A new species of genus Chlaenius BONELLI 1810, subgenus Haplochlaenius LUTSHNIK 1933 from China (Coleoptera, Carabidae, Chlaeniini) and notes on two species previously described
Author
D. W
text
Linzer biologische Beiträge
2012
2012-12-28
44
2
1195
1205
journal article
10.5281/zenodo.5329203
0253-116X
5329203
Chlaenius
(
Haplochlaenius
)
nanlingensis
DEUVE & TIAN 2005
M a t e r i a l e x a m i n e d:
China
:
Guangxi
:1, 7: NE
Guangxi
, Xing`an Co., Mao’er Shan, Mao`er Mountain Nature Reserve,
823 m
, N 25.51.497’/E 110.29.803’,
V 2007
, M. Häckel & R. Sehnal leg. (cHÄCK, cSEHN, cWR, NHW). 1, 4: Mao’ er Shan`,
493 m
/ N 25.51.11’/ E 110. 29.15’, R. Sehnal leg. (cSEHN, cWR). 2: Mao’er Shan,
800-1100 m
,
25°52N
/
110°29E
(dense primary forest with strong bamboo, steep slopes),
28.V.-9.VI.2009
, Chr. Reuter leg. (cREUT, cWR). -
Guizhou
: 2: SE.
Guizhou
, Dushan, Gengding Shan env.,
1445 m
,
N 25°52.5‘
/
E 107°38’
, M. Häckel & R. Sehnal leg. (cSEHN, cWR.)
The description based on
one male
, coming from the slopes of the Mount Shikengkong situated in the Nanling Nature reserve (Dadongshan, Lianzhou) in the Chinese Province of
Guangdong
, close to the border to
Hunan
. The material from the Mao’er Shan fits very well the description, and, though not having investigated the type which is stored in the collection of the South
China
Agricultural University, I refer the name
Chl
.
nanlingensis
to the material noted above. A habitus photograph of the
holotype
kindly provided by M. Tian enabled the species to be recognised beyond doubt without having to examine the specimen itself. The species is brachypterous (this fact was not mentioned in the description),
one specimen
investigated has the hindwings hardly as long as half of elytral length, but it is obviously able to have a wider range, the distance between the Mount Shikengkong and the Mao’er Shan is about
270 km
, with a landscape surely forming no considerable geographical barriers which could stop an expansion of brachypterous species, also the findings in the Gengding Shan in the southeast of the Province Guizou, still more north-westwards, speaks for this assumption. Just recently I received some habitus photographs of specimens stored in the Institute of Zoology, Chinese Academy of Sciences,
Beijing
and identified as
Chl. nanlingensis
, coming from
Hunan
(Shaoshan-shihu) and from
Sichuan
(Ya’an), confirming that the species has a wider range than originally thought. New records for the Chinese provinces of
Guangxi
,
Guizhou
,
Hunan
, and
Sichuan
.
N o t e s: The genus
Vachinius
CASALE 1984
was founded on several species:
V. subglaber
(ANDREWES 1937)
,
type
species from
Sikkim
,
V. pseudoglaber
CASALE 1984
, from northern
India
,
V. holzschuhi
CASALE 1984
from eastern
Nepal
, and
V. pilosus
CASALE 1984
from
Vietnam
, underlining that these species are typical
Chlaeniini
, near to species of the genus
Macrochlaenites
KUNTZEN 1919
but are distinguished by having a cordate pronotum, strongly sinuate toward base, a dimorphic apex and integument of elytra in males and females, a different construction of the male genitalia, and reduced wings. Later, further species were described, one from
Thailand
:
V. thailandensis
MORVAN 1991
, three from
China
:
V. deuvei
MORVAN 1997
(
Guangxi
)
,
V. hunanus
MORVAN 1997
(
Hunan
)
, and
V. wrasei
KIRSCHENHOFER 2003
(
Guangdong
)
, one from Sulawesi:
V. baehri
KIRSCHENHOFER 1998
, and one from
Burma
:
V. burmanensis
LASSALLE 2001
. Investigating a larger number of
Chlaenius
species
belonging to subgenus
Haplochlaenius
LUTSHNIK 1933
(a part of them placed under
Macrochlaenites
in older taxonomic classifications) I found partly the same characters peculiar to species of
Vachinius
, such as the construction of the mentum, the sinuate pronotum, the partly occurring carinate elytral intervals, a reduction of the hindwings and the morphology of the male genitalia. The classification of the new species described above posed problems and I decided to describe it in the genus
Chlaenius
, subgenus
Haplochlaenius
. So it seems necessary to clarify the systematic position and the taxonomic rank of the genus
Vachinius
within the tribe
Chlaeniini
and especially its relationship to
Haplochlaenius
by careful examination of all taxa (see also the remarks in DEUVE & TIAN 2005), including also genetic techniques. Just recently
KIRSCHENHOFER (2012)
described two further
Vachinius
species from
Laos
:
V. hajeki
and
V. laosensis
, and at this occasion he transfered
Chl. klapperichi
to the genus
Vachinius
, subgenus
Sphodromimus
CASALE 1984
, on the basis of exoskeletal characters ("aufgrund der ektoskelettalen Merkmale") but without conclusively substantiated discussion. As the problem of the systematic position of
Vachinius
is not yet solved this decision is not followed here.