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<document id="E179419A55691B08E867973D62F57360" ID-CLB-Dataset="28527" ID-DOI="10.11646/zootaxa.4549.1.1" ID-GBIF-Dataset="5d41ffeb-8b0d-4381-836d-ccd12a52142a" ID-ISSN="1175-5326" ID-Zenodo-Dep="2584949" ID-ZooBank="A97BF119-20C4-4428-9A00-68DC9F2BE34F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1550785238369" checkinUser="plazi" docAuthor="Stekolnikov, Alexandr A., Saboori, Alireza, Shamsi, Mohsen &amp; Hakimitabar, Masoud" docDate="2019" docId="C84D87CD8374D250FF18125764F6F8D6" docLanguage="en" docName="zootaxa.4549.1.1.pdf" docOrigin="Zootaxa 4549 (1)" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Helenicula goodorziani Kudryashova 1973" docType="treatment" docVersion="9" lastPageNumber="26" masterDocId="3474FFB5836DD249FF8F171F660FFF8E" masterDocTitle="Chigger mites (Acariformes: Trombiculidae) of Iran" masterLastPageNumber="66" masterPageNumber="1" pageNumber="26" updateTime="1698699467168" updateUser="ExternalLinkService">
<mods:mods id="0843FBFE5FDA45DAD1788E5AD07B1476" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="9BB9FCD94F731052E481F15FE552DAE4">Chigger mites (Acariformes: Trombiculidae) of Iran</mods:title>
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<mods:namePart id="10BA855CBFD79B74A0A9FE931C20B1A4">Stekolnikov, Alexandr A.</mods:namePart>
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<mods:namePart id="F3F8A64D8C4E6AAFDE9859CB0D7EC848">Saboori, Alireza</mods:namePart>
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<mods:namePart id="52241282CF2489DB6F99E88B816CE699">Shamsi, Mohsen</mods:namePart>
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<mods:namePart id="346E77C1C4DBE255FB61163BCA7E4F6A">Hakimitabar, Masoud</mods:namePart>
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<mods:title id="A772AC64C62E466E4B96F9F02578D121">Zootaxa</mods:title>
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<mods:date id="3DE00146B2584B57FDBC5AF5CEB47EAF">2019</mods:date>
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<mods:number id="5D21CB64FFBA90B58AE968A2F6F8E4E6">4549</mods:number>
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<treatment id="C84D87CD8374D250FF18125764F6F8D6" ID-DOI="http://doi.org/10.5281/zenodo.5250866" ID-GBIF-Taxon="154007179" ID-Zenodo-Dep="5250866" LSID="urn:lsid:plazi:treatment:C84D87CD8374D250FF18125764F6F8D6" httpUri="http://treatment.plazi.org/id/C84D87CD8374D250FF18125764F6F8D6" lastPageNumber="26" pageId="25" pageNumber="26">
<subSubSection id="08FE65508374D250FF18125764A2FAED" box="[151,685,1352,1379]" pageId="25" pageNumber="26" type="nomenclature">
<paragraph id="405B36DB8374D250FF18125764A2FAED" blockId="25.[151,685,1352,1379]" box="[151,685,1352,1379]" pageId="25" pageNumber="26">
<heading id="1B1381B78374D250FF18125764A2FAED" bold="true" box="[151,685,1352,1379]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
<taxonomicName id="87E44D588374D250FF18125764A2FAED" authority="Kudryashova, 1973" authorityName="Kudryashova" authorityYear="1973" box="[151,685,1352,1379]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="goodorziani">
<emphasis id="7290EAC98374D250FF18125764A2FAED" bold="true" box="[151,685,1352,1379]" pageId="25" pageNumber="26">
<emphasis id="7290EAC98374D250FF18125767BCFAEC" bold="true" box="[151,435,1352,1378]" italics="true" pageId="25" pageNumber="26">Helenicula goodorziani</emphasis>
Kudryashova, 1973
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="08FE65508374D250FF18129067CEFA4A" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="405B36DB8374D250FF1812906393FA2B" blockId="25.[151,1437,1423,1881]" box="[151,1436,1423,1446]" pageId="25" pageNumber="26">
<treatmentCitationGroup id="60F411F58374D250FF1812906393FA2B" box="[151,1436,1423,1446]" pageId="25" pageNumber="26">
<taxonomicName id="87E44D588374D250FF1812906466FA2B" authority="Kudryashova, 1973" authorityName="Kudryashova" authorityYear="1973" box="[151,617,1423,1445]" class="Arachnida" family="Trombiculidae" genus="Helenicula" higherTaxonomySource="GBIF" kingdom="Animalia" order="Prostigmata" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="goodorziani">
<emphasis id="7290EAC98374D250FF1812906782FA2B" box="[151,397,1423,1445]" italics="true" pageId="25" pageNumber="26">Helenicula goodorziani</emphasis>
Kudryashova,
<quantity id="871C9B3E8374D250FDB912906466FA2B" box="[566,617,1423,1445]" metricMagnitude="1" metricUnit="m" metricValue="5.011419999999999" pageId="25" pageNumber="26" unit="in" value="1973.0">1973</quantity>
</taxonomicName>
in:
<treatmentCitation id="C14510CA8374D250FD111290620CFA2B" author="Kudryashova, N. I. &amp; Neronov, V. M. &amp; Farang-Azad, A." box="[670,1027,1423,1446]" page="1725" pageId="25" pageNumber="26" year="1973">
<bibRefCitation id="24754B2A8374D250FD11129065B1FA2B" author="Kudryashova, N. I. &amp; Neronov, V. M. &amp; Farang-Azad, A." box="[670,958,1423,1446]" pageId="25" pageNumber="26" pagination="1725 - 1728" refId="ref54088" refString="Kudryashova, N. I., Neronov, V. M. &amp; Farang-Azad, A. (1973 b) New species and new findings of chiggers of Helenicula (Acariformes, Trombiculidae) from Iran. Zoologicheskiy Zhurnal, 52, 1725 - 1728. [in Russian]" type="journal article" year="1973">
Kudryashova
<emphasis id="7290EAC98374D250FCBC128E6562FA2B" box="[819,877,1423,1446]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
1973b
</bibRefCitation>
: 1725
</treatmentCitation>
, fig.;
<treatmentCitation id="C14510CA8374D250FBC312906398FA2B" author="Kudryashova, N. I. &amp; Neronov, V. M. &amp; Farang-Azad, A." box="[1100,1431,1423,1446]" page="158" pageId="25" pageNumber="26" year="1978">
<bibRefCitation id="24754B2A8374D250FBC312906352FA2B" author="Kudryashova, N. I. &amp; Neronov, V. M. &amp; Farang-Azad, A." box="[1100,1373,1423,1446]" pageId="25" pageNumber="26" pagination="92 - 180" refId="ref54194" refString="Kudryashova, N. I., Neronov, V. M. &amp; Farang-Azad, A. (1978) Mites of the family Trombiculidae (Acariformes) from small mammals from Iran. Sbornik Trudov Zoologicheskogo Muzeya MGU, 16, 92 - 180. [in Russian]" type="journal article" year="1978">
Kudryashova
<emphasis id="7290EAC98374D250FB6F128E6314FA2B" box="[1248,1307,1423,1446]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
1978
</bibRefCitation>
: 158
</treatmentCitation>
;
</treatmentCitationGroup>
</paragraph>
<paragraph id="405B36DB8374D250FF4812B167CEFA4A" blockId="25.[151,1437,1423,1881]" box="[199,449,1454,1476]" pageId="25" pageNumber="26">
<bibRefCitation id="24754B2A8374D250FF4812B16785FA4A" author="Kudryashova, N. I." box="[199,394,1454,1476]" pageId="25" pageNumber="26" refId="ref53933" refString="Kudryashova, N. I. (1998) Chigger mites (Acariformes, Trombiculidae) of East Palaearctics. KMK Scientific Press, Moscow, 342 pp. [in Russian]" type="book" year="1998">Kudryashova 1998</bibRefCitation>
: 276.
