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<document id="AB78C42961A3FD1B2DEC45384665FF85" ID-DOI="10.5281/zenodo.192017" ID-GBIF-Dataset="5b26ee34-55ff-4da4-a200-444541506cd0" ID-ISSN="1175-5326" ID-Zenodo-Dep="192017" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461140117641" checkinUser="plazi" docAuthor="Imura, Jimpei &amp; Ishikawa, Tadashi" docDate="2009" docId="03CE87E8AB276767FF6E840E10DEFB95" docLanguage="en" docName="zt02315p030.pdf" docOrigin="Zootaxa 2315" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Schiodtella Signoret 1882" docType="treatment" docVersion="9" lastPageNumber="21" masterDocId="FFF7FF90AB266765FFF98370112EFFA2" masterDocTitle="A review of Schiodtella formosana, with some remarks on the genus Schiodtella and description of a Japanese new species (Hemiptera: Heteroptera: Cydnidae)" masterLastPageNumber="30" masterPageNumber="19" pageNumber="20" updateTime="1698598840229" updateUser="plazi">
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<mods:title id="7FA4A0B3836DEFE98F9D1131E0C09A79">A review of Schiodtella formosana, with some remarks on the genus Schiodtella and description of a Japanese new species (Hemiptera: Heteroptera: Cydnidae)</mods:title>
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<mods:namePart id="B56101129E70AA8EA9F2404CBC184FEC">Imura, Jimpei</mods:namePart>
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<mods:namePart id="B000128797FA8827072DC1E5B5ED99FA">Ishikawa, Tadashi</mods:namePart>
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<treatment id="03CE87E8AB276767FF6E840E10DEFB95" ID-DOI="http://doi.org/10.5281/zenodo.6226893" ID-GBIF-Taxon="119627846" ID-Zenodo-Dep="6226893" LSID="urn:lsid:plazi:treatment:03CE87E8AB276767FF6E840E10DEFB95" httpUri="http://treatment.plazi.org/id/03CE87E8AB276767FF6E840E10DEFB95" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<subSubSection id="C37D6575AB276764FF6E840E1371F83A" box="[151,607,1918,1944]" pageId="1" pageNumber="20" type="nomenclature">
<paragraph id="8BD836FEAB276764FF6E840E1371F83A" blockId="1.[151,607,1918,1944]" box="[151,607,1918,1944]" pageId="1" pageNumber="20">
<heading id="D0908192AB276764FF6E840E1371F83A" bold="true" box="[151,607,1918,1944]" fontSize="11" level="1" pageId="1" pageNumber="20" reason="1">
<emphasis id="B913EAECAB276764FF6E840E1371F83A" bold="true" box="[151,607,1918,1944]" pageId="1" pageNumber="20">
The genus
<taxonomicName id="4C674D7DAB276764FEE7840E1371F83A" authority="Signoret, 1882" authorityName="Signoret" authorityYear="1882" box="[286,607,1918,1944]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB276764FEE7840E108BF83A" bold="true" box="[286,421,1918,1944]" italics="true" pageId="1" pageNumber="20">Schiodtella</emphasis>
<bibRefCitation id="EFF64B0FAB276764FE55840E1371F83A" author="Signoret" box="[428,607,1918,1944]" pageId="1" pageNumber="20" refString="Signoret, V (1882) Note. Bulletin de la Societe Entomologique de France 1881, 218 - 219 [also published in Annales de la Societe Entomologique de France (6), 1 (1881), clvi - clviii]." type="journal article" year="1882">Signoret, 1882</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
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<paragraph id="8BD836FEAB276767FF6E84B81532FF0D" blockId="1.[151,1436,1992,2015]" lastBlockId="2.[151,1436,152,271]" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<treatmentCitationGroup id="AB7711D0AB276767FF6E84B81532FF0D" lastPageId="2" lastPageNumber="21" pageId="1" pageNumber="20">
<taxonomicName id="4C674D7DAB276764FF6E84B810DEF87D" authority="Signoret, 1882: 218" authorityName="Signoret" authorityPageNumber="218" authorityYear="1882" box="[151,496,1992,2015]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB276764FF6E84B81021F87D" box="[151,271,1992,2015]" italics="true" pageId="1" pageNumber="20">Schiodtella</emphasis>
<treatmentCitation id="0AC610EFAB276764FEEE84B810DEF87D" author="Signoret" box="[279,496,1992,2015]" page="218" pageId="1" pageNumber="20" year="1882">
<bibRefCitation id="EFF64B0FAB276764FEEE84B81097F87D" author="Signoret" box="[279,441,1992,2015]" pageId="1" pageNumber="20" refString="Signoret, V (1882) Note. Bulletin de la Societe Entomologique de France 1881, 218 - 219 [also published in Annales de la Societe Entomologique de France (6), 1 (1881), clvi - clviii]." type="journal article" year="1882">Signoret, 1882</bibRefCitation>
: 218
</treatmentCitation>
</taxonomicName>
,
<typeStatus id="54DC885CAB276764FE0584B81304F87D" box="[508,554,1992,2015]" pageId="1" pageNumber="20">type</typeStatus>
species by subsequent designation (
<treatmentCitation id="0AC610EFAB276764FC4C84B8156BF87D" author="Lis" box="[949,1093,1992,2015]" page="25" pageId="1" pageNumber="20" year="1994">
<bibRefCitation id="EFF64B0FAB276764FC4C84B8156BF87D" author="Lis" box="[949,1093,1992,2015]" pageId="1" pageNumber="20" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae). Upper Silesian Museum, Bytom. 349 pp." type="book" year="1994">Lis 1994: 25</bibRefCitation>
</treatmentCitation>
):
<taxonomicName id="4C674D7DAB276764FBA084B814B2F87D" authority="Schiodte" authorityName="Schiodte" box="[1113,1436,1992,2015]" class="Insecta" family="Cydnidae" genus="Stibaropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="20" phylum="Arthropoda" rank="species" species="tabulatus">
<emphasis id="B913EAECAB276764FBA084B81418F87D" box="[1113,1334,1992,2015]" italics="true" pageId="1" pageNumber="20">Stibaropus tabulatus</emphasis>
Schiødte
</taxonomicName>
(syn. by
<treatmentCitation id="0AC610EFAB246767FEDA83E81082FF0D" author="Lis" box="[291,428,152,175]" page="24" pageId="2" pageNumber="21" year="1994">
<bibRefCitation id="EFF64B0FAB246767FEDA83E810ABFF0D" author="Lis" box="[291,389,152,175]" pageId="2" pageNumber="21" refString="Lis, J. A. (1994) A revision of Oriental burrower bugs (Heteroptera: Cydnidae). Upper Silesian Museum, Bytom. 349 pp." type="book" year="1994">Lis 1994</bibRefCitation>
: 24
</treatmentCitation>
, with
<taxonomicName id="4C674D7DAB246767FE1483E813C1FF0D" authority="Dallas, 1851" authorityName="Dallas" authorityYear="1851" box="[493,751,152,175]" class="Insecta" family="Cydnidae" genus="Stibaropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FE1483E81371FF0D" box="[493,607,152,175]" italics="true" pageId="2" pageNumber="21">Stibaropus</emphasis>
<bibRefCitation id="EFF64B0FAB246767FD9F83E813C1FF0D" author="Dallas" box="[614,751,152,175]" pageId="2" pageNumber="21" refString="Dallas, W. S. (1851) List of specimens of hemipterous insects in the collection of the British Museum. Part I. Taylor, London, 368 pp, 11 pls." type="book" year="1851">Dallas, 1851</bibRefCitation>
</taxonomicName>
; restored by Lis 1999: 128).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD836FEAB246767FF6E83C8105AFEAD" blockId="2.[151,1436,152,271]" pageId="2" pageNumber="21">
<emphasis id="B913EAECAB246767FF6E83C8101FFF6D" box="[151,305,184,207]" italics="true" pageId="2" pageNumber="21">Neostibaropus</emphasis>
<bibRefCitation id="EFF64B0FAB246767FEC283C813ACFF6D" author="Takano" box="[315,642,184,207]" pageId="2" pageNumber="21" refString="Takano, S. &amp; Yanagihara, M. (1939) Researches on injurious and beneficial animals of sugarcane. Taiwan Sugar Experimental Station, Extra Reports, 2, 311 pp., 18 pls." type="book" year="1939">Takano and Yanagihara, 1939</bibRefCitation>
: 135,
<typeStatus id="54DC885CAB246767FD3E83C813DBFF6D" box="[711,757,184,207]" pageId="2" pageNumber="21">type</typeStatus>
species by monotypy:
<taxonomicName id="4C674D7DAB246767FC0C83C81011FF4D" authority="Takano and Yanagihara" authorityName="Takano and Yanagihara" class="Insecta" family="Cydnidae" genus="Neostibaropus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="species" species="formosanus">
<emphasis id="B913EAECAB246767FC0C83C8143BFF6D" box="[1013,1301,184,207]" italics="true" pageId="2" pageNumber="21">Neostibaropus formosanus</emphasis>
