageId="15" pageNumber="30">Pronotum (Fig. 8B, C). Subquadrate, PW/PL 1.18-1.30. Dorsolateral portions lightly impressed. Dorsal surface with relatively dense punctuation, no punctures around posterior margin, comparatively sparse punctuation on ventral surface. Pubescence composed of many short setae and fine long setae on teeth on lateral margins and anterior and posterior angles. Each anterior angle with a distinct group of a few very small teeth, each lateral margin with four small teeth of almost the same size, each posterior angle with a small tooth, almost the same size as those on lateral margins.</paragraph>
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<paragraph pageId="15" pageNumber="30">Elytra (Fig. 8E). Oval, EW/BL 0.32-0.45. Rows of punctures wider than interstices. Pubescence composed of many short setae, no long setae.</paragraph>
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<paragraph pageId="15" pageNumber="30">9th abdominal sternite (Fig. 14D). Strut Y-shaped, cut deeply at anterior 1/5, diverging for posterior 1/4. Lateral sclerites elongate.</paragraph>
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<paragraph lastPageId="16" lastPageNumber="31" pageId="15" pageNumber="30">
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Aedeagus (Fig. 14E, F). Parameres cone-shaped with almost even sparse punctuation, sparser on bases, a few long setae around apical portions, a few short setae distributed sparsely. Phallobase tube-like, consisting of two layers, anterior margin rounded, dorsal surface around anterior margin thin, protuberances of upper layer directed towards
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<pageBreakToken pageId="16" pageNumber="31" start="start">anterior</pageBreakToken>
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portion, small protuberances at beginning of divergence of upper layer. Penis stout, punctuated on posterior 1/9, rather coarsely on ventral surface.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="31" type="type series">
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<paragraph pageId="16" pageNumber="31">Type series.</paragraph>
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<paragraph pageId="16" pageNumber="31">
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Holotype: male, Yoshin, Tanzawa, Kanagawa Prefecture, Japan, 26
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<normalizedToken originalValue="–V–">-V-</normalizedToken>
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1989, Y. Hirano leg. (EUMJ). Paratypes: [Hokkaido Pref.] 2 exs.,
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<normalizedToken originalValue="Chûbisei">Chubisei</normalizedToken>
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, Memuro Town, 25
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<normalizedToken originalValue="–VIII–">-VIII-</normalizedToken>
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1995, S. Hisamatsu leg. (EUMJ); 1 ex., Mt. Sapporo-dake, Sapporo City, 5
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<normalizedToken originalValue="–VIII–">-VIII-</normalizedToken>
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1970, S. Kinoshita leg. (EUMJ). [Niigata Pref.] 1 ex., Mikuni Touge, 1
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1967, K. Baba leg. (HUSE). [Kanagawa Pref.] 1 ex., Yoshin, Tanzawa, 26
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<normalizedToken originalValue="–V–">-V-</normalizedToken>
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1989, Y. Hirano leg. (YHC). [Nagano Pref.] 1 ex., Tokugo Touge, 29
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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1955, T. Nakane leg. (HUSE); 5 exs.,
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<normalizedToken originalValue="Ôbora">Obora</normalizedToken>
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, Ueda City, 24
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<normalizedToken originalValue="–VII–">-VII-</normalizedToken>
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2013, T. Yoshida leg. (ELKU). [Oita Pref.] 6 ex., Mt. Sobo-san, 7
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<normalizedToken originalValue="–VI–">-VI-</normalizedToken>
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2009, S. Yamamoto leg. (ELKU).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="31" type="distribution">
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<paragraph pageId="16" pageNumber="31">Distribution.</paragraph>
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<paragraph pageId="16" pageNumber="31">JAPAN: Hokkaido, Honshu, Kyushu.</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="31" type="etymology">
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<paragraph pageId="16" pageNumber="31">Etymology.</paragraph>
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<paragraph pageId="16" pageNumber="31">
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The specific name is from the god of the north wind of ancient Greek mythology. Most
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="16" pageNumber="31" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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species are distributed in tropical or subtropical zones, however, this new species is exceptionally distributed in Hokkaido or on mountains of high altitude located in Honshu and Kyushu.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="31" type="remarks">
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<paragraph pageId="16" pageNumber="31">Remarks.</paragraph>
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<paragraph pageId="16" pageNumber="31">
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<taxonomicName class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Psammoecus" order="Coleoptera" pageId="16" pageNumber="31" phylum="Arthropoda" rank="genus">Psammoecus</taxonomicName>
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sp. 3 illustrated by
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Kanagawa-chuho" pageId="27" pageNumber="42" pagination="57 - 83" title="Notes on Japanese Silvanidae (Nihonsan hosohiratamushi-ka ni tsuite)." volume="168" year="2009">Hirano (2009)</bibRefCitation>
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and
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<bibRefCitation author="Hirano, Y" journalOrPublisher="Roppon-Ashi Entomological Books, Tokyo" pageId="27" pageNumber="42" title="Cucujoidea of Japan Vol. 2 Silvanidae, Byturidae, Biphyllidae." year="2010">Hirano (2010)</bibRefCitation>
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was conspecific with this species and named same Japanese name proposed by him.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |