194 lines
15 KiB
XML
194 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.692.14706" ID-GBIF-Dataset="81f065e7-a342-495c-9855-21bc26539311" ID-PMC="PMC5672734" ID-Pensoft-Pub="1313-2970-692-1" ID-PubMed="29133987" ID-ZBK="1B7C3F517CF543338EAB1CB1BD9D8A07" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-692-1" ModsDocOrigin="ZooKeys 692" ModsDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" checkinTime="1503352913593" checkinUser="pensoft" docAuthor="Foon, Junn Kitt & Liew, Thor-Seng" docDate="2017" docId="C8E6DCB5D19CE079DE4AFEAA526D5717" docLanguage="en" docName="ZooKeys 692: 1-81" docOrigin="ZooKeys 692" docSource="http://dx.doi.org/10.3897/zookeys.692.14706" docTitle="Alycaeus altis subsp. pirus Moellendorff 1902" docType="treatment" docVersion="5" lastPageNumber="12" masterDocId="FF8E6C15A17CDA48FFA7FFE36C7CFFA2" masterDocTitle="A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia" masterLastPageNumber="81" masterPageNumber="1" pageNumber="9" updateTime="1668164728433" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A review of the land snail genus Alycaeus (Gastropoda, Alycaeidae) in Peninsular Malaysia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Foon, Junn Kitt</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liew, Thor-Seng</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>692</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>81</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.692.14706</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.692.14706</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-692-1</mods:identifier>
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<mods:identifier type="ZBK">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
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<mods:identifier type="ZooBank">1B7C3F517CF543338EAB1CB1BD9D8A07</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="132786247" LSID="urn:lsid:plazi:treatment:C8E6DCB5D19CE079DE4AFEAA526D5717" httpUri="http://treatment.plazi.org/id/C8E6DCB5D19CE079DE4AFEAA526D5717" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
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<subSubSection pageId="8" pageNumber="9" type="nomenclature">
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName authority="Moellendorff, 1902" authorityName="Moellendorff" authorityYear="1902" class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus altis" order="Architaenioglossa" pageId="8" pageNumber="9" phylum="Mollusca" rank="subSpecies" species="altis" subSpecies="pirus">
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Alycaeus altis pirus
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902
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</taxonomicName>
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Figures 9, 31U
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="10" type="reference_group">
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<paragraph pageId="9" pageNumber="10">
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="subSpecies" species="perakensis" subSpecies="altispirus">
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<pageBreakToken pageId="9" pageNumber="10" start="start">Alycaeus</pageBreakToken>
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perakensis altispirus
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</taxonomicName>
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902: 144-145;
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<bibRefCitation pageId="9" pageNumber="10">Maassen 2001</bibRefCitation>
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: 23;
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<bibRefCitation author="Tarruella, A" journalOrPublisher="Spira" pageId="53" pageNumber="54" pagination="71 - 76" title="Listado taxonomico ilustrado de la familia Cyclophoridae J. E. Gray, 1847 (Mollusca: Gastropoda): Parte 2. La subfamilia Alycaeinae J. E. Gray, 1850." url="http://www.molluscat.com/assets/spira_4_12_7.pdf" volume="4" year="2011">
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Tarruella and
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<normalizedToken originalValue="Domènech">Domenech</normalizedToken>
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2011
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</bibRefCitation>
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: 72.
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</paragraph>
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<paragraph pageId="9" pageNumber="10">
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus (Alycaeus) perakensis" order="Architaenioglossa" pageId="9" pageNumber="10" phylum="Mollusca" rank="subSpecies" species="perakensis" subGenus="Alycaeus" subSpecies="altispirus">Alycaeus (Alycaeus) perakensis altispirus</taxonomicName>
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:
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<bibRefCitation pageId="9" pageNumber="10">Zilch 1957</bibRefCitation>
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: 147, plate 6, figure 32.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="10" type="type locality">
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<paragraph pageId="9" pageNumber="10">Type locality.</paragraph>
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<paragraph pageId="9" pageNumber="10">Kelantan.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="type material">
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<paragraph pageId="10" pageNumber="11">
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<pageBreakToken pageId="10" pageNumber="11" start="start">Type</pageBreakToken>
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material.
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</paragraph>
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName genus="Lectotype" lsidName="Lectotype" pageId="10" pageNumber="11" rank="genus">Lectotype</taxonomicName>
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. Kelantan: SMF 109738 (Seen). Paralectotypes. Kelantan: SMF 109739/3(Not seen).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="description">
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<paragraph pageId="10" pageNumber="11">Description.</paragraph>
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<paragraph pageId="10" pageNumber="11">Protoconch. Smooth.</paragraph>
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<paragraph pageId="10" pageNumber="11">Shell shape. Conical. Shell height: 6.40 mm. Shell width: 6.30 mm.</paragraph>
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<paragraph pageId="10" pageNumber="11">
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Spire. Spire height: 2.37 mm. Spire width: 2.68 mm. Number of whorls: up to 5
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<normalizedToken originalValue="½">1/2</normalizedToken>
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. Spire shape: oblong conical. Whorl periphery rounded. Umbilicus open.
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</paragraph>
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<paragraph pageId="10" pageNumber="11">
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Whorl constriction. At about 5
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<normalizedToken originalValue="¼">1/4</normalizedToken>
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whorls posterior of protoconch.
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</paragraph>
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<paragraph pageId="10" pageNumber="11">Breathing tube. Length: 0.77 mm.</paragraph>
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<paragraph pageId="10" pageNumber="11">Aperture and peristome. Aperture circular, moderately expanded. Aperture height: 2.87 mm. Aperture width: 3.36 mm. Peristome double, not thickened, notched at suture, upper palatal section folded posteriorly into a wing-like structure. Peristome orientation 36° oblique with respect to the coiling axis.</paragraph>
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<paragraph pageId="10" pageNumber="11">Spiral lines. Absent.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<pageBreakToken pageId="11" pageNumber="12" start="start">Radial</pageBreakToken>
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ribs running anterior of breathing tube. Radial ribs pronounced, evenly spaced. Approximately 8 ribs per 1 mm.
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</paragraph>
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<paragraph pageId="11" pageNumber="12">Radial ribs running perpendicular to breathing tube. Radial ribs pronounced, white and thicker than those anterior of breathing tube, evenly spaced. Approximately 16 ribs per 1 mm.</paragraph>
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<paragraph pageId="11" pageNumber="12">Radial ribs running posterior of breathing tube. Radial ribs absent immediately posterior of constriction, becoming pronounced and evenly spaced from the middle section until prior to aperture. Approximately 10 ribs per 1 mm.</paragraph>
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<paragraph pageId="11" pageNumber="12">Operculum. Unknown.</paragraph>
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<paragraph pageId="11" pageNumber="12">Shell colour. Whorls yellow. Peristome white.</paragraph>
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<paragraph pageId="11" pageNumber="12">Living animal. Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="habitat">
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<paragraph pageId="11" pageNumber="12">Habitat and ecology.</paragraph>
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<paragraph pageId="11" pageNumber="12">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="distribution">
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<paragraph pageId="11" pageNumber="12">Distribution range.</paragraph>
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<paragraph pageId="11" pageNumber="12">Unknown. Appears to be restricted to Kelantan.</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="differential diagnosis">
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<paragraph pageId="11" pageNumber="12">Differential diagnosis.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus altispirus" order="Architaenioglossa" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="altispirus">Alycaeus altispirus</taxonomicName>
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is most similar to
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus regalis" order="Architaenioglossa" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="regalis">Alycaeus regalis</taxonomicName>
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sp. n. in having a large shell (shell height: 6.40-7.18 mm, shell width: 5.86-6.87 mm) and tall spire (spire height: 2.32-2.90 mm) but differs in the ultimate whorl being more keeled-like, upper palatal section folded posteriorly into a wing-like structure and peristome orientated oblique to the coiling axis. Unlike
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus perakensis" order="Architaenioglossa" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="perakensis">Alycaeus perakensis</taxonomicName>
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,
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus altispirus" order="Architaenioglossa" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="altispirus">Alycaeus altispirus</taxonomicName>
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is smaller (smaller by about 0.23 mm in shell height, 0.25 mm in shell width) and has a much less expanded ultimate whorl.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="11" pageNumber="12" type="discussion">
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<paragraph pageId="11" pageNumber="12">Discussion.</paragraph>
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<paragraph pageId="11" pageNumber="12">