</paragraph>
</subSubSection>
<subSubSection id="08FE65508374D250FF1812D362ECF948" pageId="25" pageNumber="26" type="diagnosis">
<paragraph id="405B36DB8374D250FF1812D362ECF948" blockId="25.[151,1437,1423,1881]" pageId="25" pageNumber="26">
<emphasis id="7290EAC98374D250FF1812D3671EFA6B" bold="true" box="[151,273,1484,1509]" pageId="25" pageNumber="26">Diagnosis.</emphasis>
SIF = 5B-B-3-1111.0000; fPp = B/B/BNB; fsp = 7.7.7; fCx = 1.1.3; fSt = 2.4; fSc: AL&gt;
<materialsCitation id="F08C3C868374D250FA8C12D1652DF987" ID-GBIF-Occurrence="2005760954" collectingDate="2012-06-04" collectingDateMax="2012-06-08" collectingDateMin="2012-06-04" collectionCode="PL, AM" pageId="25" pageNumber="26" specimenCount="1">
<collectionCode id="26F5AE1E8374D250FA8C12D1632BFA6B" box="[1283,1316,1486,1509]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="25" pageNumber="26" type="Museum">PL</collectionCode>
&gt;
<collectionCode id="26F5AE1E8374D250FACD12D2637CFA6B" box="[1346,1395,1485,1509]" country="Australia" name="Australian Museum" pageId="25" pageNumber="26" type="Museum">AM</collectionCode>
; fD = 4H-8-8-4-8-4-4-6-6-
<date id="345A101B8374D250FE1912EF67F7F987" box="[406,504,1520,1545]" pageId="25" pageNumber="26" value="2012-06-08" valueMax="2012-06-04" valueMin="2012-06-08">
<collectingDate id="241EE9F38374D250FE1912EF67F7F987" box="[406,504,1520,1545]" pageId="25" pageNumber="26" value="2012-06-08" valueMax="2012-06-04" valueMin="2012-06-08">8-4-12-6</collectingDate>
</date>
; DS = 72–95; VS = 55–71
</materialsCitation>
; NDV = 134–163; Ip = 793–862; eyes 2 + 2; S
<subScript id="DC60349E8374D250FAA612E0633EF983" attach="left" box="[1321,1329,1535,1549]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
apical, at level of ST; f
<subScript id="DC60349E8374D250FEA51139673DF9BA" attach="left" box="[298,306,1574,1588]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
posterior to S
<subScript id="DC60349E8374D250FE5F113967D7F9BA" attach="right" box="[464,472,1574,1588]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
; tibialae I apical; f
<subScript id="DC60349E8374D250FD23113964BBF9BA" attach="left" box="[684,692,1574,1588]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
anterior to S
<subScript id="DC60349E8374D250FCCA11396542F9BA" attach="right" box="[837,845,1574,1588]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
; S
<subScript id="DC60349E8374D250FCE41139657CF9BA" attach="left" box="[875,883,1574,1588]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
&gt; S
<subScript id="DC60349E8374D250FC2F113965A7F9BA" attach="left" box="[928,936,1574,1588]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
. Measurements of type series (
<bibRefCitation id="24754B2A8374D250FA8B1108671CF9D9" author="Kudryashova, N. I. &amp; Neronov, V. M. &amp; Farang-Azad, A." pageId="25" pageNumber="26" pagination="92 - 180" refId="ref54194" refString="Kudryashova, N. I., Neronov, V. M. &amp; Farang-Azad, A. (1978) Mites of the family Trombiculidae (Acariformes) from small mammals from Iran. Sbornik Trudov Zoologicheskogo Muzeya MGU, 16, 92 - 180. [in Russian]" type="journal article" year="1978">
Kudryashova
<emphasis id="7290EAC98374D250FF18112066C0F9D9" box="[151,207,1598,1623]" italics="true" pageId="25" pageNumber="26">et al.</emphasis>
1978
</bibRefCitation>
): AW 56–61,
<materialsCitation id="F08C3C868374D250FE2211206427F9D9" ID-GBIF-Occurrence="2005760940" box="[429,552,1598,1623]" collectionCode="PW" pageId="25" pageNumber="26" specimenCode="PW 69-78" specimenCount="1">
<specimenCode id="10429EA08374D250FE2211206427F9D9" box="[429,552,1598,1623]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="25" pageNumber="26">PW 69–78</specimenCode>
</materialsCitation>
, SB 8–11, ASB 28–31, PSB 14–17, SD 42–45, AP 19–25,
<materialsCitation id="F08C3C868374D250FB3111206335F9D9" ID-GBIF-Occurrence="2005760910" box="[1214,1338,1598,1623]" collectionCode="AM" pageId="25" pageNumber="26" specimenCode="AM 30-36" specimenCount="1">
<specimenCode id="10429EA08374D250FB3111206335F9D9" box="[1214,1338,1598,1623]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="25" pageNumber="26" type="Museum">AM 30–36</specimenCode>
</materialsCitation>
, AL 42– 50,
<materialsCitation id="F08C3C868374D250FF48117C6731F9F5" ID-GBIF-Occurrence="2005760939" box="[199,318,1634,1659]" collectionCode="PL" pageId="25" pageNumber="26" specimenCode="PL 39-45" specimenCount="1">
<specimenCode id="10429EA08374D250FF48117C6731F9F5" box="[199,318,1634,1659]" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="25" pageNumber="26" type="Museum">PL 39–45</specimenCode>
</materialsCitation>
, S 25, H 36–45, D
<subScript id="DC60349E8374D250FDA3116E644BF9F1" attach="left" box="[556,580,1649,1663]" fontSize="6" pageId="25" pageNumber="26">min</subScript>
19, D
<subScript id="DC60349E8374D250FD19116E64BEF9F1" attach="left" box="[662,689,1649,1663]" fontSize="6" pageId="25" pageNumber="26">max</subScript>
36, V
<subScript id="DC60349E8374D250FC8C116E6515F9F1" attach="left" box="[771,794,1649,1663]" fontSize="6" pageId="25" pageNumber="26">min</subScript>
17, V
<subScript id="DC60349E8374D250FCE3116E6588F9F1" attach="left" box="[876,903,1649,1663]" fontSize="6" pageId="25" pageNumber="26">max</subScript>
33, pa 281–308, pm 231–258, pp 281–300. Measurements of
<typeStatus id="9F5F88798374D250FEEE119667C8F92C" box="[353,455,1673,1698]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
: AW 58,
<materialsCitation id="F08C3C868374D250FDBF1195648FF92C" ID-GBIF-Occurrence="2005760933" box="[560,640,1674,1698]" collectionCode="PW" pageId="25" pageNumber="26" specimenCode="PW 72" specimenCount="1">
<specimenCode id="10429EA08374D250FDBF1195648FF92C" box="[560,640,1674,1698]" collectionCode="PW" country="Thailand" name="Paleontological Collections" pageId="25" pageNumber="26">PW 72</specimenCode>
</materialsCitation>
, SB 10, ASB 29, PSB 18, SD 47, P-PL 23, AP 23,
<materialsCitation id="F08C3C868374D250FB4A11956317F92C" ID-GBIF-Occurrence="2005760962" box="[1221,1304,1674,1698]" collectionCode="AM" pageId="25" pageNumber="26" specimenCode="AM 34" specimenCount="1">
<specimenCode id="10429EA08374D250FB4A11956317F92C" box="[1221,1304,1674,1698]" collectionCode="AM" country="Australia" name="Australian Museum" pageId="25" pageNumber="26" type="Museum">AM 34</specimenCode>
</materialsCitation>
, AL 42,
<materialsCitation id="F08C3C868374D250FAF4119566BAF948" ID-GBIF-Occurrence="2005760950" collectionCode="PL" pageId="25" pageNumber="26" specimenCode="PL 41" specimenCount="1">
<specimenCode id="10429EA08374D250FAF4119566BAF948" collectionCode="PL" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="25" pageNumber="26" type="Museum">PL 41</specimenCode>
</materialsCitation>
, H 42, D
<subScript id="DC60349E8374D250FE9511A3673DF944" attach="left" box="[282,306,1724,1738]" fontSize="6" pageId="25" pageNumber="26">min</subScript>
25, D
<subScript id="DC60349E8374D250FEF811A3679DF944" attach="left" box="[375,402,1724,1738]" fontSize="6" pageId="25" pageNumber="26">max</subScript>
38, V
<subScript id="DC60349E8374D250FE5811A367E0F944" attach="left" box="[471,495,1724,1738]" fontSize="6" pageId="25" pageNumber="26">min</subScript>
19, V
<subScript id="DC60349E8374D250FDBB11A36440F944" attach="left" box="[564,591,1724,1738]" fontSize="6" pageId="25" pageNumber="26">max</subScript>
32, pa 288, pm 254, pp 288, Ip 830, TaIIIL 76, TaIIIW 18.
</paragraph>
</subSubSection>
<subSubSection id="08FE65508374D250FF4811CB659EF89E" pageId="25" pageNumber="26" type="materials_examined">
<paragraph id="405B36DB8374D250FF4811CB659EF89E" blockId="25.[151,1437,1423,1881]" pageId="25" pageNumber="26">
<emphasis id="7290EAC98374D250FF4811CB67F9F963" bold="true" box="[199,502,1748,1773]" pageId="25" pageNumber="26">Type material examined.</emphasis>
<materialsCitation id="F08C3C868374D250FD8F11CB659EF89E" ID-GBIF-Occurrence="2005760921" collectingDate="1969-10-14" collectionCode="ZMMU" collectorName="V. M. Neronov." elevation="1600" pageId="25" pageNumber="26" specimenCount="1" typeStatus="holotype">
<typeStatus id="9F5F88798374D250FD8F11CB6467F963" box="[512,616,1748,1773]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
larva (
<collectionCode id="26F5AE1E8374D250FD3211CB6519F962" box="[701,790,1748,1772]" country="Russia" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="25" pageNumber="26" type="Museum">ZMMU</collectionCode>
Tdt-205, I-108-1865) from
<taxonomicName id="87E44D588374D250FBD011CA66FCF89F" authority=", Delijan, Anarbar" authorityName=", Delijan, Anarbar" authorityYear="1600" class="Mammalia" family="Muridae" genus="Meriones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="persicus">
<emphasis id="7290EAC98374D250FBD011CA633FF962" box="[1119,1328,1749,1772]" italics="true" pageId="25" pageNumber="26">Meriones persicus</emphasis>
, Delijan, Anarbar
</taxonomicName>
valley,
<elevation id="CBC9D1E88374D250FEC511E767D3F89F" box="[330,476,1784,1809]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="25" pageNumber="26" unit="m" value="1600.0">
<quantity id="871C9B3E8374D250FEC511E76790F89E" box="[330,415,1784,1809]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="25" pageNumber="26" unit="m" value="1600.0">1600 m</quantity>
a.s.l.
</elevation>
,
<date id="345A101B8374D250FE6711E764A8F89F" box="[488,679,1784,1809]" pageId="25" pageNumber="26" value="1969-10-14">
<collectingDate id="241EE9F38374D250FE6711E764A8F89F" box="[488,679,1784,1809]" pageId="25" pageNumber="26" value="1969-10-14">14 October 1969</collectingDate>
</date>
, coll.
<collectorName id="ED11530D8374D250FD6511E6659EF89E" box="[746,913,1785,1808]" pageId="25" pageNumber="26">V.M. Neronov.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="08FE65508374D250FF4810036496F8BA" box="[199,665,1820,1845]" pageId="25" pageNumber="26" type="biology_ecology">
<paragraph id="405B36DB8374D250FF4810036496F8BA" blockId="25.[151,1437,1423,1881]" box="[199,665,1820,1845]" pageId="25" pageNumber="26">
<emphasis id="7290EAC98374D250FF481003671EF8BB" bold="true" box="[199,273,1820,1845]" pageId="25" pageNumber="26">Hosts.</emphasis>
<emphasis id="7290EAC98374D250FE971002647DF8BB" box="[280,626,1820,1845]" italics="true" pageId="25" pageNumber="26">
<taxonomicName id="87E44D588374D250FE97100267E7F8BA" authorityName=", Delijan, Anarbar" authorityYear="1600" box="[280,488,1821,1844]" class="Mammalia" family="Muridae" genus="Meriones" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="persicus">Meriones persicus</taxonomicName>
,
<taxonomicName id="87E44D588374D250FE7B1003647DF8BB" box="[500,626,1820,1845]" class="Mammalia" family="Muridae" genus="Chionomys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="25" pageNumber="26" phylum="Chordata" rank="genus">Chionomys</taxonomicName>
</emphasis>
sp.
</paragraph>
</subSubSection>
<subSubSection id="08FE65508374D250FF48105F64F6F8D6" box="[199,761,1856,1881]" pageId="25" pageNumber="26" type="distribution">
<paragraph id="405B36DB8374D250FF48105F64F6F8D6" blockId="25.[151,1437,1423,1881]" box="[199,761,1856,1881]" pageId="25" pageNumber="26">
<emphasis id="7290EAC98374D250FF48105F676FF8D7" bold="true" box="[199,352,1856,1881]" pageId="25" pageNumber="26">Distribution.</emphasis>
<collectingCountry id="38F3764B8374D250FEE8105E679BF8D6" box="[359,404,1857,1880]" name="Iran" pageId="25" pageNumber="26">Iran</collectingCountry>
(Delijan,
<collectingRegion id="8220F8398374D250FD8A105E646DF8D6" box="[517,610,1857,1880]" country="Iran" name="Kerman" pageId="25" pageNumber="26">Kerman</collectingRegion>
, Mashhad 2).
</paragraph>
</subSubSection>
</treatment>
</document>
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