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<mods:title id="4B42492F391A4E9E5F5FE32509D12BCE">Contributions on a small collection of the former Subulinidae Fischer &amp; Crosse, 1877 (Eupulmonata, Achatinoidea) with catalogue of the Glessula and Rishetia species recorded from Myanmar</mods:title>
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<mods:affiliation id="B5308D9CAA3DC9721407DD6C3846AA12">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok, 10330, Thailand &amp; Department of Zoology, University of Yangon, University Avenue Road, Kamayut Township 11041, Yangon, Myanmar</mods:affiliation>
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<heading id="1B0C615727F51ADEF3635A2DFFBAC791" reason="label">Genus</heading>
<taxonomicName id="9E2A08508B25D5B322FDA275CC9FF2CA" authority="Albers, 1850" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Subulinidae" genus="Opeas" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="658058FAE8D5C2DB7CAB3603EEDFD628" italics="true">Opeas</emphasis>
<bibRefCitation id="AC69714E8100ED514DE9738FF001916A" DOI="10.5962/bhl.title.11507" author="Albers JC" journalOrPublisher="Enslin, Berlin" refId="B2" refString="Albers JC (1850) Die Heliceen nach naturlicher Verwandtschaft systematisch geordnet. Enslin, Berlin, 262 pp. https: // doi. org / 10.5962 / bhl. title. 11507" title="Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet." year="1850">Albers, 1850</bibRefCitation>
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<treatmentCitation id="B85BA814A9EC9FC81AE85D0453308690" author="Albers JC" page="175" year="1850">
<taxonomicName id="40F7015EF3E7F8EE10AC2D10047F3BC2" authority="Albers, 1850: 175" authorityName="Albers" authorityPageNumber="175" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Bulimus" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subGenus" subGenus="Opeas">
<emphasis id="3EDD5A2CCFFB11D45F8F6B40EE06F0AF" italics="true">Bulimus</emphasis>
(
<emphasis id="7790455E4DE78FEA273BD9529361AFDF" italics="true">Opeas</emphasis>
)
<comment id="9A38E195A86778B952A77D81C11C6FB2">
<bibRefCitation id="9565F1C02767F586191D6ED33A2460B2" DOI="10.5962/bhl.title.11507" author="Albers JC" journalOrPublisher="Enslin, Berlin" refId="B2" refString="Albers JC (1850) Die Heliceen nach naturlicher Verwandtschaft systematisch geordnet. Enslin, Berlin, 262 pp. https: // doi. org / 10.5962 / bhl. title. 11507" title="Die Heliceen nach natürlicher Verwandtschaft systematisch geordnet." year="1850">Albers, 1850: 175</bibRefCitation>
, 176.
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</paragraph>
<paragraph id="1960A74257DADFECA014150EFD30CAE9">
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<taxonomicName id="6981C41D64CC9A99C9A2C42A1F71E336" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Stenogyra" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="subGenus" subGenus="Opeas">
<emphasis id="233FB8654415DCE60693CEAB223ADFBD" italics="true">Stenogyra</emphasis>
(
<emphasis id="BC73A38C2FFC5704B0B12FCBD5EA9B01" italics="true">Opeas</emphasis>
)
</taxonomicName>
<comment id="DFD15699A5EE9D6E2B5BEA83F8B1D9E2">
<treatmentCitation id="3CE0D921A564D866F13F58D29FC361AF" author="von Martens E" page="265" year="1860">
<bibRefCitation id="0B1D233DC64328A403796583C1DD0C7E" author="von Martens E" firstAuthor="von Martens" journalOrPublisher="Proceedings of the Zoological Society of London" page="265" pagination="6-18" refId="B114" refString="von Martens E (1860) On the Mollusca of Siam. Proceedings of the Zoological Society of London 28: 618. https://www.biodiversitylibrary.org/page/12866504" title="On the Mollusca of Siam." url="https://www.biodiversitylibrary.org/page/12866504" volume="28" year="1860">von Martens 1860: 265</bibRefCitation>
</treatmentCitation>
, 266.
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</paragraph>
<paragraph id="B1DBCF83187CDBD4DD427532F00C4C28">
<treatmentCitationGroup id="B536C0763CE8E26A3A0F36BC2234CFAC">
<taxonomicName id="75266C6DFC90EF9313CC005F1DA5DB60" ID-CoL="7PB9P" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="DD70DA8B104C8AB6ADF74BBC5DF84514" italics="true">Opeas</emphasis>
</taxonomicName>
<comment id="C79A383AB1ECD6518ACA4EC0A3CE0DD6">
<treatmentCitation id="C78426D0858B4FAB6FE1452DDF8A1737" author="Fischer P &amp; Crosse H" page="592" year="1877">
<bibRefCitation id="629C476BE24D6474A6C96AD9B48BEB81" author="Fischer P &amp; Crosse H" firstAuthor="Fischer" journalOrPublisher="Recherches Zoologiques, Partie 7, Volume 1" page="592" refId="B33" refString="Fischer P, Crosse H (1877) Études sur les mollusques terrestres et fluviatiles du Mexique et du Guatemala. Mission scientifique au Mexique et dans ľAmerique Centrale. Recherches Zoologiques, Partie 7, Volume 1, 702 pp. https://www.biodiversitylibrary.org/item/216870" title="Études sur les mollusques terrestres et fluviatiles du Mexique et du Guatemala. Mission scientifique au Mexique et dans ľAmerique Centrale." url="https://www.biodiversitylibrary.org/item/216870" year="1877">Fischer and Crosse 1877: 592</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="22F787152213CF7B9387D704CEC05A69" author="Pilsbry HA" page="122" year="1906">
<bibRefCitation id="9BB9A425597FAFC3DAD65D302DEFB71F" DOI="10.1175/1520-0493(1907)35%3C357a:LP%3E2.0.CO;2" author="Pilsbry HA" firstAuthor="Pilsbry" page="122" refId="B86" refString="Pilsbry HA (1906) [19061907] Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2, Pulmonata. Volume 18: Achatinidae: Stenogyrinae and Coeliaxinae. Conchological Section, Academy of Natural Sciences of Philadelphia, Philadelphia, 1357. [pls 151] [pp 1272, pls 134 (1906); pp 273357, pls 3551 (1907)] https://doi.org/10.1175/1520-0493 (1907) 35 % 3 C 357 a: LP % 3 E 2.0. CO; 2" year="1906">Pilsbry 1906: 122</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="000E83BAEF0916BB3297AE26F953AC9E" author="Gude GK" page="354" year="1914">
<bibRefCitation id="FA60D288D3AECF74A268C84FA14C5516" DOI="10.5962/bhl.title.12891" author="Gude GK" firstAuthor="Gude" journalOrPublisher="Taylor &amp; Francis, London" page="354" refId="B42" refString="Gude GK (1914) Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, 520 pp. https://doi.org/10.5962/bhl.title.12891" title="Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma." year="1914">Gude 1914: 354</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="9E79E93B216C4329EBDC9EF640A6B706" author="Zilch A" page="351" year="1959">
<bibRefCitation id="DC58BB29293EC3882DF2049CEF6F5580" author="Zilch A" editor="Schindewolf OH" firstAuthor="Zilch" journalOrPublisher="Gebrüder Borntraeger, Berlin" page="351" pagination="201-400" refId="B124" refString="Zilch A (1959) Gastropoda, Euthyneura. In: Schindewolf OH (Ed.) Handbuch der Paläozoologie, Band 6, Gastropoda. Gebrüder Borntraeger, Berlin, 201400." title="Gastropoda, Euthyneura." volumeTitle="Handbuch der Paläozoologie, Band 6, Gastropoda." year="1959">Zilch 1959: 351</bibRefCitation>
</treatmentCitation>
, 352.
<treatmentCitation id="4E95A12DDCDB71B2F2A11EEC54EBEB34" author="van Benthem Jutting WSS" page="378" year="1952">
<bibRefCitation id="CFFCC20D83E66DF0A9331BC1A31EC114" author="van Benthem Jutting WSS" firstAuthor="van Benthem Jutting" journalOrPublisher="Treubia" page="378" pagination="291-435" refId="B110" refString="van Benthem Jutting WSS (1952) Systematic studies on the non-marine Mollusca of the Ind-Australian archipelago. Treubia 21: 291435." title="Systematic studies on the non-marine Mollusca of the Ind-Australian archipelago." volume="21" year="1952">van Benthem Jutting 1952: 378</bibRefCitation>
</treatmentCitation>
.
<treatmentCitation id="5EE35B4F5497A76BFC3034E124AF4F40" author="Schileyko AA" page="492" year="1999">
<bibRefCitation id="F962BB8E48F370E25C7E97EE9D0AAE0C" author="Schileyko AA" firstAuthor="Schileyko" page="492" refId="B95" refString="Schileyko AA (1999) Treatise on recent terrestrial pulmonate molluscs Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae. Ruthenica (Supplement 2): 437564." year="1999">Schileyko 1999: 492</bibRefCitation>
</treatmentCitation>
.
</comment>
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</paragraph>
</subSubSection>
<subSubSection id="SECID0E5LBK" type="type species">
<paragraph id="92D2DF0C7CC8272850E2321ECE437878">
<heading id="69670144692C49FF05AC72B2894F6920" reason="title">Type species.</heading>
</paragraph>
<paragraph id="2B28A317779877CFDA590940D3B1E538">
<treatmentCitationGroup id="AD9B906021EEC0AE77D9AF99CD3FD55C">
<taxonomicName id="CAC8038E4B435185737555E2C877EC31" authority="Miller, 1822" authorityName="Miller" authorityYear="1822" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="goodallii">
<emphasis id="A4AEA2EAAB39EFF77642E0B267EF58AF" italics="true">Helix goodallii</emphasis>
<bibRefCitation id="DB3C03E66F2C34F2E254DE4772AE76E9" author="Miller JS" issue="5" journalOrPublisher="Annals of Philosophy, New Series" pagination="376-381" refId="B60" refString="Miller JS (1822) A list of the freshwater and landshells occurring in the environment of Bristol, with observations. Annals of Philosophy, New Series 3 (5): 376 - 381. https: // www. biodiversitylibrary. org / page / 2498134" title="A list of the freshwater and landshells occurring in the environment of Bristol, with observations." url="https://www.biodiversitylibrary.org/page/2498134" volume="3" year="1822">Miller, 1822</bibRefCitation>
</taxonomicName>
[junior homonym of
<bibRefCitation id="2BE5C4A147572F74BB344B2F81986979" author="Férussac AE dAudebard de baron de" firstAuthor="Ferussac" refId="B32" refString="Férussac AE dAudebard de baron de (1821) [18211822] Tableaux systématiques des animaux mollusques suivis ďun Prodrome général pour tous les mollusques ou fluviatiles, vivantes ou fossiles, 114 pp. [pp 1114 (1821); pp i xlvii (1822)] https://www.biodiversitylibrary.org/bibliography/10558" url="https://www.biodiversitylibrary.org/bibliography/10558" year="1821">Férussac (1821)</bibRefCitation>
] accepted as
<taxonomicName id="D2BC1C87BA1FA4118721A5FD93057A61" authority="(Rang, 1831)" baseAuthorityName="Rang" baseAuthorityYear="1831" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="species" species="hannense">
<emphasis id="C54196F74E2807C8D3393584C1838510" italics="true">Opeas hannense</emphasis>
(
<bibRefCitation id="C3B992FAF3E3BCFF9C0D36E725CAB997" author="Rang S" issue="93" journalOrPublisher="Annales des Sciences Naturelles" pagination="5-63" refId="B92" refString="Rang S (1831) Description des coquilles terrestres recueillies pendant un voyage a la cote occidentale dAfrique, et au Bresil. Annales des Sciences Naturelles 24 (93): 5 - 63." title="Description des coquilles terrestres recueillies pendant un voyage à la côte occidentale ďAfrique, et au Brésil." volume="24" year="1831">Rang, 1831</bibRefCitation>
)
</taxonomicName>
, subsequent designation by von Martens in
<treatmentCitation id="74999B7C999ACF5D727A5CD0B4E9729F" author="Albers JC" page="265" year="1860">