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To date, this species is only known from the four type specimens collected by John Waterstradt (
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<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="52" pageNumber="53" pagination="135 - 149" title="Binnenmollusken aus Hinterindien 1. Landschnecken von Kelantan, Ostkueste der Halbinsel Malacca." url="http://www.biodiversitylibrary.org/part/145333/summary" volume="34" year="1902">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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1902
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</bibRefCitation>
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,
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<bibRefCitation pageId="11" pageNumber="12">Zilch 1957</bibRefCitation>
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) presumably from the same locality as
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<taxonomicName lsidName="A. kelantanensis" pageId="11" pageNumber="12" rank="species" species="kelantanensis">A. kelantanensis</taxonomicName>
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, on Pulai Princess Cave hill (
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<geoCoordinate direction="north" orientation="latitude" precision="15" value="4.793889">4°47'38"N</geoCoordinate>
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,
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<geoCoordinate direction="east" orientation="longitude" precision="15" value="101.94195">101°56'31"E</geoCoordinate>
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), Kelantan (see Discussion under
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<taxonomicName lsidName="A. kelantanensis" pageId="11" pageNumber="12" rank="species" species="kelantanensis">A. kelantanensis</taxonomicName>
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;
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<bibRefCitation author="Waterstradt, J" journalOrPublisher="Journal of the Straits Branch of the Royal Asiatic Society" pageId="54" pageNumber="55" pagination="1 - 27" title="Kelantan and my trip to Gunong Tahan." url="http://www.biodiversitylibrary.org/part/145327/summary" volume="37" year="1902">Waterstradt 1902</bibRefCitation>
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,
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<bibRefCitation author="Liew, T" journalOrPublisher="ZooKeys" pageId="51" pageNumber="52" pagination="1 - 107" title="A cybertaxonomic revision of the micro-landsnail genus Plectostoma Adam (Mollusca, Caenogastropoda, Diplommatinidae), from Peninsular Malaysia, Sumatra and Indochina." url="https://doi.org/10.3897/zookeys.393.6717" volume="393" year="2014">Liew et al. 2014</bibRefCitation>
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). Yet no recent shells have been found on surveyed limestone hills near the locality (about 5 km from Pulai Princess Cave hill) or elsewhere in Kelantan. At the time of its description,
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<bibRefCitation author="Moellendorff, von OF" journalOrPublisher="Nachrichtsblatt der Deutschen Malakozoologischen Gesellschaft" pageId="52" pageNumber="53" pagination="135 - 149" title="Binnenmollusken aus Hinterindien 1. Landschnecken von Kelantan, Ostkueste der Halbinsel Malacca." url="http://www.biodiversitylibrary.org/part/145333/summary" volume="34" year="1902">
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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(1902)
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</bibRefCitation>
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considered
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<taxonomicName lsidName="A. altispirus" pageId="11" pageNumber="12" rank="species" species="altispirus">A. altispirus</taxonomicName>
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a subspecies of
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<taxonomicName lsidName="A. perakensis" pageId="11" pageNumber="12" rank="species" species="perakensis">A. perakensis</taxonomicName>
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but did not elaborate why. We disagree with this decision because the two taxa differ markedly from each other in shell shape especially the degree of expansion and shape of the ultimate whorl as well as their very disjunct distribution range. As such, we consider
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<taxonomicName lsidName="A. altispirus" pageId="11" pageNumber="12" rank="species" species="altispirus">A. altispirus</taxonomicName>
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a species on its own.
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</paragraph>
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<caption pageId="11" pageNumber="12">
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<paragraph pageId="11" pageNumber="12">
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Figure 9.
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<taxonomicName class="Gastropoda" family="Alycaeidae" genus="Alycaeus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Alycaeus altispirus" order="Architaenioglossa" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="altispirus">Alycaeus altispirus</taxonomicName>
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(
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1902)
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<normalizedToken originalValue="A–E">A-E</normalizedToken>
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Shell of lectotype, SMF 109738
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<normalizedToken originalValue="F–H">F-H</normalizedToken>
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Close up of shell of lectotype, SMF 109738. Scale bars:
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<normalizedToken originalValue="A–G">A-G</normalizedToken>
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1 mm; H 0.5 mm. All photographs by
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<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
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Barna, courtesy of R. Janssen, SMF.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |