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<document id="F877AB44A5F6BF2B4C6537D246D3E0BB" ID-CLB-Dataset="4042" ID-DOI="10.5852/ejt.2021.772.1505" ID-GBIF-Dataset="b65bb5a9-bbe7-49a4-af44-4b4c03121288" ID-ISSN="2118-9773" ID-Zenodo-Dep="5536410" ID-ZooBank="9B82A32F-0A07-47E3-8684-FED7C8EBF1E9" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1632919496065" checkinUser="felipe" docAuthor="Miranda, Gustavo Silva de, Giupponi, Alessandro P. L., Prendini, Lorenzo &amp; Scharff, Nikolaj" docDate="2021" docId="8F431375FF78FF31A511FDA8FACEDB2A" docLanguage="en" docName="ejt-772_1-409.pdf" docOrigin="European Journal of Taxonomy 772" docSource="http://dx.doi.org/10.5852/ejt.2021.772.1505" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sarax pakistanus Miranda &amp; Giupponi &amp; Prendini &amp; Scharff 2021, comb. nov." docType="treatment" docVersion="14" lastPageNumber="227" masterDocId="737A6B0DFF99FFD3A710FFEFFFC6DC01" masterDocTitle="Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi)" masterLastPageNumber="409" masterPageNumber="1" pageNumber="226" updateTime="1699416367315" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="5CF340E9B07B7C885CE4D7D799D83FFA">Systematic revision of the pantropical whip spider family Charinidae Quintero, 1986 (Arachnida, Amblypygi)</mods:title>
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<mods:namePart id="1CE039F5100E1C3DC6A222A329024980">Miranda, Gustavo Silva de</mods:namePart>
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<mods:affiliation id="9D59AB56DC87A9F13D7E59F7ED5E84EC">Arachnology Lab, Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA.</mods:affiliation>
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<mods:affiliation id="16F7724103B1B43BFEAFA96E5958CBB6">Center for Macroecology, Evolution and Climate, Natural History Museum of Denmark, University of Copenhagen, Copenhagen, Denmark. &amp; Entomology Department, National Museum of Natural History, Smithsonian Institution, Washington DC, USA. &amp; Zoology Section, Natural History Museum of Denmark, University of Copenhagen, Denmark.</mods:affiliation>
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<heading id="5C1D150FFF78FF32A511FDA8FBF5DE60" box="[513,1075,582,610]" centered="true" fontSize="11" level="2" pageId="225" pageNumber="226" reason="2">
<taxonomicName id="C0EAD9E0FF78FF32A511FDA8FC6CDE63" ID-CoL="8VNGG" authority="(Weygoldt, 2005)" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt" baseAuthorityYear="2005" box="[513,938,582,610]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="pakistanus" status="comb. nov.">
<emphasis id="359E7E71FF78FF32A511FDA8FD13DE60" bold="true" box="[513,725,582,609]" italics="true" pageId="225" pageNumber="226">Sarax pakistanus</emphasis>
(
<bibRefCitation id="637BDF92FF78FF32A5F4FDA8FC64DE63" author="Weygoldt P." box="[740,930,583,610]" firstAuthor="Weygoldt" journalOrPublisher="Senckenbergiana Biologica" pageId="225" pageNumber="226" pagination="1 - 14" part="85" refId="ref156233" refString="Weygoldt P. 2005. Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae). Senckenbergiana Biologica 85: 1 - 14." title="Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae)" type="journal article" year="2005">Weygoldt, 2005</bibRefCitation>
)
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<taxonomicNameLabel id="2EADC30AFF78FF32A4A2FDA8FBF5DE60" box="[946,1075,583,609]" pageId="225" pageNumber="226" rank="species">comb. nov.</taxonomicNameLabel>
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<subSubSection id="4FF0F1E8FF78FF32A587FD85FC5BDE85" box="[663,925,618,645]" pageId="225" pageNumber="226" type="description">
<paragraph id="0755A263FF78FF32A587FD85FC5BDE85" blockId="225.[513,1075,582,645]" box="[663,925,618,645]" pageId="225" pageNumber="226">
<figureCitation id="9FD1BEE6FF78FF32A587FD85FD38DE84" box="[663,766,618,645]" captionStart="Fig" captionStartId="390.[189,232,1237,1263]" captionTargetBox="[192,1396,268,1195]" captionTargetId="figure-21@390.[189,1398,265,1198]" captionTargetPageId="390" captionText="Fig. 135. Map plotting known distributions of species of Sarax Simon, 1892 in Africa, the Middle East and South Asia." figureDoi="http://doi.org/10.5281/zenodo.5536776" httpUri="https://zenodo.org/record/5536776/files/figure.png" pageId="225" pageNumber="226">Figs 135</figureCitation>
,
<figureCitation id="9FD1BEE6FF78FF32A41BFD85FCFCDE85" box="[779,826,618,644]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="225" pageNumber="226">144</figureCitation>
;
<tableCitation id="4A6897D8FF78FF32A456FD85FC5BDE85" box="[838,925,618,644]" captionStart="Table 9" captionStartId="213.[181,251,576,605]" captionTargetId="graphics-0@213.[181,1934,690,1746]" captionTargetPageId="213" captionText="Table 9 (continued on next page). Measurements of all African, Middle Eastern and South Asian Sarax Simon, 1892." pageId="225" pageNumber="226">Table 9</tableCitation>
</paragraph>
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<treatmentCitation id="864B8472FF78FF32A7ADFD43FD74DEC7" author="Weygoldt P." box="[189,690,684,711]" page="8" pageId="225" pageNumber="226" year="2005">
<taxonomicName id="C0EAD9E0FF78FF32A7ADFD43FD74DEC7" ID-CoL="69PNY" authority="Weygoldt, 2005: 6 - 8" authorityName="Weygoldt" authorityPageNumber="6 - 8" authorityYear="2005" box="[189,690,684,711]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="pakistanus">
<emphasis id="359E7E71FF78FF32A7ADFD43FE74DEC7" box="[189,434,684,710]" italics="true" pageId="225" pageNumber="226">Charinus pakistanus</emphasis>
<bibRefCitation id="637BDF92FF78FF32A6A9FD43FD74DEC7" author="Weygoldt P." box="[441,690,684,711]" firstAuthor="Weygoldt" journalOrPublisher="Senckenbergiana Biologica" page="6 - 8" pageId="225" pageNumber="226" pagination="1 - 14" part="85" refId="ref156233" refString="Weygoldt P. 2005. Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae). Senckenbergiana Biologica 85: 1 - 14." title="Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae)" type="journal article" year="2005">Weygoldt, 2005: 68</bibRefCitation>
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, figs 1015.
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<emphasis id="359E7E71FF78FF32A7ADFD02FE74DF06" box="[189,434,749,775]" italics="true" pageId="225" pageNumber="226">Charinus pakistanus</emphasis>
</taxonomicName>
<treatmentCitation id="864B8472FF78FF32A6DFFD02FD3CDF09" author="Blick T. &amp; Seiter M." box="[463,762,749,776]" page="590" pageId="225" pageNumber="226" year="2016">
<bibRefCitation id="637BDF92FF78FF32A6DFFD02FD3CDF09" author="Blick T. &amp; Seiter M." box="[463,762,749,776]" firstAuthor="Blick" journalOrPublisher="Zootaxa" page="590" pageId="225" pageNumber="226" pagination="586 - 592" part="4161" refId="ref147053" refString="Blick T. &amp; Seiter M. 2016. Whip spiders (Amblypygi, Arachnida) of the Western Palaearctic - a review. Zootaxa 4161 (4): 586 - 592. https: // doi. org / 10.11646 / zootaxa. 4161.4.11" title="Whip spiders (Amblypygi, Arachnida) of the Western Palaearctic - a review" type="journal article" year="2016">Blick &amp; Seiter 2016: 590</bibRefCitation>
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.
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</paragraph>
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<emphasis id="359E7E71FF78FF32A7ADFCC0FEF0DF48" bold="true" box="[189,310,815,841]" pageId="225" pageNumber="226">Diagnosis</emphasis>
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This species may be separated from other species of
<taxonomicName id="C0EAD9E0FF78FF32A432FCB1FCA0DF79" authorityName="Simon" authorityYear="1892" box="[802,870,862,888]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="225" pageNumber="226" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FF78FF32A432FCB1FCA0DF79" box="[802,870,862,888]" italics="true" pageId="225" pageNumber="226">Sarax</emphasis>
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in Africa, the Middle East and South Asia by the following combination of characters: distal margin of female genital operculum with small median projection; ventral sac cover absent; female gonopod finger-like, with slender apex; sexual dimorphism in pedipalps apparently absent; pedipalp patella with large dorsal spine, not setiferous tubercle, distal to spine I, two-thirds length of spine I, two ventral spines, and one seta between spine I and distal margin (
<figureCitation id="9FD1BEE6FF78FF32A7D6FBE0FEA7D828" box="[198,353,1039,1065]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="225" pageNumber="226">Fig. 144EF</figureCitation>
); pedipalp tibia with base of two dorsal spines situated very close to each other; leg IV basitibia with three articles.