<bibRefCitation id="980800F6A4AFD89EED3276F9C4FC3AFF" DOI="10.5962/bhl.title.11218" author="Albers JC" firstAuthor="Albers" journalOrPublisher="Wilhelm Englemann, Leipzig" page="265" refId="B3" refString="Albers JC (1860) Die Heliceen nach Natürlicher Verwandtschaft Systematisch Geordnet, 2 nd edn. (manuscript edited by E. von Martens). Wilhelm Englemann, Leipzig, 359 pp. https://doi.org/10.5962/bhl.title.11218" title="Die Heliceen nach Natürlicher Verwandtschaft Systematisch Geordnet, 2 nd edn. (manuscript edited by E. von Martens)." year="1860">Albers (1860: 265)</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
</subSubSection>
<subSubSection id="SECID0ECNBK" type="diagnosis">
<paragraph id="A426BD0924889FEEAECD73D49F1BDBBE">
<heading id="A7FC2F0EE592AE64B8B704B0DB7FA4E7" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="EECD52BB308AFD2FD6B544B4C69B1C69">Shell small, slender, and conical; spire high, turreted, cylindrically, and gradually attenuated; embryonic whorls smooth; subsequent whorls with fine radial striations or growth lines. Aperture vertical, narrow, oblong, columellar margin expanded, and columella straight or concave. Umbilicus narrowly opened or closed. Penis cylindrical tube with short epiphallus, and flagellum and penial sheath absent; vagina long ~ 1 / 2 of penis length.</paragraph>
</subSubSection>
<subSubSection id="SECID0EHNBK" type="remarks">
<paragraph id="40B9F50FFCE780C3AA9DC2353464C407">
<heading id="08F3D3EB6AADC9C3B6E9B803ADB4BFAD" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="DCEA939D3EDB3BFDCE511BF7CF93BDAC">
<taxonomicName id="A370F1F7EF740769AA89B2151CE547B9" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="D98DC52C1DBFE5C31BAF8023161967A3" italics="true">Opeas</emphasis>
</taxonomicName>
and
<taxonomicName id="4EFF699A8E06D210D8CB8A0CAED9167F" authority="Pilsbry, 1906" authorityName="Pilsbry" authorityYear="1906" class="Gastropoda" family="Achatinidae" genus="Tortaxis" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="5E0FFE59461570499B3564F2A5945B48" italics="true">Tortaxis</emphasis>
<bibRefCitation id="0167EC8983DEAFC9E3F7D91DD405DF76" DOI="10.1175/1520-0493(1907)35%3C357a:LP%3E2.0.CO;2" author="Pilsbry HA" refId="B86" refString="Pilsbry HA (1906) [1906 - 1907] Manual of Conchology, Structural and Systematic, with Illustrations of the Species, Series 2, Pulmonata. Volume 18: Achatinidae: Stenogyrinae and Coeliaxinae. Conchological Section, Academy of Natural Sciences of Philadelphia, Philadelphia, 1 - 357. [pls 1 - 51] [pp 1 - 272, pls 1 - 34 (1906); pp 273 - 357, pls 35 - 51 (1907)] https: // doi. org / 10.1175 / 1520 - 0493 (1907) 35 % 3 C 357 a: LP % 3 E 2.0. CO; 2" year="1906">Pilsbry, 1906</bibRefCitation>
</taxonomicName>
generally resemble one another in having slender conical or cylindrical shells, nearly smooth to fine striations, and flatly convex whorls (Table
<tableCitation id="9C55F1D0693001A01E91D48E6DE2DD56" captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of shell morphology among subulinid genera recorded in Myanmar. The superscript numbers are references; 1 = Pilsbry (1906), 2 = Gude (1914), 3 = Godwin-Austen (1920), 4 = Naggs (1994), 5 = Schileyko (1999), and 6 = Budha et al. (2017)." httpUri="http://table.plazi.org/id/7320BC8BE0D4AF3B31AE4617F5EE0127" tableUuid="7320BC8BE0D4AF3B31AE4617F5EE0127">2</tableCitation>
). However,
<taxonomicName id="03EB1BA74279A1099BA119E20D5EE601" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="6BB8207DBDEFD81A81190062B05AAF76" italics="true">Opeas</emphasis>
</taxonomicName>
has a smaller size, straight columella, and mostly oblong aperture, whereas
<taxonomicName id="33D2A2E18246B5099DC16FD8951A6053" authorityName="Pilsbry" authorityYear="1906" class="Gastropoda" family="Achatinidae" genus="Tortaxis" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="8034E04AB015F149D6A9055086FF24D4" italics="true">Tortaxis</emphasis>
</taxonomicName>
displays mostly larger size, nearly straight or slightly concave columella with a spiral fold, and more or less obliqued aperture. Likewise,
<taxonomicName id="F9D72B820CDADA6E62B638CCC3C4977D" authorityName="Albers" authorityYear="1850" class="Gastropoda" family="Achatinidae" genus="Opeas" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="DE556D1C24F957FD3BC70CC1F9073E69" italics="true">Opeas</emphasis>
</taxonomicName>
is distinctly differentiated from
<taxonomicName id="B56AF2F65EDF317ED243A63EC4552363" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Achatinidae" genus="Bacillum" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="5DB033102AD6682164851C4B4F0911D8" italics="true">Bacillum</emphasis>
</taxonomicName>
in that it is smaller in size, has a straight columella, fine shell sculptures and embryonic whorls which are narrowly rounded. In contrast,
<taxonomicName id="F48F63953085ABDD7AE721A3C4D83725" authorityName="Theobald" authorityYear="1870" class="Gastropoda" family="Achatinidae" genus="Bacillum" kingdom="Animalia" order="Stylommatophora" phylum="Mollusca" rank="genus">
<emphasis id="0C3063CB45CFDDF9B089E41160175D0B" italics="true">Bacillum</emphasis>
</taxonomicName>
exhibits a larger shell size, truncated and concave columella, stronger shell sculptures, and the embryonic whorls cylindrically rounded.
</paragraph>
<paragraph id="B878A0BBF370994CFA408ED6C954EE42">
This genus is distributed in tropical and subtropical regions in Europe, Asia, Africa, and North America, and comprises nearly 200 species (
<bibRefCitation id="B742EC0095E1AF4727D4AB9189DE687D" author="Schileyko" firstAuthor="Schileyko" refId="B95" refString="Schileyko AA (1999) Treatise on recent terrestrial pulmonate molluscs Part 4. Draparnaudiidae, Caryodidae, Macrocyclidae, Acavidae, Clavatoridae, Dorcasiidae, Sculptariidae, Corillidae, Plectopylidae, Megalobulimidae, Strophocheilidae, Cerionidae, Achatinidae, Subulinidae, Glessulidae, Micractaeonidae, Ferrussaciidae. Ruthenica (Supplement 2): 437564." year="1999">Schileyko 1999</bibRefCitation>
;
<bibRefCitation id="99417EA6BA55A7FFF6FF8FAFA35B751D" DOI="10.14284/448" author="MolluscaBase" firstAuthor="MolluscaBase" refId="B61" refString="MolluscaBase eds (2023) MolluscaBase. https://www.molluscabase.org [in 202307 17] https://doi.org/10.14284/448" url="https://www.molluscabase.org" year="2023">MolluscaBase 2023</bibRefCitation>
). In
<collectingCountry id="941CFAD7848D9C435D32BB50886B88E9" name="Myanmar">Myanmar</collectingCountry>
, two species are recorded (
<bibRefCitation id="11B5D8124C750DBB892AFF9AC3A013D9" DOI="10.5962/bhl.title.12891" author="Gude" firstAuthor="Gude" journalOrPublisher="Taylor &amp; Francis, London" refId="B42" refString="Gude GK (1914) Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma. Taylor &amp; Francis, London, 520 pp. https://doi.org/10.5962/bhl.title.12891" title="Mollusca - II. Trochomorphidae - Janellidae. In: Shipley AE, Marshall GAK (Eds) The Fauna of British India, including Ceylon and Burma." year="1914">Gude 1914</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>