203 lines
23 KiB
XML
203 lines
23 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.544.6074" ID-GBIF-Dataset="0e5e47a0-f8f3-4039-b0e1-c159fef0b9f6" ID-PMC="PMC4714384" ID-Pensoft-Pub="1313-2970-544-1" ID-PubMed="26798289" ID-ZBK="C5978BD0145B40F8ACDEB27371B7B9A4" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-544-1" ModsDocOrigin="ZooKeys 544" ModsDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" checkinTime="1451243640953" checkinUser="pensoft" docAuthor="Liebherr, James K." docDate="2015" docId="B7F2080CAC8EC5164AE116238F12CFBC" docLanguage="en" docName="ZooKeys 544: 1-407" docOrigin="ZooKeys 544" docSource="http://dx.doi.org/10.3897/zookeys.544.6074" docTitle="Mecyclothorax scarites Liebherr, 2015, sp. n." docType="treatment" docUuid="3A931DB6-433E-4A1D-A333-31FE391055A6" docUuidSource="ZooBank" docVersion="5" lastPageNumber="192" masterDocId="FF92FFC2FF9FFFC7191EFFB8FF91AF5D" masterDocTitle="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation" masterLastPageNumber="407" masterPageNumber="1" pageNumber="190" updateTime="1668162300918" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala-, Maui: Keystone of a hyperdiverse Hawaiian radiation</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liebherr, James K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2015</mods:date>
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<mods:detail type="volume">
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<mods:number>544</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>407</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.544.6074</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.544.6074</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-544-1</mods:identifier>
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<mods:identifier type="ZBK">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
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<mods:identifier type="ZooBank">C5978BD0145B40F8ACDEB27371B7B9A4</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127853560" LSID="urn:lsid:zoobank.org:act:3A931DB6-433E-4A1D-A333-31FE391055A6" httpUri="http://treatment.plazi.org/id/B7F2080CAC8EC5164AE116238F12CFBC" lastPageId="191" lastPageNumber="192" pageId="189" pageNumber="190">
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<subSubSection pageId="189" pageNumber="190" type="multiple">
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<paragraph pageId="189" pageNumber="190">
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<pageBreakToken pageId="189" pageNumber="190" start="start">Taxon</pageBreakToken>
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classification Animalia Coleoptera Carabidae
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</paragraph>
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</subSubSection>
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<subSubSection pageId="189" pageNumber="190" type="nomenclature">
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<paragraph pageId="189" pageNumber="190">
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(079)
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<taxonomicName LSID="http://zoobank.org/3A931DB6-433E-4A1D-A333-31FE391055A6" class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scarites" order="Coleoptera" pageId="189" pageNumber="190" phylum="Arthropoda" rank="species" species="scarites">Mecyclothorax scarites</taxonomicName>
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<taxonomicNameLabel pageId="189" pageNumber="190">sp. n.</taxonomicNameLabel>
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Figs 95D, 96
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<normalizedToken originalValue="D–E">D-E</normalizedToken>
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, 97C, 98C, 99
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</paragraph>
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</subSubSection>
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<subSubSection pageId="190" pageNumber="191" type="reference_group">
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<paragraph pageId="190" pageNumber="191">
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<taxonomicName class="Insecta" family="Carabidae" genus="Metrothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Metrothorax perkinsianus" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="perkinsianus">
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<pageBreakToken pageId="190" pageNumber="191" start="start">Metrothorax</pageBreakToken>
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perkinsianus
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</taxonomicName>
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Sharp,
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<bibRefCitation author="Swezey, OH" journalOrPublisher="The Hawaiian Botanical Society and The Hawaiian Entomological Society, Honolulu" pageId="310" pageNumber="311" title="Forest Entomology in Hawaii: An annotated check-list of the insect faunas of the various components of the Hawaiian forests" year="1954">Swezey 1954</bibRefCitation>
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: 53 (misidentification,
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<taxonomicName class="Polypodiopsida" family="Cibotiaceae" genus="Cibotium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cibotium" order="Cyatheales" pageId="190" pageNumber="191" phylum="Tracheophyta" rank="genus">Cibotium</taxonomicName>
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associate).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="190" pageNumber="191" type="diagnosis">
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<paragraph pageId="190" pageNumber="191">Diagnosis.</paragraph>
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<paragraph pageId="190" pageNumber="191">
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This species (Fig. 95D) is assigned to the setal formula 1 0 2 0 along with two other
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<normalizedToken originalValue="Haleakalā">Haleakala</normalizedToken>
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species-
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax molops" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="molops">Mecyclothorax molops</taxonomicName>
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(Fig. 95A) and
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax macrops" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="macrops">Mecyclothorax macrops</taxonomicName>
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(Fig. 95B)-yet it is more similar in pronotal configuration, overall body proportions, and elytral striation to
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
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(Fig. 95C), a species characterized by presence of both supraorbital setae. In addition to the supraorbital setae,
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scarites" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="scarites">Mecyclothorax scarites</taxonomicName>
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can be distinguished from
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
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by the shallow but distinct transverse microsculpture on the elytral disc that is composed of sculpticells 3
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<normalizedToken originalValue="–4×">-4x</normalizedToken>
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broad as long plus transverse lines not joined into a mesh. The discal elytral striae are not punctate, with the sutural stria irregularly impressed along its discal portion, and striae 2-4 variable impressed among individuals, but smooth to slightly irregular across the disc in all individuals. The elytra are slightly more ovoid than those of
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="190" pageNumber="191" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
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, with the humeral angles more distant relative to overall elytral width; MEW/HuW = 1.93-2.0. Standardized body length 5.3-5.5 mm.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="191" lastPageNumber="192" pageId="190" pageNumber="191" type="description">
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<paragraph pageId="190" pageNumber="191">Description</paragraph>
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<paragraph lastPageId="191" lastPageNumber="192" pageId="190" pageNumber="191">
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(n = 5). Head capsule with frontal grooves broad, sinuous, a broad lateral convexity before eye, groove nearly joined to postocular groove anterad posterior supraorbital seta; dorsal impression of neck slightly concave; eyes moderately convex, covering much of ocular lobe, ocular ratio = 1.47-1.52, ocular lobe ratio = 0.84-0.88; labral anterior margin very shallowly emarginate medially; antennae filiform, antennomeres 1-3 glabrous; mentum tooth with sides acute, apex tightly rounded. Pronotum moderately transverse, MPW/PL = 1.20-1.22, base moderately constricted, MPW/BPW = 1.47-1.55, lateral margin sinuate for 0.1
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<normalizedToken originalValue="×">x</normalizedToken>
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pronotal length anterad obtuse hind angles, their apex rounded; median base coplanar with disc medially, ~14 small, isolated punctures each side; basal margin moderately convex between hind angles; median longitudinal impression very shallowly incised, traceable to obsolete; anterior transverse impression broad, shallow medially, narrow and finely incised laterally; anterior callosity slightly convex, smooth; front angles slightly projected, tightly rounded; pronotal apical and basal widths subequal, APW/BPW = 0.94-1.05; lateral marginal depression very narrow, edge beaded even at front angle, disc very convex; laterobasal depression smooth, convex with U-shaped groove along lateral and basal margins of convexity. Proepisternum with 5 minute punctures along hind marginal groove; prosternal process with very shallow depression medially. Elytra ovoid, greatest width slightly behind midlength; disc very convex, slightly depressed along suture, sides distinctly sloped to near vertical inside marginal depression; basal groove distinctly curved to subangulate humerus; parascutellar seta present; parascutellar striole shallow, with 3-4 elongate punctures, smooth between punctures; sutural interval slightly more convex than intervals 2-4 throughout their length; sutural stria narrow, moderately deep apically, 2nd stria obsolete, traceable at apex; 8th interval laterad 7th stria of similar convexity to fused apical portion of striae 5 + 7; 2 dorsal elytral setae at 0.19
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<normalizedToken originalValue="–0.23×">-0.23x</normalizedToken>
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and 0.52
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<normalizedToken originalValue="–0.53×">-0.53x</normalizedToken>
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elytral length, setal impressions very small, crossing 1/3 of interval 3; lateral elytral setae arranged in anterior series of 7 setae and posterior series of 6 setae; elytral marginal depression very narrow throughout, a low bead present at humerus, a more distinct bead near subapical sinuation; subapical sinuation very shallow, less concave than width of marginal bead. Mesepisternum with ~10 punctures in 2-3 rows; metepisternal width to length ratio = 0.73; metepisternum/metepimeron suture distinct. Abdomen with irregular lateral wrinkles on ventrites 1-6, lateral depressions on ventrites 3-6; suture between ventrites 2 and 3 effaced; apical male ventrite with 2 marginal setae and apical female ventrite with 4 equally spaced setae plus median trapezoid of 4 subequal, short setae. Legs-metatarsomere 1/metatibial length ratio = 0.20; metatarsomere 4 length along outer lobe 1.5
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<normalizedToken originalValue="×">x</normalizedToken>
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medial tarsomere length, apical and subapical setae present, the lateral subapical seta short; metatarsal dorsolateral sulci broad, shallow, medially subcarinate on metatarsomere 1 only. Microsculpture of vertex obsolete transverse mesh, sculpticell breadth 2
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<normalizedToken originalValue="×">x</normalizedToken>
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length, surface glossy; pronotal disc with obsolete transverse mesh, sculpticell breadth 3
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<normalizedToken originalValue="–4×">-4x</normalizedToken>