</paragraph>
<paragraph id="0755A263FF78FF32A7ADFB9AFB51D8B0" blockId="225.[188,1399,1140,1202]" pageId="225" pageNumber="226">
<taxonomicName id="C0EAD9E0FF78FF32A7ADFB9AFE43D88F" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt" baseAuthorityYear="2005" box="[189,389,1140,1167]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="pakistanus" status="comb. nov.">
<emphasis id="359E7E71FF78FF32A7ADFB9AFE43D88F" box="[189,389,1140,1167]" italics="true" pageId="225" pageNumber="226">Sarax pakistanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF78FF32A69CFB9BFDCBD88E" box="[396,525,1140,1167]" pageId="225" pageNumber="226" rank="species">comb. nov.</taxonomicNameLabel>
resembles
<taxonomicName id="C0EAD9E0FF78FF32A59EFB9AFCF0D88F" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" baseAuthorityYear="2021" box="[654,822,1140,1167]" class="Arachnida" family="Charinidae" genus="Charinides" kingdom="Animalia" order="Amblypygi" pageId="225" pageNumber="226" phylum="Arthropoda" rank="species" species="bengalensis" status="comb. nov.">
<emphasis id="359E7E71FF78FF32A59EFB9AFCF0D88F" box="[654,822,1140,1167]" italics="true" pageId="225" pageNumber="226">S. bengalensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF78FF32A42CFB9BFC7BD88E" box="[828,957,1140,1167]" pageId="225" pageNumber="226" rank="species">comb. nov.</taxonomicNameLabel>
but bears more teeth on the cheliceral claw, more spines on the pedipalp, and the eyes are not as developed, than the latter.
</paragraph>
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<emphasis id="359E7E71FF78FF32A7ADFB36FE82D8F2" bold="true" box="[189,324,1241,1267]" pageId="225" pageNumber="226">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF78FF32A7ADFAE7FAA7D922" blockId="225.[189,1377,1288,1315]" box="[189,1377,1288,1315]" pageId="225" pageNumber="226">
An adjective referring to
<collectingCountry id="7FFDE2F3FF78FF32A6F5FAE7FD8DD923" box="[485,587,1288,1314]" name="Pakistan" pageId="225" pageNumber="226">Pakistan</collectingCountry>
, the country in which the
<typeStatus id="D8511CC1FF78FF32A46CFAE6FC69D922" box="[892,943,1289,1315]" pageId="225" pageNumber="226">type</typeStatus>
locality is situated (
<bibRefCitation id="637BDF92FF78FF32A38AFAE7FA93D922" author="Weygoldt P." box="[1178,1365,1288,1315]" firstAuthor="Weygoldt" journalOrPublisher="Senckenbergiana Biologica" pageId="225" pageNumber="226" pagination="1 - 14" part="85" refId="ref156233" refString="Weygoldt P. 2005. Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae). Senckenbergiana Biologica 85: 1 - 14." title="Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae)" type="journal article" year="2005">Weygoldt 2005</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF78FF32A7ADFAA5FE6FDAA9" pageId="225" pageNumber="226" type="materials_examined">
<paragraph id="0755A263FF78FF32A7ADFAA5FEABD962" blockId="225.[189,365,1353,1380]" box="[189,365,1353,1380]" pageId="225" pageNumber="226">
<heading id="5C1D150FFF78FF32A7ADFAA5FEABD962" bold="true" box="[189,365,1353,1380]" fontSize="11" level="3" pageId="225" pageNumber="226" reason="3">
<emphasis id="359E7E71FF78FF32A7ADFAA5FEABD962" bold="true" box="[189,365,1353,1380]" pageId="225" pageNumber="226">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF78FF32A7FAFA96FD03D9FC" blockId="225.[189,1398,1401,1533]" pageId="225" pageNumber="226">
<materialsCitation id="B782A83EFF78FF32A7FAFA96FD03D9FC" ID-GBIF-Occurrence="3383565393" collectingDate="1983-06-03" collectionCode="MHNG" collectorName="C. Besuchet &amp; I. Lobl" country="Pakistan" elevation="1500" latitude="34.6875" location="Malkandi" longLatPrecision="15" longitude="73.44965" pageId="225" pageNumber="226" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="359E7E71FF78FF32A7FAFA96FE9AD992" bold="true" box="[234,348,1401,1427]" pageId="225" pageNumber="226">
<typeStatus id="D8511CC1FF78FF32A7FAFA96FE9AD992" box="[234,348,1401,1427]" pageId="225" pageNumber="226" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="7FFDE2F3FF78FF32A7ADFA73FE8FD9B7" box="[189,329,1436,1462]" name="Pakistan" pageId="225" pageNumber="226">PAKISTAN</collectingCountry>
<specimenCount id="11EC69EAFF78FF32A666FA73FE49D9B7" box="[374,399,1436,1462]" pageId="225" pageNumber="226" type="male"></specimenCount>
;
<location id="0235F4B8FF78FF32A6B5FA73FDDED9B7" LSID="urn:lsid:plazi:treatment:8F431375FF78FF31A511FDA8FACEDB2A:0235F4B8FF78FF32A6B5FA73FDDED9B7" box="[421,536,1436,1462]" country="Pakistan" latitude="34.6875" longLatPrecision="15" longitude="73.44965" name="Malkandi" pageId="225" pageNumber="226">Malkandi</location>
,
<locationDeviation id="16E7BB41FF78FF32A53EFA73FAB4D9B7" box="[558,1394,1436,1462]" firstLocation="Kawai" pageId="225" pageNumber="226" secondLocation="Mahandri, Kaghan Valley, Mansehra District">between Kawai and Mahandri, Kaghan Valley, Mansehra District</locationDeviation>
; [
<geoCoordinate id="62DEC4A4FF78FF32A7D4FA51FE9BD9DB" box="[196,349,1470,1498]" degrees="34" direction="north" minutes="41" orientation="latitude" pageId="225" pageNumber="226" precision="15" seconds="15" value="34.6875">34°4115″ N</geoCoordinate>
,
<geoCoordinate id="62DEC4A4FF78FF32A67AFA51FDE5D9DB" box="[362,547,1470,1498]" degrees="73" direction="east" minutes="26" orientation="longitude" pageId="225" pageNumber="226" precision="1" seconds="58.75" value="73.44965">73°2658.75″ E</geoCoordinate>
];
<elevation id="8CC74550FF78FF32A529FA50FD12D9D8" box="[569,724,1471,1498]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="225" pageNumber="226" unit="m" value="1500.0">
<quantity id="C0120F86FF78FF32A529FA50FD55D9DB" box="[569,659,1471,1498]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="225" pageNumber="226" unit="m" value="1500.0">1500 m</quantity>
a.s.l.
</elevation>
;
<date id="735484A3FF78FF32A5F2FA50FCA9D9D8" box="[738,879,1471,1498]" pageId="225" pageNumber="226" value="1983-06-03">
<collectingDate id="63107D4BFF78FF32A5F2FA50FCA9D9D8" box="[738,879,1471,1498]" pageId="225" pageNumber="226" value="1983-06-03">3 Jun. 1983</collectingDate>
</date>
;
<collectorName id="AA1FC7B5FF78FF32A46CFA50FBCDD9D8" box="[892,1035,1471,1497]" pageId="225" pageNumber="226">C. Besuchet</collectorName>
and
<collectorName id="AA1FC7B5FF78FF32A356FA2FFB50D9D8" box="[1094,1174,1471,1498]" pageId="225" pageNumber="226">I. Löbl</collectorName>
leg.; sifting leaves and old stumps in hardwood forest;
<collectionCode id="61FB3AA6FF78FF32A573FA0CFD07D9FC" box="[611,705,1507,1533]" pageId="225" pageNumber="226">MHNG</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="0755A263FF78FF32A7FAF9CBFE6FDAA9" blockId="225.[189,1399,1572,1705]" pageId="225" pageNumber="226">
<materialsCitation id="B782A83EFF78FF32A7FAF9CBFE63DAA9" ID-GBIF-Occurrence="3383565408" accessionNumber="SMF 40168" collectedFrom="forest, under stones" collectingDate="2003-07-10" collectionCode="SMF" collectorName="S. V. Ovchinnikov" country="Pakistan" elevation="800" latitude="33.738335" location="Khyber Pakhtunkhwa" longLatPrecision="774" longitude="73.05" pageId="225" pageNumber="226" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Khyber Pakhtunkhwa" typeStatus="paratype">
<emphasis id="359E7E71FF78FF32A7FAF9CBFEA1DA3F" bold="true" box="[234,359,1572,1598]" pageId="225" pageNumber="226">
<typeStatus id="D8511CC1FF78FF32A7FAF9CBFEA1DA3F" box="[234,359,1572,1598]" pageId="225" pageNumber="226" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="7FFDE2F3FF78FF32A7ADF9A7FE8FDA63" box="[189,329,1608,1634]" name="Pakistan" pageId="225" pageNumber="226">PAKISTAN</collectingCountry>
<specimenCount id="11EC69EAFF78FF32A673F9A7FE60DA63" box="[355,422,1608,1634]" pageId="225" pageNumber="226" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="11EC69EAFF78FF32A6A4F9A7FE3FDA63" box="[436,505,1608,1634]" pageId="225" pageNumber="226" type="male">2 ♂♂</specimenCount>
;
<collectingRegion id="C52E6C81FF78FF32A516F9A7FCC0DA63" box="[518,774,1608,1634]" country="Pakistan" name="Khyber Pakhtunkhwa" pageId="225" pageNumber="226">Khyber Pakhtunkhwa</collectingRegion>
[former North-West Frontier Province],
<locationDeviation id="16E7BB41FF78FF32A3F6F9A7FE9CDA84" pageId="225" pageNumber="226">
ca
<quantity id="C0120F86FF78FF32A218F9A7FA9FDA63" box="[1288,1369,1608,1635]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="225" pageNumber="226" unit="km" value="1.5">1.5 km</quantity>
N of Islamabad
</locationDeviation>
;
<geoCoordinate id="62DEC4A4FF78FF32A676F985FE2FDA87" box="[358,489,1642,1670]" degrees="33" direction="north" minutes="44.3" orientation="latitude" pageId="225" pageNumber="226" precision="92" value="33.738335">33°44.3 N</geoCoordinate>
,
<geoCoordinate id="62DEC4A4FF78FF32A6E6F985FDA6DA87" box="[502,608,1642,1670]" degrees="73" direction="east" minutes="03" orientation="longitude" pageId="225" pageNumber="226" precision="925" value="73.05">73°03 E</geoCoordinate>
; ca
<elevation id="8CC74550FF78FF32A580F984FCDDDA84" box="[656,795,1643,1670]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="225" pageNumber="226" unit="m" value="800.0">
<quantity id="C0120F86FF78FF32A580F984FD1DDA87" box="[656,731,1643,1670]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="225" pageNumber="226" unit="m" value="800.0">800 m</quantity>
a.s.l.
</elevation>
;
<date id="735484A3FF78FF32A43AF984FC79DA84" box="[810,959,1643,1669]" pageId="225" pageNumber="226" value="2003-07-10">
<collectingDate id="63107D4BFF78FF32A43AF984FC79DA84" box="[810,959,1643,1669]" pageId="225" pageNumber="226" value="2003-07-10">10 Jul. 2003</collectingDate>
</date>
;
<collectorName id="AA1FC7B5FF78FF32A4DCF984FB5BDA84" box="[972,1181,1643,1669]" pageId="225" pageNumber="226">S.V. Ovchinnikov</collectorName>
leg.;
<collectedFrom id="B4F6E19EFF78FF32A3F1F984FECCDAA8" pageId="225" pageNumber="226">forest, under stones</collectedFrom>
; SMF 40168
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF78FF31A7ADF93FFE8BD8EE" lastPageId="226" lastPageNumber="227" pageId="225" pageNumber="226" type="description">
<paragraph id="0755A263FF78FF32A7ADF93FFDD7DAEB" blockId="225.[189,529,1744,1770]" box="[189,529,1744,1770]" pageId="225" pageNumber="226">
<heading id="5C1D150FFF78FF32A7ADF93FFDD7DAEB" bold="true" box="[189,529,1744,1770]" fontSize="11" level="3" pageId="225" pageNumber="226" reason="3">
<emphasis id="359E7E71FF78FF32A7ADF93FFDD7DAEB" bold="true" box="[189,529,1744,1770]" pageId="225" pageNumber="226">Supplementary description</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF78FF32A7ADF910FB2EDB61" blockId="225.[189,1399,1791,1888]" pageId="225" pageNumber="226">
CARAPACE. Median eyes and median ocular tubercle reduced (
<figureCitation id="9FD1BEE6FF78FF32A484F910FBD6DB18" box="[916,1040,1791,1817]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="225" pageNumber="226">Fig. 144A</figureCitation>
); frontal process with rounded apex (
<figureCitation id="9FD1BEE6FF78FF32A618F8CDFE43DB3C" box="[264,389,1826,1853]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="225" pageNumber="226">Fig. 144C</figureCitation>
); one seta, not pair, posterior to median eyes; lateral eyes well developed, one seta posterior to lateral ocular triad; lateral ocular triad well separated from margin of carapace.
</paragraph>
<paragraph id="0755A263FF78FF32A7ADF867FDA3DBE9" blockId="225.[189,1399,1927,2025]" pageId="225" pageNumber="226">
STERNUM. Tritosternum projected anteriorly with typical setation (
<figureCitation id="9FD1BEE6FF78FF32A4CEF868FB9ADBA3" box="[990,1116,1927,1954]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="225" pageNumber="226">Fig. 144B</figureCitation>
); other sternal platelets small, rounded and convex, with pair of setae anteriorly; pentasternum with two setae anteriorly and without seta in membranous region.
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFEE6FC92DD22" blockId="226.[189,852,265,292]" box="[189,852,265,292]" pageId="226" pageNumber="227">OPISTHOSOMA. Ventral sacs and ventral sac cover absent.</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFEA4FEF7DDCF" blockId="226.[189,1399,330,462]" pageId="226" pageNumber="227">GENITALIA. Posterior margin of female genital operculum with small median projection; gonopod fingerlike, with slender apex, unsclerotized basally. Male gonopod sclerotized along entire margin of fistula and ventrally, near margin of genital operculum; lateral lobe 1 sclerotized apically; lateral lobe 2 not fimbriate.</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFE19FDD3DE7B" blockId="226.[189,1401,501,634]" pageId="226" pageNumber="227">CHELICERAE. Small, flat tooth projecting from retrolateral surface of basal segment, opposite to bifid tooth; retrolateral surface of claw with two patches of setae, basally and subdorsally; claw with eight teeth; row of eight setae on prolateral surface of basal segment; bifid tooth on basal segment with dorsal cusp larger than ventral cusp.</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFD4EFC45DFB0" blockId="226.[188,1401,672,945]" pageId="226" pageNumber="227">
PEDIPALPS. Coxae with seta encircled by round carina and four setae on margin. Femur with four dorsal spines and four ventral spines (
<figureCitation id="9FD1BEE6FF7BFF31A526FD2BFD17DEDF" box="[566,721,708,734]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="226" pageNumber="227">Fig. 144EF</figureCitation>
); two setiferous tubercles between spine 1 and proximal margin. Patella with three dorsal spines and two prominent setiferous tubercles proximal to spine 3; large spine distal to spine I, two-thirds length of spine I; three ventral spines; prominent setiferous tubercle between spine I and distal margin. Tibia with base of two dorsal spines situated very close to each other; three ventral spines distally and seta between spine and distal margin. Tarsus with two dorsal spines, distal spine long, more than half length of tarsus, proximal spine two-thirds length of distal spine (
<figureCitation id="9FD1BEE6FF7BFF31A7D5FC78FE87DFB0" box="[197,321,919,945]" captionStart="Fig" captionStartId="399.[189,232,1892,1918]" captionTargetBox="[222,1366,265,1859]" captionTargetId="figure-60@399.[222,1366,265,1859]" captionTargetPageId="399" captionText="Fig. 144. Sarax pakistanus (Weygoldt, 2005) comb. nov. (MHNG), general morphology,♂. A. Carapace, dorsal view. B. Sternum, ventral view. C. Frontal process. D. Pedipalp tarsus, frontal view. E. Pedipalp, dorsal view. F. Pedipalp, ventral view. Scale bars: AB, EF = 1 mm; C = 0.1 mm; D = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5536806" httpUri="https://zenodo.org/record/5536806/files/figure.png" pageId="226" pageNumber="227">Fig. 144D</figureCitation>
); cleaning organ with 3034 setae in ventral row.
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFC37FD3CD87E" blockId="226.[189,1399,984,1151]" pageId="226" pageNumber="227">
LEGS. Tibia of leg I with 21 articles; tarsus I with 35 articles; first tarsal article slightly longer than second (mean length of first article:
<quantity id="C0120F86FF7BFF31A56CFC13FD2AD817" box="[636,748,1020,1046]" metricMagnitude="-4" metricUnit="m" metricValue="3.2" pageId="226" pageNumber="227" unit="mm" value="0.32">0.32 mm</quantity>
; mean length of second article:
<quantity id="C0120F86FF7BFF31A368FC13FB2ED817" box="[1144,1256,1020,1046]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" pageId="226" pageNumber="227" unit="mm" value="0.22">0.22 mm</quantity>
, n =3). Leg IV basitibia with three pseudo-articles, trichobothrium
<emphasis id="359E7E71FF7BFF31A440FBF0FCA1D838" box="[848,871,1055,1081]" italics="true" pageId="226" pageNumber="227">bt</emphasis>
situated in proximal third of pseudo-article; distitibia trichobothrium
<emphasis id="359E7E71FF7BFF31A6F9FBADFDC3D85D" box="[489,517,1090,1116]" italics="true" pageId="226" pageNumber="227">bc</emphasis>
situated closer to
<emphasis id="359E7E71FF7BFF31A5F8FBADFD39D85D" box="[744,767,1090,1116]" italics="true" pageId="226" pageNumber="227">bf</emphasis>
than to s
<emphasis id="359E7E71FF7BFF31A464FBADFC4DD85D" box="[884,907,1090,1116]" italics="true" pageId="226" pageNumber="227">bf</emphasis>
,
<emphasis id="359E7E71FF7BFF31A48DFBACFC73D85D" box="[925,949,1091,1116]" italics="true" pageId="226" pageNumber="227">sc</emphasis>
and
<emphasis id="359E7E71FF7BFF31A4E5FBADFBCFD85D" box="[1013,1033,1090,1116]" italics="true" pageId="226" pageNumber="227">sf</emphasis>
series with five trichobothria; proximal-most trichobothria of
<emphasis id="359E7E71FF7BFF31A520FB89FD8ED87E" box="[560,584,1126,1151]" italics="true" pageId="226" pageNumber="227">sc</emphasis>
and
<emphasis id="359E7E71FF7BFF31A591FB8AFD53D87E" box="[641,661,1125,1151]" italics="true" pageId="226" pageNumber="227">sf</emphasis>
aligned.
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFB48FEB3D8C0" blockId="226.[189,373,1191,1217]" box="[189,373,1191,1217]" pageId="226" pageNumber="227">
<heading id="5C1D150FFF7BFF31A7ADFB48FEB3D8C0" bold="true" box="[189,373,1191,1217]" fontSize="11" level="3" pageId="226" pageNumber="227" reason="3">
<emphasis id="359E7E71FF7BFF31A7ADFB48FEB3D8C0" bold="true" box="[189,373,1191,1217]" pageId="226" pageNumber="227">Measurements</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFB3AFE8BD8EE" blockId="226.[189,333,1237,1263]" box="[189,333,1237,1263]" pageId="226" pageNumber="227">
See
<tableCitation id="4A6897D8FF7BFF31A7FEFB3AFE81D8EE" box="[238,327,1237,1263]" captionStart="Table 9" captionStartId="213.[181,251,576,605]" captionTargetId="graphics-0@213.[181,1934,690,1746]" captionTargetPageId="213" captionText="Table 9 (continued on next page). Measurements of all African, Middle Eastern and South Asian Sarax Simon, 1892." pageId="226" pageNumber="227">Table 9</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF7BFF31A7ADFAF9FDABD95E" box="[189,621,1302,1376]" pageId="226" pageNumber="227" type="distribution">
<paragraph id="0755A263FF7BFF31A7ADFAF9FE90D931" blockId="226.[189,342,1302,1328]" box="[189,342,1302,1328]" pageId="226" pageNumber="227">
<heading id="5C1D150FFF7BFF31A7ADFAF9FE90D931" bold="true" box="[189,342,1302,1328]" fontSize="11" level="3" pageId="226" pageNumber="227" reason="3">
<emphasis id="359E7E71FF7BFF31A7ADFAF9FE90D931" bold="true" box="[189,342,1302,1328]" pageId="226" pageNumber="227">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFAA9FDABD95E" blockId="226.[189,621,1349,1376]" box="[189,621,1349,1376]" pageId="226" pageNumber="227">
Known only from the
<typeStatus id="D8511CC1FF7BFF31A6D2FAA9FE33D961" box="[450,501,1350,1376]" pageId="226" pageNumber="227">type</typeStatus>
localities.
</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF7BFF31A7ADFA68FAA7D9CE" pageId="226" pageNumber="227" type="biology_ecology">
<paragraph id="0755A263FF7BFF31A7ADFA68FEB8D9A1" blockId="226.[189,382,1414,1441]" box="[189,382,1414,1441]" pageId="226" pageNumber="227">
<heading id="5C1D150FFF7BFF31A7ADFA68FEB8D9A1" bold="true" box="[189,382,1414,1441]" fontSize="11" level="3" pageId="226" pageNumber="227" reason="3">
<emphasis id="359E7E71FF7BFF31A7ADFA68FEB8D9A1" bold="true" box="[189,382,1414,1441]" pageId="226" pageNumber="227">Natural history</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADFA5AFAA7D9CE" blockId="226.[189,1377,1461,1488]" box="[189,1377,1461,1488]" pageId="226" pageNumber="227">The known specimens were collected under stones and tree stumps, and by sifting leaf litter in forest.</paragraph>
</subSubSection>
<subSubSection id="4FF0F1E8FF7BFF31A7ADFA18FACEDB2A" pageId="226" pageNumber="227" type="discussion">
<paragraph id="0755A263FF7BFF31A7ADFA18FEE9DA10" blockId="226.[189,303,1527,1553]" box="[189,303,1527,1553]" pageId="226" pageNumber="227">
<heading id="5C1D150FFF7BFF31A7ADFA18FEE9DA10" bold="true" box="[189,303,1527,1553]" fontSize="11" level="3" pageId="226" pageNumber="227" reason="3">
<emphasis id="359E7E71FF7BFF31A7ADFA18FEE9DA10" bold="true" box="[189,303,1527,1553]" pageId="226" pageNumber="227">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADF9CAFAF8DA63" blockId="226.[189,1398,1573,1635]" pageId="226" pageNumber="227">
The specimen labeled “
<typeStatus id="D8511CC1FF7BFF31A6DFF9CAFDFDDA3E" box="[463,571,1573,1599]" pageId="226" pageNumber="227" type="holotype">holotype</typeStatus>
” in the vial is male not female, as indicated in the original description by
<bibRefCitation id="637BDF92FF7BFF31A7F2F9A7FE6ADA62" author="Weygoldt P." box="[226,428,1608,1635]" firstAuthor="Weygoldt" journalOrPublisher="Senckenbergiana Biologica" pageId="226" pageNumber="227" pagination="1 - 14" part="85" refId="ref156233" refString="Weygoldt P. 2005. Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae). Senckenbergiana Biologica 85: 1 - 14." title="Biogeography, systematic position, and reproduction of Charinus ioanniticus (Kritscher, 1959) with the description of a new species from Pakistan (Chelicerata, Amblypygi, Charinidae)" type="journal article" year="2005">Weygoldt (2005)</bibRefCitation>
, and the
<typeStatus id="D8511CC1FF7BFF31A504F9A6FD42DA62" box="[532,644,1609,1635]" pageId="226" pageNumber="227" type="paratype">paratypes</typeStatus>
are
<specimenCount id="11EC69EAFF7BFF31A5A7F9A7FD3CDA62" box="[695,762,1608,1635]" pageId="226" pageNumber="227" type="female">3 ♀♀</specimenCount>
,
<specimenCount id="11EC69EAFF7BFF31A417F9A7FC8DDA62" box="[775,843,1608,1635]" pageId="226" pageNumber="227" type="male">2 ♂♂</specimenCount>
, again contrary to the original description.
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADF965FCD2DAC6" blockId="226.[188,1398,1674,1735]" pageId="226" pageNumber="227">
This species was originally described in the genus
<taxonomicName id="C0EAD9E0FF7BFF31A41AF965FCB1DAA5" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" box="[778,887,1674,1700]" class="Arachnida" family="Charinidae" genus="Charinus" kingdom="Animalia" order="Amblypygi" pageId="226" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FF7BFF31A41AF965FCB1DAA5" box="[778,887,1674,1700]" italics="true" pageId="226" pageNumber="227">Charinus</emphasis>
</taxonomicName>
and is hereby transferred to
<taxonomicName id="C0EAD9E0FF7BFF31A3D4F965FACEDAA5" authorityName="Simon" authorityYear="1892" box="[1220,1288,1674,1700]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="226" pageNumber="227" phylum="Arthropoda" rank="genus">
<emphasis id="359E7E71FF7BFF31A3D4F965FACEDAA5" box="[1220,1288,1674,1700]" italics="true" pageId="226" pageNumber="227">Sarax</emphasis>
</taxonomicName>
based on the phylogenetic analyses of
<bibRefCitation id="637BDF92FF7BFF31A501F942FCD6DAC6" author="Miranda G. S. &amp; Giupponi A. P. &amp; Scharff N. &amp; Prendini L." box="[529,784,1709,1735]" etAl="et al." firstAuthor="Miranda" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="226" pageNumber="227" refId="ref151478" refString="Miranda G. S., Giupponi A. P., Scharff N. &amp; Prendini L. 2021. Phylogeny and biogeography of the pantropical whip spider family Charinidae (Arachnida: Amblypygi). Zoological Journal of the Linnean Society. https: // doi. org / 10.1093 / zoolinnean / zlaa 101" title="Phylogeny and biogeography of the pantropical whip spider family Charinidae (Arachnida: Amblypygi)" type="book" year="2021">
<collectingRegion id="C52E6C81FF7BFF31A501F942FDB2DAC6" box="[529,628,1709,1735]" country="Venezuela" name="Miranda" pageId="226" pageNumber="227">Miranda</collectingRegion>
<emphasis id="359E7E71FF7BFF31A56CF941FD71DAC6" box="[636,695,1709,1735]" italics="true" pageId="226" pageNumber="227">et al.</emphasis>
(2021)
</bibRefCitation>
.
</paragraph>
<paragraph id="0755A263FF7BFF31A7ADF901FACEDB2A" blockId="226.[189,1399,1774,1836]" pageId="226" pageNumber="227">
The male gonopod of
<taxonomicName id="C0EAD9E0FF7BFF31A6DEF900FDADDB09" authorityName="Miranda &amp; Giupponi &amp; Prendini &amp; Scharff" authorityYear="2021" baseAuthorityName="Weygoldt" baseAuthorityYear="2005" box="[462,619,1774,1801]" class="Arachnida" family="Charinidae" genus="Sarax" kingdom="Animalia" order="Amblypygi" pageId="226" pageNumber="227" phylum="Arthropoda" rank="species" species="pakistanus" status="comb. nov.">
<emphasis id="359E7E71FF7BFF31A6DEF900FDADDB09" box="[462,619,1774,1801]" italics="true" pageId="226" pageNumber="227">S. pakistanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="2EADC30AFF7BFF31A566F901FD3ADB08" box="[630,764,1774,1801]" pageId="226" pageNumber="227" rank="species">comb. nov.</taxonomicNameLabel>
is completely different from all other species in the family. Detailed SEM images are needed for comparison with the genitalia of other charinids.
</paragraph>
</subSubSection>
</treatment>
</document>