Takano and Yanagihara
</taxonomicName>
(syn. by
<bibRefCitation id="EFF64B0FAB246767FE5F83A813B2FF4D" author="Esaki" box="[422,668,216,239]" pageId="2" pageNumber="21" refString="Esaki, T. &amp; Ishihara, T (1951) Hemipetra of Shansi, North China. II. Pentatomoidea. Mushi, 22 (5), 22 - 44." type="journal article" year="1951">Esaki &amp; Ishihara 1951</bibRefCitation>
: 30, with
<taxonomicName id="4C674D7DAB246767FCF183A81522FF4D" authority="Dallas, 1851" authorityName="Dallas" authorityYear="1851" box="[776,1036,216,239]" class="Insecta" family="Cydnidae" genus="Stibaropus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FCF183A81254FF4D" box="[776,890,216,239]" italics="true" pageId="2" pageNumber="21">Stibaropus</emphasis>
<bibRefCitation id="EFF64B0FAB246767FC7B83A81522FF4D" author="Dallas" box="[898,1036,216,239]" pageId="2" pageNumber="21" refString="Dallas, W. S. (1851) List of specimens of hemipterous insects in the collection of the British Museum. Part I. Taylor, London, 368 pp, 11 pls." type="book" year="1851">Dallas, 1851</bibRefCitation>
</taxonomicName>
; by
<bibRefCitation id="EFF64B0FAB246767FBC583A81585FF4D" author="Lis" box="[1084,1195,216,239]" pageId="2" pageNumber="21" refString="Lis, J. A. (1991 a) Studies on Oriental Cydnidae IV. New species, new synonyms and new records (Heteroptera). Annals of the Upper Silesian Museum, 2: 165 - 190." type="journal article" year="1991" yearSuffix="a">Lis 1991a</bibRefCitation>
: 169, with
<taxonomicName id="4C674D7DAB246767FADD83A81046FEAD" authority="Signoret, 1882" authorityName="Signoret" authorityYear="1882" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FADD83A814B2FF4D" box="[1316,1436,216,239]" italics="true" pageId="2" pageNumber="21">Schiodtella</emphasis>
<bibRefCitation id="EFF64B0FAB246767FF3E83881046FEAD" author="Signoret" box="[199,360,248,271]" pageId="2" pageNumber="21" refString="Signoret, V (1882) Note. Bulletin de la Societe Entomologique de France 1881, 218 - 219 [also published in Annales de la Societe Entomologique de France (6), 1 (1881), clvi - clviii]." type="journal article" year="1882">Signoret, 1882</bibRefCitation>
</taxonomicName>
).
</paragraph>
<paragraph id="8BD836FEAB246767FF6E824E135DFC8A" blockId="2.[151,1437,318,1079]" pageId="2" pageNumber="21">
<bibRefCitation id="EFF64B0FAB246767FF6E824E100FFEFA" author="Lis" box="[151,289,318,344]" pageId="2" pageNumber="21" refString="Lis, J. A. (1991 a) Studies on Oriental Cydnidae IV. New species, new synonyms and new records (Heteroptera). Annals of the Upper Silesian Museum, 2: 165 - 190." type="journal article" year="1991" yearSuffix="a">Lis (1991a)</bibRefCitation>
synonymized
<emphasis id="B913EAECAB246767FE36824F1354FEFA" box="[463,634,319,344]" italics="true" pageId="2" pageNumber="21">Neostibaropus</emphasis>
with
<taxonomicName id="4C674D7DAB246767FD46824F126BFEFA" box="[703,837,319,344]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FD46824F126BFEFA" box="[703,837,319,344]" italics="true" pageId="2" pageNumber="21">Schiodtella</emphasis>
</taxonomicName>
chiefly on the basis of having the four-segmented antennae, and afterwards, diagnosed the latter as “Head narrowed anteriorly, crenulated on margins; paraclypei with peg-like setae; clypeus parallel-sided or narrowed apically, shorter than paraclypei; antennae 4-segmented, second segment as long as or longer than the third; second rostral segment simple, without foliaceous lobe; apex of peritreme with small tooth-like process; margin of abdominal sternites straight in lateral parts; tarsi of all leg present” (Lis 1999). In addition to these characters, the 1+1 abdominal trichobothrial formula on the sternites III to VII is of taxonomic value for
<taxonomicName id="4C674D7DAB246767FBF2815715BEFDE2" box="[1035,1168,551,576]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FBF2815715BEFDE2" box="[1035,1168,551,576]" italics="true" pageId="2" pageNumber="21">Schiodtella</emphasis>
</taxonomicName>
as documented by
<bibRefCitation id="EFF64B0FAB246767FA8F81561332FDC5" author="Lis" pageId="2" pageNumber="21" refString="Lis, J. A. &amp; Hohol-Kilinkiewicz, A. (2001) Abdominal trichobothrial pattern and its taxonomic and phylogenetic significance in Cephalocteinae (Hemiptera: Heteroptera: Cydnidae). Annales zoologici (Warszawa), 51 (3), 293 - 297." type="journal article" year="2001">Lis and Hohol-Kilinkiewicz (2001)</bibRefCitation>
. As
<emphasis id="B913EAECAB246767FDAA813D122DFDC4" box="[595,771,589,614]" italics="true" pageId="2" pageNumber="21">Neostibaropus</emphasis>
is clearly ascertained to have no tarsi, although it has almost all characters above, it is tempting to restitute
<emphasis id="B913EAECAB246767FCDC810412FAFD2F" box="[805,980,628,653]" italics="true" pageId="2" pageNumber="21">Neostibaropus</emphasis>
from the synonymy as a valid genus. However, we believe that maintainance of
<emphasis id="B913EAECAB246767FD5981EA1261FD11" box="[672,847,666,691]" italics="true" pageId="2" pageNumber="21">Neostibaropus</emphasis>
as a synonym of
<taxonomicName id="4C674D7DAB246767FBDE81EA1581FD11" box="[1063,1199,666,691]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FBDE81EA1581FD11" box="[1063,1199,666,691]" italics="true" pageId="2" pageNumber="21">Schiodtella</emphasis>
</taxonomicName>
, based on a careful survey on the comprehensive morphology among the species of
<taxonomicName id="4C674D7DAB246767FC6081B11530FD78" box="[921,1054,705,730]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FC6081B11530FD78" box="[921,1054,705,730]" italics="true" pageId="2" pageNumber="21">Schiodtella</emphasis>
</taxonomicName>
and its allies (see below for the detailed discussion), should be kept. Herein we provide a modified diagnosis for
<taxonomicName id="4C674D7DAB246767FB9B819815C9FCA3" box="[1122,1255,744,769]" class="Insecta" family="Cydnidae" genus="Schiodtella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B913EAECAB246767FB9B819815C9FCA3" box="[1122,1255,744,769]" italics="true" pageId="2" pageNumber="21">Schiodtella</emphasis>
</taxonomicName>
, and add some characters that are important to define it.
</paragraph>
</subSubSection>
<subSubSection id="C37D6575AB246767FF3F804510DEFB95" pageId="2" pageNumber="21" type="diagnosis">
<paragraph id="8BD836FEAB246767FF3F804510DEFB95" blockId="2.[151,1437,318,1079]" pageId="2" pageNumber="21">
<emphasis id="B913EAECAB246767FF3F80451069FCED" bold="true" box="[198,327,821,847]" italics="true" pageId="2" pageNumber="21">Diagnosis.</emphasis>
Head narrowed anteriorly, crenulated by fringe of peg-bearing tubercles on paraclypeal margin; clypeus parallel-sided or narrowed apically, shorter than paraclypei, with 2 subapical pegs. Antennae 4-segmented; segment II as long as or longer than segment III. Rostral segment II simple, without foliaceous lobe. Apical process of peritreme usually tooth-like, variable in length, but sometimes lacking. Tarsi present or absent. Posterior margin of abdominal sterna not insinuated; trichobothrial formula 1+1 on sternites III to VII (
<figureCitation id="135C2A7BAB246767FF2880861005FBB2" box="[209,299,1014,1040]" captionStart="FIGURE 17 21" captionStartId="7.[151,258,1851,1875]" captionTargetBox="[161,1434,836,1789]" captionTargetId="figure@7.[151,1436,815,1825]" captionTargetPageId="7" captionText="FIGURE 17 21. Schiodtella formosana. 17, dorsal aspect; 18 19, head, dorsal (18) and lateral (19) views; 20, left evaporatorium; 21, left side of abdominal sterna III VII, lateral view. Abbreviations: Sp, spiracle; Trb, trichobothrium. Scales: 1.0 mm for 17 &amp; 21, 0.5 mm for 18 20." httpUri="https://zenodo.org/record/192021/files/figure.png" pageId="2" pageNumber="21">Fig. 21</figureCitation>
). Penis with conjunctival appendage of various lengths; spermathecal duct with large, globate, distinctly pigmented dilation.
</paragraph>
</subSubSection>
</treatment>
</document>