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length, most parts glossy, median base glossy, indistinct transverse cells laterally; elytral apex with shallow transverse mesh, sculpticell breadth 2
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<normalizedToken originalValue="–3×">-3x</normalizedToken>
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length; metasternum with obsolete transverse mesh, glossy; laterobasal abdominal ventrites with swirling isodiametric and transverse microsculpture. Coloration of vertex rufous with a piceous cast; antennomeres 1-3 rufoflavous, 4-11 rufobrunneous; pronotal disc rufous, concolorous basally and later
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<pageBreakToken pageId="191" pageNumber="192" start="start">ally</pageBreakToken>
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, apex slightly paler; proepipleuron pale rufobrunneous, proepisternum rufobrunneous; elytral disc rufous, sutural interval concolorous basally, rufoflavous apically; elytral marginal depression and 9th interval rufoflavous apically; elytral epipleuron pale rufobrunneous, metepisternum rufobrunneous; abdomen rufobrunneous, ventrites 1-3 rufoflavous laterally, apical 1/6 of apical ventrite flavous; metafemur rufoflavous; metatibia rufoflavous with brunneous cast.
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</paragraph>
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<paragraph pageId="191" pageNumber="192">
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Male genitalia (n = 2). Aedeagal median lobe slender, distance from parameral articulation to tip 5.0
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<normalizedToken originalValue="×">x</normalizedToken>
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depth at midlength (Fig. 96D); apex distinctly expanded dorsally and broadly expanded ventrally producing an apical knob, the outline of the tip broadly rounded; internal sac elongate, sac length from ventral margin of ostial opening to base of flagellar plate 0.78
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<normalizedToken originalValue="×">x</normalizedToken>
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parameral articulation-tip distance, without ornamentation (Fig. 96E); flagellar plate very short, nearly as broad as long, length 0.24
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<normalizedToken originalValue="×">x</normalizedToken>
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parameral articulation-tip distance.
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</paragraph>
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<paragraph pageId="191" pageNumber="192">
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Female reproductive tract (n = 1). Bursa copulatrix columnar with narrower apex, broader base, length 0.91 mm, apical breadth 0.23 mm, basal breadth 0.41 mm (Fig. 97C); bursal walls translucent, thinly wrinkled; gonocoxite 1 with 3-4 apical fringe setae (Fig. 98C), a thick, curved seta at medioapical angle and 10-11 smaller setae along medial surface; gonocoxite 2 falcate with acuminate tip, base broadly extended laterally with curved terminus, 2 broad lateral ensiform setae, apical nematiform setae on medioventral surface at 0.66
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<normalizedToken originalValue="×">x</normalizedToken>
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gonocoxite length.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="191" pageNumber="192" type="holotype">
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<paragraph pageId="191" pageNumber="192">Holotype.</paragraph>
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<paragraph pageId="191" pageNumber="192">
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Male (BPBM) dissected and labeled: Kula Pipe / Line, Maui / 4200 ft. / Jan. 14, 1926 // R H VanZwal- / uwenburg // blue square //
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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/
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<taxonomicName lsidName="scarites" pageId="191" pageNumber="192" rank="species" species="scarites">scarites</taxonomicName>
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/ ♂ #2 / det. J.K. Liebherr 2014 // 1 // HOLOTYPE /
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="genus">Mecyclothorax</taxonomicName>
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/
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<taxonomicName lsidName="scarites" pageId="191" pageNumber="192" rank="species" species="scarites">scarites</taxonomicName>
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/ Liebherr / det. J.K. Liebherr 2015 (black-margined red label).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="191" pageNumber="192" type="paratypes">
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<paragraph pageId="191" pageNumber="192">Paratypes.</paragraph>
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<paragraph pageId="191" pageNumber="192">
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HI: Maui, Haleakala, Olinda,
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<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus" order="Rosales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
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, 1280 m el., 27-ii-1926, Swezey (BPBM, 1),
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<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Cheirodendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cheirodendron" order="Apiales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="genus">Cheirodendron</taxonomicName>
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, 1280 m el., 27-ii-1926, Swezey (BPBM, 1),
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<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Acacia" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Acacia koa" order="Fabales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="species" species="koa">Acacia koa</taxonomicName>
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, 1280 m el., 10-x-1926, Swezey (BPBM, 1; CUIC, 1); Koolau For. Res., Kula Pipeline Rd.,
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<taxonomicName class="Polypodiopsida" family="Cibotiaceae" genus="Cibotium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cibotium menziesii" order="Cyatheales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="species" species="menziesii">Cibotium menziesii</taxonomicName>
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, 975-1210 m el., 27-ii-1935, Swezey (BPBM, 1).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="191" pageNumber="192" type="etymology">
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<paragraph pageId="191" pageNumber="192">Etymology.</paragraph>
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<paragraph pageId="191" pageNumber="192">
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The cylindrical body shape of
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
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appears to be the basis for
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<bibRefCitation author="Blackburn, T" journalOrPublisher="The Entomologist's Monthly Magazine" pageId="307" pageNumber="308" pagination="156 - 158" title="Characters of new genera and descriptions of new species of Geodephaga from the Hawaiian Islands." volume="15" year="1878 b">Blackburn (1878b)</bibRefCitation>
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using a species epithet derived from the Greek skaritis, that stem used by
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<bibRefCitation author="Fabricius, JC" journalOrPublisher="Officina Libraria Kortii, Flensburg and Leipzig" pageId="308" pageNumber="309" title="Systema entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus" url="10.5962/bhl.title.36510" year="1775">Fabricius (1775)</bibRefCitation>
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to name the large-bodied
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<taxonomicName class="Insecta" family="Carabidae" genus="Scarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scarites" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="genus">Scarites</taxonomicName>
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beetles of the Holarctic. This species is given the species epithet
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<taxonomicName lsidName="scarites" pageId="191" pageNumber="192" rank="species" species="scarites">scarites</taxonomicName>
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to complete the cycle with the generic name
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<taxonomicName class="Insecta" family="Carabidae" genus="Scarites" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Scarites" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="genus">Scarites</taxonomicName>
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F.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="191" pageNumber="192" type="distribution and habitat">
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<paragraph pageId="191" pageNumber="192">Distribution and habitat.</paragraph>
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<paragraph pageId="191" pageNumber="192">
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scarites" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scarites">Mecyclothorax scarites</taxonomicName>
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is known from two localities peripheral to the known localities of
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
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(Fig. 99) in the leeward forests uphill (mauka) from Olinda. Based on the careful ecological labeling of specimens by O.H. Swezey, this species is known to have occurred on
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<taxonomicName lsidName="koa" pageId="191" pageNumber="192" rank="species" species="koa">koa</taxonomicName>
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,
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<taxonomicName class="Magnoliopsida" family="Araliaceae" genus="Cheirodendron" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cheirodendron" order="Apiales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="genus">Cheirodendron</taxonomicName>
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('ōlapa),
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<taxonomicName class="Polypodiopsida" family="Cibotiaceae" genus="Cibotium" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Cibotium menziesii" order="Cyatheales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="species" species="menziesii">Cibotium menziesii</taxonomicName>
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(
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<normalizedToken originalValue="hāpu‘u">hapu'u</normalizedToken>
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), and
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<taxonomicName class="Magnoliopsida" family="Rosaceae" genus="Rubus" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Rubus" order="Rosales" pageId="191" pageNumber="192" phylum="Tracheophyta" rank="genus">Rubus</taxonomicName>
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(
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<normalizedToken originalValue="‘ākala">'akala</normalizedToken>
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).
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<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scarites" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scarites">Mecyclothorax scarites</taxonomicName>
|
||
is closely related yet clearly distinct from
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
|
||
. The two species were parapatrically distributed, with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scarites" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scarites">Mecyclothorax scarites</taxonomicName>
|
||
appearing in the collecting record subsequent to the last appearance of any specimen of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Mecyclothorax" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mecyclothorax scaritoides" order="Coleoptera" pageId="191" pageNumber="192" phylum="Arthropoda" rank="species" species="scaritoides">Mecyclothorax scaritoides</taxonomicName>
|
||
. Their patchwork appearances in the scientific record are consistent with a haphazard pattern of habitat destruction along the leeward edge of the 19th Century Koa-
|
||
<normalizedToken originalValue="‘Ōhi‘a">'Ōhi'a</normalizedToken>
|
||
Mesic Forest.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |