276 lines
33 KiB
XML
276 lines
33 KiB
XML
<document id="FEE7172529C1C900FF31D01F94EFC95B" ID-DOI="10.5281/zenodo.5329203" ID-ISSN="0253-116X" ID-Zenodo-Dep="5329203" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630162199424" checkinUser="carolina" docAuthor="D. W" docDate="2012" docId="0386227E1938FFBBFF21B08DFC9030CC" docLanguage="en" docName="LinzerbiolBeitr.2012.44.2.1195-1205.pdf" docOrigin="Linzer biologische Beiträge 44 (2)" docStyle="DocumentStyle:24E5EB772D355DE2EE23B1E8D07853E0.9:LinzerbiolBeitr.2006-.journal_article" docStyleId="24E5EB772D355DE2EE23B1E8D07853E0" docStyleName="LinzerbiolBeitr.2006-.journal_article" docStyleVersion="9" docTitle="Chlaenius (Haplochlaenius) nanlingensis DEUVE & TIAN 2005" docType="treatment" docVersion="4" lastPageNumber="1203" masterDocId="FFBF5A061930FFB2FF93B055FFB0316C" masterDocTitle="A new species of genus Chlaenius BONELLI 1810, subgenus Haplochlaenius LUTSHNIK 1933 from China (Coleoptera, Carabidae, Chlaeniini) and notes on two species previously described" masterLastPageNumber="1205" masterPageNumber="1195" pageNumber="1203" updateTime="1699169689368" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="59DFC9F7430E3C4AE66AF3A885190270">A new species of genus Chlaenius BONELLI 1810, subgenus Haplochlaenius LUTSHNIK 1933 from China (Coleoptera, Carabidae, Chlaeniini) and notes on two species previously described</mods:title>
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<mods:name id="E6D03B2628282A2B385774EA9C0EA59A" type="personal">
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<mods:namePart id="437F065A6BBB1BF2B7126096010A59A2">D. W</mods:namePart>
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<mods:typeOfResource id="BB3AF499847C7A9E39F01B7D9D3D781F">text</mods:typeOfResource>
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<mods:title id="A0CFFC0292595F3A255924B641E51E29">Linzer biologische Beiträge</mods:title>
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<mods:part id="26A5FFCAA934BCEF83C2978891D3878F">
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<mods:date id="D56D18442463A9C7A772ED845F224AAD">2012</mods:date>
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<mods:number id="1BE8F614906C2CD97EDC2569C7A7E1B8">2012-12-28</mods:number>
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<mods:detail id="742251DEE539564E50AA91889665D52E" type="volume">
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<mods:number id="ACB3C8CAEE00090E3C42EA84CA405B2B">44</mods:number>
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<mods:start id="F549415C1D80E1CC3B563DDD1C9AF65D">1195</mods:start>
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<mods:end id="13C640E083FF1F445352B74C2A5429C5">1205</mods:end>
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<mods:identifier id="06332E3DB04FD070CA403ABB450C927A" type="DOI">10.5281/zenodo.5329203</mods:identifier>
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<mods:identifier id="AA20F14AE5B9A54197559FEB8E16D255" type="ISSN">0253-116X</mods:identifier>
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<mods:identifier id="C8A819DBCCF4CE38C7127A2E2A4ACDC8" type="Zenodo-Dep">5329203</mods:identifier>
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<treatment id="0386227E1938FFBBFF21B08DFC9030CC" ID-DOI="http://doi.org/10.5281/zenodo.5397064" ID-Zenodo-Dep="5397064" LSID="urn:lsid:plazi:treatment:0386227E1938FFBBFF21B08DFC9030CC" httpUri="http://treatment.plazi.org/id/0386227E1938FFBBFF21B08DFC9030CC" lastPageId="9" lastPageNumber="1203" pageId="8" pageNumber="1203">
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<subSubSection id="C335C0E31938FFBAFF21B08DFCA830C0" pageId="8" pageNumber="1203" type="nomenclature">
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<paragraph id="8B9093681938FFBAFF21B08DFCF53182" blockId="8.[178,837,216,238]" box="[178,837,216,238]" pageId="8" pageNumber="1203">
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<heading id="D0D824041938FFBAFF21B08DFCF53182" box="[178,837,216,238]" fontSize="8" level="2" pageId="8" pageNumber="1203" reason="2">
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<taxonomicName id="4C2FE8EB1938FFBAFF21B08DFCF53182" ID-CoL="5XVJR" authority="DEUVE & TIAN 2005" authorityName="DEUVE & TIAN" authorityYear="2005" box="[178,837,216,238]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="nanlingensis" subGenus="Haplochlaenius">
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<emphasis id="B95B4F7A1938FFBAFF21B08DFCF53182" bold="true" box="[178,837,216,238]" pageId="8" pageNumber="1203">
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<emphasis id="B95B4F7A1938FFBAFF21B08DFEAD3182" bold="true" box="[178,285,216,238]" italics="true" pageId="8" pageNumber="1203">Chlaenius</emphasis>
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(
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<emphasis id="B95B4F7A1938FFBAFEBFB08DFE623182" bold="true" box="[300,466,216,238]" italics="true" pageId="8" pageNumber="1203">Haplochlaenius</emphasis>
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)
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<emphasis id="B95B4F7A1938FFBAFE73B08DFDD63182" bold="true" box="[480,614,216,238]" italics="true" pageId="8" pageNumber="1203">nanlingensis</emphasis>
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DEUVE & TIAN 2005
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B9093681938FFBAFF21B15DFCA830C0" blockId="8.[178,1087,264,428]" pageId="8" pageNumber="1203">
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M a t e r i a l e x a m i n e d:
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<collectingCountry id="F338D3F81938FFBAFE69B15DFD863070" box="[506,566,264,284]" name="China" pageId="8" pageNumber="1203">China</collectingCountry>
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:
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<collectingRegion id="49EB5D8A1938FFBAFDAEB15DFD213070" box="[573,657,264,284]" country="China" name="Guangxi" pageId="8" pageNumber="1203">Guangxi</collectingRegion>
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:1, 7: NE
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<collectingRegion id="49EB5D8A1938FFBAFCB5B15DFCC33070" box="[806,883,264,284]" country="China" name="Guangxi" pageId="8" pageNumber="1203">Guangxi</collectingRegion>
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, Xing`an Co., Mao’er Shan, Mao`er Mountain Nature Reserve,
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<quantity id="4CD73E8D1938FFBAFDD8B175FD363058" box="[587,646,288,308]" metricMagnitude="2" metricUnit="m" metricValue="8.23" pageId="8" pageNumber="1203" unit="m" value="823.0">823 m</quantity>
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, N 25.51.497’/E 110.29.803’,
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<date id="FF91B5A81938FFBAFC01B175FC663058" box="[914,982,288,308]" pageId="8" pageNumber="1203" value="2007-05">V 2007</date>
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, M. Häckel & R. Sehnal leg. (cHÄCK, cSEHN, cWR, NHW). 1, 4: Mao’ er Shan`,
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<quantity id="4CD73E8D1938FFBAFC15B16DFC0E3020" box="[902,958,312,332]" metricMagnitude="2" metricUnit="m" metricValue="4.93" pageId="8" pageNumber="1203" unit="m" value="493.0">493 m</quantity>
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/ N 25.51.11’/ E 110. 29.15’, R. Sehnal leg. (cSEHN, cWR). 2: Mao’er Shan,
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<quantity id="4CD73E8D1938FFBAFCA3B105FC2C3008" box="[816,924,336,356]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="8" pageNumber="1203" unit="m" value="950.0" valueMax="1100.0" valueMin="800.0">800-1100 m</quantity>
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,
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<geoCoordinate id="EE1BF5AF1938FFBAFC35B105FC5A3008" box="[934,1002,336,356]" degrees="25" direction="north" minutes="52" orientation="latitude" pageId="8" pageNumber="1203" precision="925" value="25.866667">25°52N</geoCoordinate>
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/
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<geoCoordinate id="EE1BF5AF1938FFBAFC62B105FB8D3008" box="[1009,1085,336,356]" degrees="110" direction="east" minutes="29" orientation="longitude" pageId="8" pageNumber="1203" precision="925" value="110.48333">110°29E</geoCoordinate>
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(dense primary forest with strong bamboo, steep slopes),
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<date id="FF91B5A81938FFBAFC97B13DFC263010" box="[772,918,360,380]" pageId="8" pageNumber="1203" value="2009-05-28" valueMax="2009-06-09" valueMin="2009-05-28">28.V.-9.VI.2009</date>
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, Chr. Reuter leg. (cREUT, cWR). -
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<collectingRegion id="49EB5D8A1938FFBAFE07B1D5FE5330F8" box="[404,483,384,404]" country="China" name="Guizhou" pageId="8" pageNumber="1203">Guizhou</collectingRegion>
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: 2: SE.
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<collectingRegion id="49EB5D8A1938FFBAFDDCB1D5FD2E30F8" box="[591,670,384,404]" country="China" name="Guizhou" pageId="8" pageNumber="1203">Guizhou</collectingRegion>
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, Dushan, Gengding Shan env.,
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<quantity id="4CD73E8D1938FFBAFC41B1D5FBAD30F8" box="[978,1053,384,404]" metricMagnitude="3" metricUnit="m" metricValue="1.4449999999999998" pageId="8" pageNumber="1203" unit="m" value="1445.0">1445 m</quantity>
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,
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<geoCoordinate id="EE1BF5AF1938FFBAFBBFB1D5FE9930C0" degrees="25" direction="north" minutes="52.5" orientation="latitude" pageId="8" pageNumber="1203" precision="92" value="25.875">N 25°52.5‘</geoCoordinate>
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/
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<geoCoordinate id="EE1BF5AF1938FFBAFEA3B1CDFE3C30C0" box="[304,396,408,428]" degrees="107" direction="east" minutes="38" orientation="longitude" pageId="8" pageNumber="1203" precision="925" value="107.63333">E 107°38’</geoCoordinate>
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, M. Häckel & R. Sehnal leg. (cSEHN, cWR.)
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</paragraph>
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</subSubSection>
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<subSubSection id="C335C0E31938FFBBFF21B1EEFC9030CC" lastPageId="9" lastPageNumber="1204" pageId="8" pageNumber="1203" type="description">
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<paragraph id="8B9093681938FFBAFF21B1EEFE5A328C" blockId="8.[178,1085,443,992]" pageId="8" pageNumber="1203">
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The description based on
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<specimenCount id="9D2958E11938FFBAFE2DB1EEFDAE30BD" box="[446,542,443,465]" pageId="8" pageNumber="1203" type="male">one male</specimenCount>
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, coming from the slopes of the Mount Shikengkong situated in the Nanling Nature reserve (Dadongshan, Lianzhou) in the Chinese Province of
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<collectingRegion id="49EB5D8A1938FFBAFF42B1A0FEFA3367" box="[209,330,501,523]" country="China" name="Guangdong" pageId="8" pageNumber="1203">Guangdong</collectingRegion>
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, close to the border to
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<collectingRegion id="49EB5D8A1938FFBAFDD5B1A3FD3D3367" box="[582,653,502,523]" country="China" name="Hunan" pageId="8" pageNumber="1203">Hunan</collectingRegion>
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. The material from the Mao’er Shan fits very well the description, and, though not having investigated the type which is stored in the collection of the South
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<collectingCountry id="F338D3F81938FFBAFE63B265FD9C332A" box="[496,556,560,582]" name="China" pageId="8" pageNumber="1203">China</collectingCountry>
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Agricultural University, I refer the name
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<taxonomicName id="4C2FE8EB1938FFBAFB81B265FE82330F" authorityName="DEUVE & TIAN" authorityYear="2005" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="nanlingensis">
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<emphasis id="B95B4F7A1938FFBAFB81B265FB86332A" box="[1042,1078,560,582]" italics="true" pageId="8" pageNumber="1203">Chl</emphasis>
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.
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<emphasis id="B95B4F7A1938FFBAFF21B218FE82330F" box="[178,306,589,611]" italics="true" pageId="8" pageNumber="1203">nanlingensis</emphasis>
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</taxonomicName>
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to the material noted above. A habitus photograph of the
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<typeStatus id="54942DCA1938FFBAFC09B218FC43330F" box="[922,1011,589,611]" pageId="8" pageNumber="1203" type="holotype">holotype</typeStatus>
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kindly provided by M. Tian enabled the species to be recognised beyond doubt without having to examine the specimen itself. The species is brachypterous (this fact was not mentioned in the description),
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<specimenCount id="9D2958E11938FFBAFEE4B2F0FDB133D7" box="[375,513,677,699]" pageId="8" pageNumber="1203" type="generic">one specimen</specimenCount>
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investigated has the hindwings hardly as long as half of elytral length, but it is obviously able to have a wider range, the distance between the Mount Shikengkong and the Mao’er Shan is about
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<quantity id="4CD73E8D1938FFBAFD4DB28AFC833399" box="[734,819,735,757]" metricMagnitude="5" metricUnit="m" metricValue="2.7" pageId="8" pageNumber="1203" unit="km" value="270.0">270 km</quantity>
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, with a landscape surely forming no considerable geographical barriers which could stop an expansion of brachypterous species, also the findings in the Gengding Shan in the southeast of the Province Guizou, still more north-westwards, speaks for this assumption. Just recently I received some habitus photographs of specimens stored in the Institute of Zoology, Chinese Academy of Sciences,
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<collectingRegion id="49EB5D8A1938FFBAFE65B327FDF132E4" box="[502,577,882,904]" country="China" name="Beijing" pageId="8" pageNumber="1203">Beijing</collectingRegion>
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and identified as
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<taxonomicName id="4C2FE8EB1938FFBAFD69B327FC1B32E4" authorityName="DEUVE & TIAN" authorityYear="2005" box="[762,939,882,904]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="nanlingensis">
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<emphasis id="B95B4F7A1938FFBAFD69B327FC1B32E4" box="[762,939,882,904]" italics="true" pageId="8" pageNumber="1203">Chl. nanlingensis</emphasis>
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</taxonomicName>
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, coming from
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<collectingRegion id="49EB5D8A1938FFBAFF21B3C5FF4532C9" box="[178,245,912,933]" country="China" name="Hunan" pageId="8" pageNumber="1203">Hunan</collectingRegion>
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(Shaoshan-shihu) and from
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<collectingRegion id="49EB5D8A1938FFBAFDB7B3DAFDC532C9" box="[548,629,911,933]" country="China" name="Sichuan" pageId="8" pageNumber="1203">Sichuan</collectingRegion>
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(Ya’an), confirming that the species has a wider range than originally thought. New records for the Chinese provinces of
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<collectingRegion id="49EB5D8A1938FFBAFC4CB3F9FB8932AE" box="[991,1081,940,962]" country="China" name="Guangxi" pageId="8" pageNumber="1203">Guangxi</collectingRegion>
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,
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<collectingRegion id="49EB5D8A1938FFBAFF21B39FFEBC328C" box="[178,268,970,992]" country="China" name="Guizhou" pageId="8" pageNumber="1203">Guizhou</collectingRegion>
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,
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<collectingRegion id="49EB5D8A1938FFBAFE85B39FFEEC32B3" box="[278,348,970,991]" country="China" name="Hunan" pageId="8" pageNumber="1203">Hunan</collectingRegion>
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, and
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<collectingRegion id="49EB5D8A1938FFBAFE02B39FFE56328C" box="[401,486,970,992]" country="China" name="Sichuan" pageId="8" pageNumber="1203">Sichuan</collectingRegion>
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.
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</paragraph>
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<paragraph id="8B9093681938FFBBFF21B3A5FC9030CC" blockId="8.[178,1085,1007,1585]" lastBlockId="9.[178,1085,219,416]" lastPageId="9" lastPageNumber="1204" pageId="8" pageNumber="1203">
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N o t e s: The genus
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<taxonomicName id="4C2FE8EB1938FFBAFE34B3BAFD1B3569" authority="CASALE 1984" authorityName="CASALE" authorityYear="1984" box="[423,683,1007,1029]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="genus">
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<emphasis id="B95B4F7A1938FFBAFE34B3BAFDBC3569" box="[423,524,1007,1029]" italics="true" pageId="8" pageNumber="1203">Vachinius</emphasis>
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<bibRefCitation id="EFBEEE991938FFBAFD8AB3BAFD1B3569" author="CASALE A" box="[537,683,1007,1029]" pageId="8" pageNumber="1203" pagination="371 - 382" refId="ref5537" refString="CASALE A. (1984): The new Asiatic genus Vachinius (Carabidae: Callistinae) with three new species. - Bollettino del Museo Regionale di Scienze Naturali, Torino 2: 371 - 382." type="journal article" year="1984">CASALE 1984</bibRefCitation>
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</taxonomicName>
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was founded on several species:
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<taxonomicName id="4C2FE8EB1938FFBAFBB4B3A5FE6E354E" authority="(ANDREWES 1937)" baseAuthorityName="ANDREWES" baseAuthorityYear="1937" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="subglaber">
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<emphasis id="B95B4F7A1938FFBAFBB4B3A5FEA7354E" italics="true" pageId="8" pageNumber="1203">V. subglaber</emphasis>
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(ANDREWES 1937)
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</taxonomicName>
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,
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<typeStatus id="54942DCA1938FFBAFE79B458FDA6354E" box="[490,534,1037,1058]" pageId="8" pageNumber="1203">type</typeStatus>
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species from
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<collectingRegion id="49EB5D8A1938FFBAFD35B459FD42354E" box="[678,754,1036,1058]" country="India" name="Sikkim" pageId="8" pageNumber="1203">Sikkim</collectingRegion>
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,
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<taxonomicName id="4C2FE8EB1938FFBAFD6DB458FB89354E" authority="CASALE 1984" authorityName="CASALE" authorityYear="1984" box="[766,1081,1036,1058]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="pseudoglaber">
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<emphasis id="B95B4F7A1938FFBAFD6DB458FC14354E" box="[766,932,1036,1058]" italics="true" pageId="8" pageNumber="1203">V. pseudoglaber</emphasis>
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<bibRefCitation id="EFBEEE991938FFBAFC38B459FB89354E" author="CASALE A" box="[939,1081,1036,1058]" pageId="8" pageNumber="1203" pagination="371 - 382" refId="ref5537" refString="CASALE A. (1984): The new Asiatic genus Vachinius (Carabidae: Callistinae) with three new species. - Bollettino del Museo Regionale di Scienze Naturali, Torino 2: 371 - 382." type="journal article" year="1984">CASALE 1984</bibRefCitation>
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</taxonomicName>
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, from northern
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<collectingCountry id="F338D3F81938FFBAFEDCB47CFE363553" box="[335,390,1065,1087]" name="India" pageId="8" pageNumber="1203">India</collectingCountry>
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,
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<taxonomicName id="4C2FE8EB1938FFBAFE06B47FFD0C3553" authority="CASALE 1984" authorityName="CASALE" authorityYear="1984" box="[405,700,1065,1087]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="holzschuhi">
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<emphasis id="B95B4F7A1938FFBAFE06B47FFD923553" box="[405,546,1065,1087]" italics="true" pageId="8" pageNumber="1203">V. holzschuhi</emphasis>
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<bibRefCitation id="EFBEEE991938FFBAFDBEB47CFD0C3553" author="CASALE A" box="[557,700,1065,1087]" pageId="8" pageNumber="1203" pagination="371 - 382" refId="ref5537" refString="CASALE A. (1984): The new Asiatic genus Vachinius (Carabidae: Callistinae) with three new species. - Bollettino del Museo Regionale di Scienze Naturali, Torino 2: 371 - 382." type="journal article" year="1984">CASALE 1984</bibRefCitation>
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</taxonomicName>
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from eastern
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<collectingCountry id="F338D3F81938FFBAFCC5B47CFC253553" box="[854,917,1065,1087]" name="Nepal" pageId="8" pageNumber="1203">Nepal</collectingCountry>
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, and
|
||
<taxonomicName id="4C2FE8EB1938FFBAFC47B47FFE8D3531" authority="CASALE 1984" authorityName="CASALE" authorityYear="1984" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="pilosus">
|
||
<emphasis id="B95B4F7A1938FFBAFC47B47FFB8D3553" box="[980,1085,1065,1087]" italics="true" pageId="8" pageNumber="1203">V. pilosus</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFF21B412FE8D3531" author="CASALE A" box="[178,317,1095,1117]" pageId="8" pageNumber="1203" pagination="371 - 382" refId="ref5537" refString="CASALE A. (1984): The new Asiatic genus Vachinius (Carabidae: Callistinae) with three new species. - Bollettino del Museo Regionale di Scienze Naturali, Torino 2: 371 - 382." type="journal article" year="1984">CASALE 1984</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="F338D3F81938FFBAFEE9B412FE633531" box="[378,467,1095,1117]" name="Vietnam" pageId="8" pageNumber="1203">Vietnam</collectingCountry>
|
||
, underlining that these species are typical
|
||
<taxonomicName id="4C2FE8EB1938FFBAFC12B412FC5D3531" authorityName="BRULLE" authorityYear="1834" box="[897,1005,1095,1117]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
|
||
, near to species of the genus
|
||
<emphasis id="B95B4F7A1938FFBAFE18B431FD853516" box="[395,565,1124,1146]" italics="true" pageId="8" pageNumber="1203">Macrochlaenites</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFDADB430FD6B3516" author="KUNTZEN H" box="[574,731,1124,1146]" pageId="8" pageNumber="1203" pagination="91 - 156" refId="ref5843" refString="KUNTZEN H. (1919): Die Carabidenfauna Deutsch-Sudwestafrikas. - Mitteilungen aus dem Zoologischen Museum in Berlin 9 (2): 91 - 156." type="journal article" year="1919">KUNTZEN 1919</bibRefCitation>
|
||
but are distinguished by having a cordate pronotum, strongly sinuate toward base, a dimorphic apex and integument of elytra in males and females, a different construction of the male genitalia, and reduced wings. Later, further species were described, one from
|
||
<collectingCountry id="F338D3F81938FFBAFC84B4EEFCCA35BE" box="[791,890,1211,1234]" name="Thailand" pageId="8" pageNumber="1203">Thailand</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C2FE8EB1938FFBAFC19B4EEFEE93583" authority="MORVAN 1991" authorityName="MORVAN" authorityYear="1991" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="thailandensis">
|
||
<emphasis id="B95B4F7A1938FFBAFC19B4EEFB8D35BE" box="[906,1085,1211,1234]" italics="true" pageId="8" pageNumber="1203">V. thailandensis</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFF21B48DFEE93583" author="MORVAN P" box="[178,345,1240,1263]" pageId="8" pageNumber="1203" pagination="55 - 62" refId="ref5924" refString="MORVAN P. [D]. (1991): Contribution a la connaissance des coleopteres Carabidae de Thailand. - Elytron 5: 55 - 62." type="journal article" year="1991">MORVAN 1991</bibRefCitation>
|
||
</taxonomicName>
|
||
, three from
|
||
<collectingCountry id="F338D3F81938FFBAFE70B48CFD933583" box="[483,547,1241,1263]" name="China" pageId="8" pageNumber="1203">China</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C2FE8EB1938FFBAFDA1B48DFC073583" authority="MORVAN 1997" authorityName="MORVAN" authorityYear="1997" box="[562,951,1240,1263]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="deuvei">
|
||
<emphasis id="B95B4F7A1938FFBAFDA1B48DFD2A3583" box="[562,666,1240,1263]" italics="true" pageId="8" pageNumber="1203">V. deuvei</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFD35B48FFCF13583" author="MORVAN P" box="[678,833,1241,1263]" pageId="8" pageNumber="1203" pagination="1 - 23" refId="ref5953" refString="MORVAN P. [D]. (1997): Etude faunistique des coleopteres du Nepal avec extension aux provinces chinoises du Yunnan et du Sichuan. Genre Andrewesius JEDLICKA et Vachinius CASALE. - Loened Aziad Amprevaned Feuraskelleged C'Hwiledig 2: 1 - 23." type="journal article" year="1997">MORVAN 1997</bibRefCitation>
|
||
(
|
||
<collectingRegion id="49EB5D8A1938FFBAFCC0B48CFC1F3583" box="[851,943,1241,1263]" country="China" name="Guangxi" pageId="8" pageNumber="1203">Guangxi</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2FE8EB1938FFBAFC55B48FFE1C3460" authority="MORVAN 1997" authorityName="MORVAN" authorityYear="1997" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="hunanus">
|
||
<emphasis id="B95B4F7A1938FFBAFC55B48FFB8D3583" box="[966,1085,1241,1263]" italics="true" pageId="8" pageNumber="1203">V. hunanus</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFF21B4A2FEFC3460" author="MORVAN P" box="[178,332,1270,1292]" pageId="8" pageNumber="1203" pagination="1 - 23" refId="ref5953" refString="MORVAN P. [D]. (1997): Etude faunistique des coleopteres du Nepal avec extension aux provinces chinoises du Yunnan et du Sichuan. Genre Andrewesius JEDLICKA et Vachinius CASALE. - Loened Aziad Amprevaned Feuraskelleged C'Hwiledig 2: 1 - 23." type="journal article" year="1997">MORVAN 1997</bibRefCitation>
|
||
(
|
||
<collectingRegion id="49EB5D8A1938FFBAFECEB4A3FE153460" box="[349,421,1270,1292]" country="China" name="Hunan" pageId="8" pageNumber="1203">Hunan</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2FE8EB1938FFBAFE7AB4A2FC603460" authority="KIRSCHENHOFER 2003" authorityName="KIRSCHENHOFER" authorityYear="2003" box="[489,976,1270,1292]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="wrasei">
|
||
<emphasis id="B95B4F7A1938FFBAFE7AB4A2FDFC3460" box="[489,588,1271,1292]" italics="true" pageId="8" pageNumber="1203">V. wrasei</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFDC5B4A2FC8C3460" author="KIRSCHENHOFER E" box="[598,828,1270,1292]" pageId="8" pageNumber="1203" pagination="29 - 60" refId="ref5763" refString="KIRSCHENHOFER E. (2003): Uber neue und wenig bekannte Carabidae aus der athiopischen und orientalischen Region (Coleoptera: Carabidae, Chlaeniinae, Pterostichinae). - Entomofauna 24 (3): 29 - 60." type="journal article" year="2003">KIRSCHENHOFER 2003</bibRefCitation>
|
||
(
|
||
<collectingRegion id="49EB5D8A1938FFBAFCDEB4A3FC783460" box="[845,968,1270,1292]" country="China" name="Guangdong" pageId="8" pageNumber="1203">Guangdong</collectingRegion>
|
||
)
|
||
</taxonomicName>
|
||
, one from Sulawesi:
|
||
<taxonomicName id="4C2FE8EB1938FFBAFEB0B540FD343446" authority="KIRSCHENHOFER 1998" authorityName="KIRSCHENHOFER" authorityYear="1998" box="[291,644,1300,1322]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="baehri">
|
||
<emphasis id="B95B4F7A1938FFBAFEB0B540FE393446" box="[291,393,1300,1322]" italics="true" pageId="8" pageNumber="1203">V. baehri</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFE0BB541FD343446" author="KIRSCHENHOFER E" box="[408,644,1300,1322]" pageId="8" pageNumber="1203" pagination="317 - 332" refId="ref5732" refString="KIRSCHENHOFER E. (1998): Neue Chlaeniinae der palaearktischen und orientalischen Region (Coleoptera, Carabidae). - Entomofauna 19 (20): 317 - 332." type="journal article" year="1998">KIRSCHENHOFER 1998</bibRefCitation>
|
||
</taxonomicName>
|
||
, and one from
|
||
<collectingCountry id="F338D3F81938FFBAFCAEB541FC363445" box="[829,902,1300,1321]" name="Myanmar" pageId="8" pageNumber="1203">Burma</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C2FE8EB1938FFBAFC04B540FEE7342B" authority="LASSALLE 2001" authorityName="LASSALLE" authorityYear="2001" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="burmanensis">
|
||
<emphasis id="B95B4F7A1938FFBAFC04B540FB8D3446" box="[919,1085,1300,1322]" italics="true" pageId="8" pageNumber="1203">V. burmanensis</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFF21B567FEE7342B" author="LASSALLE B" box="[178,343,1329,1351]" pageId="8" pageNumber="1203" pagination="239 - 243" refId="ref5871" refString="LASSALLE B. (2001): Chasses en Birmanie (Coleoptera Carabidae). - L'Entomologiste 57 (6): 239 - 243." type="journal article" year="2001">LASSALLE 2001</bibRefCitation>
|
||
</taxonomicName>
|
||
. Investigating a larger number of
|
||
<taxonomicName id="4C2FE8EB1938FFBAFD27B564FCDA342B" box="[692,874,1329,1351]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B95B4F7A1938FFBAFD27B564FCAA342B" box="[692,794,1329,1351]" italics="true" pageId="8" pageNumber="1203">Chlaenius</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
belonging to subgenus
|
||
<taxonomicName id="4C2FE8EB1938FFBAFF4CB51BFD9E3408" authority="LUTSHNIK 1933" authorityName="LUTSHNIK" authorityYear="1933" box="[223,558,1358,1380]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="subGenus" subGenus="Haplochlaenius">
|
||
<emphasis id="B95B4F7A1938FFBAFF4CB51BFECF3408" box="[223,383,1358,1380]" italics="true" pageId="8" pageNumber="1203">Haplochlaenius</emphasis>
|
||
<bibRefCitation id="EFBEEE991938FFBAFE19B51AFD9E3408" author="LUTSHNIK V" box="[394,558,1358,1380]" pageId="8" pageNumber="1203" pagination="169 - 172" refId="ref5897" refString="LUTSHNIK V. N. (1933): Synopsis subgenerum palaearcticorum generis Chlaenius BON. - Casopis Ceskoslovenske Spolecnosti Entomologicke 30: 169 - 172." type="journal article" year="1933">LUTSHNIK 1933</bibRefCitation>
|
||
</taxonomicName>
|
||
(a part of them placed under
|
||
<emphasis id="B95B4F7A1938FFBAFCE6B51BFBAF3408" box="[885,1055,1358,1380]" italics="true" pageId="8" pageNumber="1203">Macrochlaenites</emphasis>
|
||
in older taxonomic classifications) I found partly the same characters peculiar to species of
|
||
<emphasis id="B95B4F7A1938FFBAFF21B5DCFEA734F3" box="[178,279,1417,1439]" italics="true" pageId="8" pageNumber="1203">Vachinius</emphasis>
|
||
, such as the construction of the mentum, the sinuate pronotum, the partly occurring carinate elytral intervals, a reduction of the hindwings and the morphology of the male genitalia. The classification of the new species described above posed problems and I decided to describe it in the genus
|
||
<taxonomicName id="4C2FE8EB1938FFBAFDE5B5B4FD6C349B" authorityName="BONELLI" authorityYear="1810" box="[630,732,1505,1527]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B95B4F7A1938FFBAFDE5B5B4FD6C349B" box="[630,732,1505,1527]" italics="true" pageId="8" pageNumber="1203">Chlaenius</emphasis>
|
||
</taxonomicName>
|
||
, subgenus
|
||
<taxonomicName id="4C2FE8EB1938FFBAFCC4B5B4FC47349B" authorityName="LUTSHNIK" authorityYear="1933" box="[855,1015,1505,1527]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="subGenus" subGenus="Haplochlaenius">
|
||
<emphasis id="B95B4F7A1938FFBAFCC4B5B4FC47349B" box="[855,1015,1505,1527]" italics="true" pageId="8" pageNumber="1203">Haplochlaenius</emphasis>
|
||
</taxonomicName>
|
||
. So it seems necessary to clarify the systematic position and the taxonomic rank of the genus
|
||
<emphasis id="B95B4F7A1938FFBAFF21B64EFEA7375D" box="[178,279,1563,1585]" italics="true" pageId="8" pageNumber="1203">Vachinius</emphasis>
|
||
within the tribe
|
||
<taxonomicName id="4C2FE8EB1938FFBAFE2CB64EFD98375D" authorityName="BRULLE" authorityYear="1834" box="[447,552,1563,1585]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="tribe" tribe="Chlaeniini">Chlaeniini</taxonomicName>
|
||
and especially its relationship to
|
||
<taxonomicName id="4C2FE8EB1938FFBAFCEEB64EFBAD375D" authorityName="LUTSHNIK" authorityYear="1933" box="[893,1053,1563,1585]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="1203" phylum="Arthropoda" rank="subGenus" subGenus="Haplochlaenius">
|
||
<emphasis id="B95B4F7A1938FFBAFCEEB64EFBAD375D" box="[893,1053,1563,1585]" italics="true" pageId="8" pageNumber="1203">Haplochlaenius</emphasis>
|
||
</taxonomicName>
|
||
by careful examination of all taxa (see also the remarks in DEUVE & TIAN 2005), including also genetic techniques. Just recently
|
||
<bibRefCitation id="EFBEEE991939FFBBFDC3B0ADFCFA3062" author="KIRSCHENHOFER E" box="[592,842,248,270]" pageId="9" pageNumber="1204" pagination="83 - 90" refId="ref5803" refString="KIRSCHENHOFER E. (2012): Neue Arten der Gattung Vachinius CASALE, 1984 von Laos (Coleoptera: Carabidae). - Mitteilungen des internationalen entomologischen Vereins 37 (1 / 2): 83 - 90." type="journal article" year="2012">KIRSCHENHOFER (2012)</bibRefCitation>
|
||
described two further
|
||
<emphasis id="B95B4F7A1939FFBBFF21B140FEA73047" box="[178,279,277,299]" italics="true" pageId="9" pageNumber="1204">Vachinius</emphasis>
|
||
species from
|
||
<collectingCountry id="F338D3F81939FFBBFE3FB143FE513047" box="[428,481,278,299]" name="Laos" pageId="9" pageNumber="1204">Laos</collectingCountry>
|
||
:
|
||
<taxonomicName id="4C2FE8EB1939FFBBFE7EB143FDFA3047" authorityName="Kirschenhofer" authorityYear="2012" box="[493,586,277,299]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1204" phylum="Arthropoda" rank="species" species="hajeki">
|
||
<emphasis id="B95B4F7A1939FFBBFE7EB143FDFA3047" box="[493,586,277,299]" italics="true" pageId="9" pageNumber="1204">V. hajeki</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C2FE8EB1939FFBBFD13B143FD4C3047" authorityName="Kirschenhofer" authorityYear="2012" box="[640,764,277,299]" class="Insecta" family="Carabidae" genus="Vachinius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1204" phylum="Arthropoda" rank="species" species="laosensis">
|
||
<emphasis id="B95B4F7A1939FFBBFD13B143FD4C3047" box="[640,764,277,299]" italics="true" pageId="9" pageNumber="1204">V. laosensis</emphasis>
|
||
</taxonomicName>
|
||
, and at this occasion he transfered
|
||
<taxonomicName id="4C2FE8EB1939FFBBFF7FB167FE213024" authorityName="JEDLI C KA" authorityYear="1956" box="[236,401,306,328]" class="Insecta" family="Carabidae" genus="Chlaenius" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="1204" phylum="Arthropoda" rank="species" species="klapperichi">
|
||
<emphasis id="B95B4F7A1939FFBBFF7FB167FE213024" box="[236,401,306,328]" italics="true" pageId="9" pageNumber="1204">Chl. klapperichi</emphasis>
|
||
</taxonomicName>
|
||
to the genus
|
||
<emphasis id="B95B4F7A1939FFBBFD8BB167FDCD3024" box="[536,637,306,328]" italics="true" pageId="9" pageNumber="1204">Vachinius</emphasis>
|
||
, subgenus
|
||
<emphasis id="B95B4F7A1939FFBBFD7CB167FC373024" box="[751,903,306,328]" italics="true" pageId="9" pageNumber="1204">Sphodromimus</emphasis>
|
||
<bibRefCitation id="EFBEEE991939FFBBFC1FB167FBA93024" author="CASALE A" box="[908,1049,306,328]" pageId="9" pageNumber="1204" pagination="371 - 382" refId="ref5537" refString="CASALE A. (1984): The new Asiatic genus Vachinius (Carabidae: Callistinae) with three new species. - Bollettino del Museo Regionale di Scienze Naturali, Torino 2: 371 - 382." type="journal article" year="1984">CASALE 1984</bibRefCitation>
|
||
, on the basis of exoskeletal characters ("aufgrund der ektoskelettalen Merkmale") but without conclusively substantiated discussion. As the problem of the systematic position of
|
||
<emphasis id="B95B4F7A1939FFBBFF21B1DFFEA730CC" box="[178,279,394,416]" italics="true" pageId="9" pageNumber="1204">Vachinius</emphasis>
|
||
is not yet solved this decision is not followed here.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |