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<document id="0D51ED8A7E0C46DFAD17235EFCB68040" ID-DOI="10.37520/fi.2024.006" ID-ISSN="2533-4069" ID-Zenodo-Dep="14722393" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1737336788380" checkinUser="felipe" docAuthor="Cleal, Christopher J., Strullu-Derrien, Christine &amp; Spencer, Alan R. T." docDate="2024" docId="03A187BCB029FFB2748CF94A54D2F9D2" docLanguage="en" docName="FossilImprint.80.1.35-67.pdf" docOrigin="Fossil Imprint 80 (1)" docStyle="DocumentStyle:0EC54BF35E227CA4627F894F6BA30634.4:FossilImprint.2016-.journal_article" docStyleId="0EC54BF35E227CA4627F894F6BA30634" docStyleName="FossilImprint.2016-.journal_article" docStyleVersion="4" docTitle="Calymmotheca divaricata STUR 1877" docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="FF98FFC4B039FFA0771DFFD7564FFFC3" masterDocTitle="Early Coal Swamp Vegetation From The Serpukhovian Lower Clackmannan Group Of Scotland" masterLastPageNumber="67" masterPageNumber="35" pageNumber="51" updateTime="1737596041418" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="B4085D109E5463F456DA589E238BF610">Early Coal Swamp Vegetation From The Serpukhovian Lower Clackmannan Group Of Scotland</mods:title>
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<mods:affiliation id="8E2FE238D2DEC3F352E7BB24EDF6ACEA">Department of Earth Sciences, University of Bristol, Bristol, BS 8 1 TQ, United Kingdom; email: christopher. cleal @ bristol. ac. uk.</mods:affiliation>
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<mods:namePart id="8EC0F88EAF8E0193FD265C3E55BC3C45">Strullu-Derrien, Christine</mods:namePart>
<mods:affiliation id="9A4A821EED6649ACB74E147C25B511E9">Institut de Systématique, Évolution, Biodiversité (UMR 7205), Muséum National dHistoire Naturelle, CNRS, Sorbonne Université, Paris, France. &amp; Science Group, The Natural History Museum, London, United Kingdom.</mods:affiliation>
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<mods:namePart id="F59E90212D5E5E750B577E9CF09D158F">Spencer, Alan R. T.</mods:namePart>
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<subSubSection id="C3126521B029FFB0748CF94A53CCF976" box="[913,1411,1693,1717]" pageId="16" pageNumber="51" type="nomenclature">
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<heading id="D0FF81C6B029FFB0748CF94A53CCF976" box="[913,1411,1693,1717]" centered="true" fontSize="8" level="2" pageId="16" pageNumber="51" reason="2">
<taxonomicName id="4C084D29B029FFB0748CF94A53CCF976" authority="(GOPP.) STUR, 1877" authorityName="STUR" authorityYear="1877" baseAuthorityName="GOPP." box="[913,1411,1693,1717]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B029FFB0748CF94A53CCF976" bold="true" box="[913,1411,1693,1717]" pageId="16" pageNumber="51">
<emphasis id="B97CEAB8B029FFB0748CF94A52E4F976" bold="true" box="[913,1195,1693,1717]" italics="true" pageId="16" pageNumber="51">Calymmotheca divaricata</emphasis>
(
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.)
<bibRefCitation id="EF994B5BB029FFB07217F94A530FF970" author="Stur, D." box="[1290,1344,1693,1717]" pageId="16" pageNumber="51" pagination="107 - 472" refId="ref24329" refString="Stur, D. (1877): Die Culm-Flora der Ostrauer und Waldenburger Schichten. - Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 8 (2): 107 - 472." type="journal article" year="1877">
<smallCapsWord id="8D51A076B029FFB07217F94A530FF970" baselines="1711,1711" box="[1290,1344,1693,1717]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Stur" pageId="16" pageNumber="51">STUR</smallCapsWord>
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, 1877
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<paragraph id="8BB736AAB029FFB0734EF969528EF915" blockId="16.[1107,1217,1726,1750]" box="[1107,1217,1726,1750]" pageId="16" pageNumber="51">
<figureCitation id="13332A2FB029FFB0734EF969528EF915" box="[1107,1217,1726,1750]" captionStart="Text-fig" captionStartId="17.[125,206,1890,1911]" captionTargetBox="[125,1440,151,1867]" captionTargetId="figure-6@17.[124,783,627,1105]" captionTargetPageId="17" captionText="Text-fig. 9. Calymmotheca divaricata (GÖPP.) STUR. a: Wimbledon Pit, Machrihanish, Campbeltown, Mull of Kintyre; shale above Main Coal, Limestone Coal Formation; BGS Kidst.3202. b, c: Machrihanish Water, 290 m west of Wimbledon Pit, Campbeltown, Mull of Kintyre; Limestone Coal Formation; BGS Kidst.3210 (b), BGS Kidst.4488 (c). Scale bars = 10 mm." figureDoi="http://doi.org/10.5281/zenodo.14722409" httpUri="https://zenodo.org/record/14722409/files/figure.png" pageId="16" pageNumber="51">Text-fig. 9</figureCitation>
</paragraph>
<paragraph id="8BB736AAB029FFB07456F93B53E1F8C1" blockId="16.[843,1482,1772,2092]" box="[843,1454,1772,1794]" pageId="16" pageNumber="51">
1836
<taxonomicName id="4C084D29B029FFB0748BF93B528AF8C1" authority="GOPP." authorityName="GOPP." box="[918,1221,1772,1794]" class="Polypodiopsida" genus="Cheilanthites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="divaricatus">
<emphasis id="B97CEAB8B029FFB0748BF93B52C7F8C2" box="[918,1160,1772,1793]" italics="true" pageId="16" pageNumber="51">Cheilanthites divaricatus</emphasis>
<smallCapsWord id="8D51A076B029FFB07393F93A52F0F8C1" baselines="1789,1790" box="[1166,1215,1773,1794]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gopp" pageId="16" pageNumber="51">GÖPP</smallCapsWord>
.
</taxonomicName>
, p. 238, pl. 12, figs 1, 2.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F8DD530BF8DC" blockId="16.[843,1482,1772,2092]" box="[843,1348,1802,1824]" pageId="16" pageNumber="51">
1836
<taxonomicName id="4C084D29B029FFB0748BF8DD52CDF8E3" authority="GOPP." authorityName="GOPP." box="[918,1154,1802,1824]" class="Polypodiopsida" family="Gleicheniaceae" genus="Gleichenites" higherTaxonomySource="GBIF" kingdom="Plantae" order="Gleicheniales" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="linkii">
<emphasis id="B97CEAB8B029FFB0748BF8DD520AF8DC" box="[918,1093,1802,1823]" italics="true" pageId="16" pageNumber="51">Gleichenites linkii</emphasis>
<smallCapsWord id="8D51A076B029FFB07351F8DD5233F8DC" baselines="1818,1819" box="[1100,1148,1802,1823]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gopp" pageId="16" pageNumber="51">GÖPP</smallCapsWord>
.
</taxonomicName>
, p. 182, pl. 2, fig. 1.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F8FF5338F8FE" blockId="16.[843,1482,1772,2092]" box="[843,1399,1832,1854]" pageId="16" pageNumber="51">
1875
<taxonomicName id="4C084D29B029FFB0748BF8FF528DF8FF" authority="STUR" authorityName="STUR (Stur" authorityYear="1875" box="[918,1218,1832,1853]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="falkenhainii">
<emphasis id="B97CEAB8B029FFB0748BF8FF52C2F8FE" box="[918,1165,1832,1853]" italics="true" pageId="16" pageNumber="51">Sphenopteris falkenhainii</emphasis>
<smallCapsWord id="8D51A076B029FFB0738FF8FF528DF8FF" baselines="1848,1848" box="[1170,1218,1832,1853]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Stur" pageId="16" pageNumber="51">STUR</smallCapsWord>
</taxonomicName>
, p. 26, pl. 6, fig. 1.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F89155A6F8BA" blockId="16.[843,1482,1772,2092]" pageId="16" pageNumber="51">
1877
<taxonomicName id="4C084D29B029FFB0748BF891536DF89A" authority="(GOPP.) STUR" authorityName="STUR" baseAuthorityName="GOPP." box="[918,1314,1862,1883]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B029FFB0748BF89152DAF898" box="[918,1173,1862,1883]" italics="true" pageId="16" pageNumber="51">Calymmotheca divaricata</emphasis>
(
<smallCapsWord id="8D51A076B029FFB073B4F8915296F898" baselines="1878,1879" box="[1193,1241,1862,1883]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gopp" pageId="16" pageNumber="51">GÖPP</smallCapsWord>
.)
<smallCapsWord id="8D51A076B029FFB073EEF891536DF89A" baselines="1878,1877" box="[1267,1314,1862,1883]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Stur" pageId="16" pageNumber="51">STUR</smallCapsWord>
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, p. 165, pl. 13, figs 13.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F8565581F877" blockId="16.[843,1482,1772,2092]" pageId="16" pageNumber="51">
1923
<taxonomicName id="4C084D29B029FFB0748BF8565281F856" authority="STUR" authorityName="STUR (Stur" authorityYear="1875" box="[918,1230,1921,1942]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="falkenhainii">
<emphasis id="B97CEAB8B029FFB0748BF85652DCF855" box="[918,1171,1921,1942]" italics="true" pageId="16" pageNumber="51">Sphenopteris falkenhainii</emphasis>
<smallCapsWord id="8D51A076B029FFB07382F8565281F856" baselines="1937,1937" box="[1183,1230,1921,1942]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Stur" pageId="16" pageNumber="51">STUR</smallCapsWord>
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; Kidston, p. 79, pl. 19, fig. 1.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F86A5263F833" blockId="16.[843,1482,1772,2092]" pageId="16" pageNumber="51">
1923
<taxonomicName id="4C084D29B029FFB0748BF86A52FAF811" authority="KIDST." authorityName="KIDST." box="[918,1205,1981,2002]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="fragilis">
<emphasis id="B97CEAB8B029FFB0748BF86A5227F811" box="[918,1128,1981,2002]" italics="true" pageId="16" pageNumber="51">Sphenopteris fragilis</emphasis>
<smallCapsWord id="8D51A076B029FFB07365F86A52E1F813" baselines="1997,1996" box="[1144,1198,1981,2002]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kidst" pageId="16" pageNumber="51">KIDST</smallCapsWord>
.
</taxonomicName>
(non Schlotheim), p. 86, pl. 17, figs 15.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F82F53C5F7CE" blockId="16.[843,1482,1772,2092]" box="[843,1418,2040,2062]" pageId="16" pageNumber="51">
1923
<taxonomicName id="4C084D29B029FFB0748BF82F52D0F7CD" authority="KIDST." authorityName="KIDST." box="[918,1183,2040,2062]" class="Pinopsida" genus="Schuetzia" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="bennieana">
<emphasis id="B97CEAB8B029FFB0748BF82F5213F7CE" box="[918,1116,2040,2061]" italics="true" pageId="16" pageNumber="51">Schuetzia bennieana</emphasis>
<smallCapsWord id="8D51A076B029FFB0737FF82E52D6F7CF" baselines="2057,2056" box="[1122,1177,2041,2062]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kidst" pageId="16" pageNumber="51">KIDST</smallCapsWord>
.
</taxonomicName>
; Kidston, pl. 107, fig. 8.
</paragraph>
<paragraph id="8BB736AAB029FFB07456F7C15385F7E8" blockId="16.[843,1482,1772,2092]" box="[843,1482,2070,2092]" pageId="16" pageNumber="51">
1924
<taxonomicName id="4C084D29B029FFB0748BF7C152F4F7E8" authority="KIDST." authorityName="KIDST." box="[918,1211,2070,2092]" class="Pinopsida" genus="Schuetzia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Plantae" pageId="16" pageNumber="51" phylum="Tracheophyta" rank="species" species="bennieana">
<emphasis id="B97CEAB8B029FFB0748BF7C155BDF7E8" box="[918,1010,2070,2091]" italics="true" pageId="16" pageNumber="51">Schuetzia</emphasis>
cf.
<emphasis id="B97CEAB8B029FFB0730BF7C15235F7E8" box="[1046,1146,2070,2091]" italics="true" pageId="16" pageNumber="51">bennieana</emphasis>
<smallCapsWord id="8D51A076B029FFB07362F7C152FAF7EA" baselines="2086,2085" box="[1151,1205,2070,2091]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Kidst" pageId="16" pageNumber="51">KIDST</smallCapsWord>
.
</taxonomicName>
; Kidston, pl. 107, figs 913.
</paragraph>
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<paragraph id="8BB736AAB028FFB17760F8B552F0F871" blockId="17.[125,1440,1890,1970]" pageId="17" pageNumber="52">
<emphasis id="B97CEAB8B028FFB17760F8B556A5F8B4" bold="true" box="[125,234,1890,1911]" pageId="17" pageNumber="52">Text-fig. 9.</emphasis>
<taxonomicName id="4C084D29B028FFB177EDF8B55435F8B4" authority="(GOPP.) STUR." authorityName="STUR." baseAuthorityName="GOPP." box="[240,634,1890,1911]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="17" pageNumber="52" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B028FFB177EDF8B5543AF8B5" bold="true" box="[240,629,1890,1911]" pageId="17" pageNumber="52">
<emphasis id="B97CEAB8B028FFB177EDF8B557A2F8B4" bold="true" box="[240,493,1890,1911]" italics="true" pageId="17" pageNumber="52">Calymmotheca divaricata</emphasis>
(
<smallCapsWord id="8D51A076B028FFB176E6F8B5547FF8B5" baselines="1906,1906" box="[507,560,1890,1911]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Gopp" pageId="17" pageNumber="52">GÖPP</smallCapsWord>
.)
<smallCapsWord id="8D51A076B028FFB1755EF8B5543AF8B5" baselines="1906,1906" box="[579,629,1890,1911]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Stur" pageId="17" pageNumber="52">STUR</smallCapsWord>
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.
</taxonomicName>
a: Wimbledon Pit, Machrihanish, Campbeltown, Mull of Kintyre; shale above Main Coal, Limestone Coal Formation; BGS Kidst.3202. b, c: Machrihanish Water, 290 m west of Wimbledon Pit, Campbeltown, Mull of Kintyre; Limestone Coal Formation; BGS Kidst.3210 (b), BGS Kidst.4488 (c). Scale bars = 10 mm.
</paragraph>
</caption>
<paragraph id="8BB736AAB02BFFB277BBFF4F544FFDB2" blockId="18.[129,768,151,625]" pageId="18" pageNumber="53">
D e s c r i p t i o n. Bifurcate fronds attached to stems up to
<quantity id="4CF09B4FB02BFFB277D2FF60576CFF0C" box="[207,291,183,207]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="53" unit="mm" value="15.0">15 mm</quantity>
wide with elongate-anastomosed surface markings. Primary rachis ca.
<quantity id="4CF09B4FB02BFFB276A1FF0F57B6FF2C" box="[444,505,216,240]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
wide, bifurcating at 35° about
<quantity id="4CF09B4FB02BFFB277DEFF2F5741FECC" box="[195,270,248,272]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="53" unit="mm" value="40.0">40 mm</quantity>
away from the attachment to the stem to
<taxonomicName id="4C084D29B02BFFB275D6FF2F56ABFEF3" form="bipinnate" pageId="18" pageNumber="53" rank="form">form bipinnate</taxonomicName>
to tripinnate primary rachis branches. Gentlytapered ultimate pinnae attached at 7090°, oppositely or suboppositely arranged, spaced up to
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apart. Pinnules up to
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long and
<quantity id="4CF09B4FB02BFFB276FDFEAF546FFE53" box="[480,544,376,400]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="18" pageNumber="53" unit="mm" value="3.0">3 mm</quantity>
wide, alternately to suboppositely arranged, and somewhat obliquely (7080°) attached to rachis by a broad but short petiole; pinnule shape deltoid to subrhomboidal. The most proximal pinnules are deeply incised with five to seven cuneate lobes, with each lobe sometimes being further shallowly divided; more distal pinnules are less divided and at the ends of the ultimate pinnae they become simple or bifid.
</paragraph>
<paragraph id="8BB736AAB02BFFB277BBFD5F5482FBC3" blockId="18.[128,768,648,1410]" pageId="18" pageNumber="53">
R e m a r k s. This fossil-species is widespread in the upper Visean and Serpukhovian of Europe (
<bibRefCitation id="EF994B5BB02BFFB2759FFD7F56F4FD23" author="Mosseichik" pageId="18" pageNumber="53" pagination="1 - 27" refId="ref22403" refString="Mosseichik [= Moseychik], Yu. V. (2010): Rannekamennougol'nye fitokhorii Severnoy Evrazii: struktura, sistema, evolyutsiya [Early Carboniferous phytochoria of the North Eurasia: structure, system, evolution]. - Lethaea rossica, 2: 1 - 27. (in Russian with English summary)" type="journal article" year="2010">Mosseichik 2010</bibRefCitation>
). Historically it has often been referred to as “
<taxonomicName id="4C084D29B02BFFB27791FD3F57A5FCC3" authority="SCHLOTH." authorityName="SCHLOTH." box="[140,490,744,768]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="fragilis">
<emphasis id="B97CEAB8B02BFFB27791FD3F5738FCC3" box="[140,375,744,768]" italics="true" pageId="18" pageNumber="53">Sphenopteris fragilis</emphasis>
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.
</taxonomicName>
”. That name was not validly published as
<bibRefCitation id="EF994B5BB02BFFB27670FCDF5403FCE3" author="Schlotheim, E. F. F. von" box="[365,588,776,800]" pageId="18" pageNumber="53" refId="ref23604" refString="Schlotheim, E. F. F. von (1820): Die Petrefactenkunde auf ihrem jetzigen Standpunkte durch die Beschreibung seiner Sammlung versteinerter und fossiler Uberreste des Thier und Pflan- zenreichs der Vorwelt erlautert. - Becker, Gotha, 504 pp." type="book" year="1820">Schlotheims (1820)</bibRefCitation>
work pre-dated the starting point for palaeobotanical nomenclature but was subsequently validated by
<bibRefCitation id="EF994B5BB02BFFB276B0FC9F5438FCA2" author="Brongniart, A." box="[429,631,840,865]" pageId="18" pageNumber="53" refId="ref16202" refString="Brongniart, A. (1828): Prodrome d'une histoire des vegetaux fossiles. - F. G. Levrault, Paris, 244 pp. https: // doi. org / 10.5962 / bhl. title. 62840" type="book" year="1828">Brongniart (1828)</bibRefCitation>
. As pointed out by
<bibRefCitation id="EF994B5BB02BFFB277CFFCBF5444FC42" author="Strullu-Derrien, C. &amp; Cleal, C. J. &amp; Ducassou, C. &amp; Spencer, A. R. T. &amp; Stolle, E. &amp; Leshyk, V. O." box="[210,523,872,897]" pageId="18" pageNumber="53" refId="ref24107" refString="Strullu-Derrien, C., Cleal, C. J., Ducassou, C., Spencer, A. R. T., Stolle, E., Leshyk, V. O. (2021): A rare late Mississippian flora from Northwestern Europe (Maine-et- - Loire Coalfield, Pays de la Loire, France). - Review of Palaeobotany and Palynology, 285: 104359 (23 pp.). https: // doi. org / 10.1016 / j. revpalbo. 2020.104359" type="book" year="2021">Strullu-Derrien et al. (2021)</bibRefCitation>
, however, the
<typeStatus id="54B38808B02BFFB275ACFCBE5490FC42" box="[689,735,873,897]" pageId="18" pageNumber="53">type</typeStatus>
of that name (
<bibRefCitation id="EF994B5BB02BFFB277E1FC5E57F5FC62" author="Schlotheim, E. F. F. von" box="[252,442,905,929]" pageId="18" pageNumber="53" refId="ref23569" refString="Schlotheim, E. F. F. von (1804): Beschreibung merkwurdi- ger Krauter-Abdrucke und Pflanzen-Versteinerungen: ein Beitrag zur Flora der Vorwelt. - Becker, Gotha, 110 pp." type="book" year="1804">Schlotheim 1804</bibRefCitation>
: pl. 10, fig. 17) is difficult to interpret and the earliest legitimate name with a
<typeStatus id="54B38808B02BFFB27582FC7E5482FC02" box="[671,717,937,961]" pageId="18" pageNumber="53">type</typeStatus>
that can definitely be assigned to this species is
<taxonomicName id="4C084D29B02BFFB2756CFC1E56B6FBC2" class="Polypodiopsida" genus="Cheilanthites" higherTaxonomySource="GBIF" kingdom="Plantae" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="divaricatus">
<emphasis id="B97CEAB8B02BFFB2756CFC1E56B6FBC2" italics="true" pageId="18" pageNumber="53">Cheilanthites divaricatus</emphasis>
</taxonomicName>
the basionym of
<taxonomicName id="4C084D29B02BFFB276AFFC3E5489FBC2" box="[434,710,1001,1025]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B02BFFB276AFFC3E5489FBC2" box="[434,710,1001,1025]" italics="true" pageId="18" pageNumber="53">Calymmotheca divaricata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736AAB02BFFB277BBFBDE549FFB22" blockId="18.[128,768,648,1410]" pageId="18" pageNumber="53">
<bibRefCitation id="EF994B5BB02BFFB277BBFBDE5702FBE2" author="Kidston, R." box="[166,333,1033,1057]" pageId="18" pageNumber="53" pagination="1 - 376" refId="ref21245" refString="Kidston, R. (1923): Fossil plants of the Carboniferous rocks of Great Britain. Parts 1 - 4. - Memoirs of the Geological Survey of Great Britain, Palaeontology, 2: 1 - 376." type="journal article" year="1923">Kidston (1923</bibRefCitation>
: pl. 17, figs 15) figured specimens of this species from the Upper Limestone Formation as
<taxonomicName id="4C084D29B02BFFB2779CFB9E5734FBA2" authority="BRONGN. He" authorityName="BRONGN. He" box="[129,379,1097,1121]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="fragilis">
<emphasis id="B97CEAB8B02BFFB2779CFB9E56A3FBA2" box="[129,236,1097,1121]" italics="true" pageId="18" pageNumber="53">S. fragilis</emphasis>
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. He
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also (
<bibRefCitation id="EF994B5BB02BFFB276DFFB9E5413FBA2" author="Kidston, R." box="[450,604,1097,1121]" pageId="18" pageNumber="53" pagination="1 - 376" refId="ref21245" refString="Kidston, R. (1923): Fossil plants of the Carboniferous rocks of Great Britain. Parts 1 - 4. - Memoirs of the Geological Survey of Great Britain, Palaeontology, 2: 1 - 376." type="journal article" year="1923">Kidston 1923</bibRefCitation>
: pl. 19, fig. 1) figured a specimen with somewhat less divided pinnules as
<taxonomicName id="4C084D29B02BFFB2779DFB5E542BFB62" authority="STUR (Stur 1875)" authorityName="STUR (Stur" authorityYear="1875" box="[128,612,1161,1185]" class="Pteridospermopsida" family="Lyginopteridaceae" genus="Sphenopteris" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lyginopteridales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="falkenhainii">
<emphasis id="B97CEAB8B02BFFB2779DFB5E57D8FB62" box="[128,407,1161,1185]" italics="true" pageId="18" pageNumber="53">Sphenopteris falkenhainii</emphasis>
<smallCapsWord id="8D51A076B02BFFB276BEFB5E5798FB5C" baselines="1179,1179" box="[419,471,1161,1185]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Stur" pageId="18" pageNumber="53">STUR</smallCapsWord>
(
<bibRefCitation id="EF994B5BB02BFFB276F6FB5E5413FB62" author="Stur, D." box="[491,604,1161,1185]" pageId="18" pageNumber="53" pagination="1 - 106" refId="ref24299" refString="Stur, D. (1875): Die Culm-Flora des mahrisch-schlesischen Dachschiefers. - Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 8 (1): 1 - 106." type="journal article" year="1875">Stur 1875</bibRefCitation>
)
</taxonomicName>
but
<bibRefCitation id="EF994B5BB02BFFB27580FB5E5687FB02" author="Patteisky, K." pageId="18" pageNumber="53" pagination="59 - 83" refId="ref22770" refString="Patteisky, K. (1957): Die phylogenetische Entwicklung der Arten von Lyginopteris und ihre Bedeutung fur die Stratigraphie. - Mitteilungen der Westfalischen Berggewerkschaftskasse, 12: 59 - 83." type="journal article" year="1957">Patteisky (1957)</bibRefCitation>
regarded this species as being merely a variety of
<taxonomicName id="4C084D29B02BFFB2779DFB1D5741FB22" box="[128,270,1225,1249]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B02BFFB2779DFB1D5741FB22" box="[128,270,1225,1249]" italics="true" pageId="18" pageNumber="53">C. divaricata</emphasis>
</taxonomicName>
and the distinction is not recognised here.
</paragraph>
<paragraph id="8BB736AAB02BFFB277BBFB3E575BFA42" blockId="18.[128,768,648,1410]" pageId="18" pageNumber="53">
Clusters of sporangia associated with
<taxonomicName id="4C084D29B02BFFB27574FB3D554FFAC2" box="[617,768,1257,1281]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B02BFFB27574FB3D554FFAC2" box="[617,768,1257,1281]" italics="true" pageId="18" pageNumber="53">C. divaricata</emphasis>
</taxonomicName>
fronds were described by
<bibRefCitation id="EF994B5BB02BFFB276BEFADD5406FAE1" author="Kidston, R." box="[419,585,1290,1314]" pageId="18" pageNumber="53" pagination="377 - 522" refId="ref21286" refString="Kidston, R. (1924): Fossil plants of the Carboniferous rocks of Great Britain. Part 5. - Memoirs of the Geological Survey of Great Britain, Palaeontology, 2: 377 - 522." type="journal article" year="1924">Kidston (1924)</bibRefCitation>
as
<taxonomicName id="4C084D29B02BFFB2756EFADD56A0FA81" class="Pinopsida" genus="Schuetzia" higherTaxonomySource="GBIF" isUncertain="true" kingdom="Plantae" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="bennieana">
<emphasis id="B97CEAB8B02BFFB2756EFADD5496FAE1" box="[627,729,1290,1314]" italics="true" pageId="18" pageNumber="53">Schuetzia</emphasis>
cf.
<emphasis id="B97CEAB8B02BFFB2779DFAFD56A0FA81" box="[128,239,1322,1346]" italics="true" pageId="18" pageNumber="53">bennieana</emphasis>
</taxonomicName>
but
<bibRefCitation id="EF994B5BB02BFFB27601FAFD57FAFA81" author="Walton, J." box="[284,437,1322,1346]" pageId="18" pageNumber="53" pagination="347 - 379" refId="ref25126" refString="Walton, J. (1931): Contributions to the knowledge of Lower Carboniferous plants. - Part III. - On the fossil-flora of the black limestones in Teila Quarry, Gwaenysgor, near Prestatyn, Flintshire, with special references to Diplopteridium teilianum Kidston sp. (gen, nov.) and some other fern-like fronds. - Philosophical Transactions of the Royal Society of London, B, 219: 347 - 379. https: // doi. org / 10.1098 / rstb. 1931.0007" type="journal article" year="1931">Walton (1931)</bibRefCitation>
showed this generic attribution to be incorrect; they are almost certainly the pollen organs of
<taxonomicName id="4C084D29B02BFFB2779DFABD5741FA41" box="[128,270,1386,1410]" class="Polypodiopsida" family="Cyatheaceae" genus="Calymmotheca" higherTaxonomySource="GBIF" kingdom="Plantae" order="Cyatheales" pageId="18" pageNumber="53" phylum="Tracheophyta" rank="species" species="divaricata">
<emphasis id="B97CEAB8B02BFFB2779DFABD5741FA41" box="[128,270,1386,1410]" italics="true" pageId="18" pageNumber="53">C. divaricata</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BB736AAB02BFFB277BBFA4E54D2F9D2" blockId="18.[128,768,1433,1553]" pageId="18" pageNumber="53">O c c u r r e n c e s. Widespread through the Limestone Coal Formation from the coal-bearing interval above the Black Metals Band, and through the Upper Limestone Formation to just below the Castlecary Limestone.</paragraph>
</subSubSection>
</treatment>
</document>