220 lines
34 KiB
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220 lines
34 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e95181" ID-Pensoft-Pub="1864-8312-81-317" ID-Pensoft-UUID="C3B28C3FAD9250BC906A4A865EAA18B4" ID-ZooBank="BE57FF40714740AC8BEA1706A95DA4ED" ModsDocID="1864-8312-81-317" checkinTime="1680679737344" checkinUser="pensoft" docAuthor="Botero-Trujillo, Ricardo, Martinez, Leonel, Iuri, Hernan Augusto, Ojanguren-Affilastro, Andres Alejandro & Carvalho, Leonardo Sousa" docDate="2023" docId="F564A87D66755CB68D71A53BECEB9B0C" docLanguage="en" docName="ArthropodSystPhyl 81: 317-344" docOrigin="Arthropod Systematics & amp; Phylogeny 81" docPubDate="2023-04-04" docSource="http://dx.doi.org/10.3897/asp.81.e95181" docTitle="Eutrecha Maury 1982" docType="treatment" docVersion="1" id="C3B28C3FAD9250BC906A4A865EAA18B4" lastPageNumber="317" masterDocId="C3B28C3FAD9250BC906A4A865EAA18B4" masterDocTitle="Revision of the genera Eutrecha and Xenotrecha (Solifugae: Ammotrechidae), taxonomic notes on Ammotrechinae, and description of a remarkable new Eutrecha from Colombia" masterLastPageNumber="344" masterPageNumber="317" pageNumber="317" updateTime="1680679737344" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the genera Eutrecha and Xenotrecha (Solifugae: Ammotrechidae), taxonomic notes on Ammotrechinae, and description of a remarkable new Eutrecha from Colombia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Botero-Trujillo, Ricardo</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6199-6572</mods:nameIdentifier>
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<mods:affiliation>Division of Invertebrate Zoology, American Museum of Natural History, Central Park West at 79 th Street, New York, NY 10024 - 5192, USA & Division Aracnologia, Museo Argentino de Ciencias Naturales " Bernardino Rivadavia " - CONICET, Avenida Angel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Martinez, Leonel</mods:namePart>
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<mods:affiliation>Division Aracnologia, Museo Argentino de Ciencias Naturales " Bernardino Rivadavia " - CONICET, Avenida Angel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Iuri, Hernan Augusto</mods:namePart>
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<mods:affiliation>Division Aracnologia, Museo Argentino de Ciencias Naturales " Bernardino Rivadavia " - CONICET, Avenida Angel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ojanguren-Affilastro, Andres Alejandro</mods:namePart>
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<mods:affiliation>Division Aracnologia, Museo Argentino de Ciencias Naturales " Bernardino Rivadavia " - CONICET, Avenida Angel Gallardo 470, CP: 1405 DJR, C. A. B. A., Buenos Aires, Argentina</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Carvalho, Leonardo Sousa</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-4700-5610</mods:nameIdentifier>
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<mods:affiliation>Universidade Federal do Piaui, Campus Amilcar Ferreira Sobral, BR 343, km 3.5, Bairro Meladao, s / no. CEP 64800 - 000, Floriano, PI, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">carvalho@ufpi.edu.br</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-04-04</mods:number>
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<mods:detail type="volume">
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<mods:number>81</mods:number>
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<mods:start>317</mods:start>
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<mods:end>344</mods:end>
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<mods:url>http://dx.doi.org/10.3897/asp.81.e95181</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/asp.81.e95181</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1864-8312-81-317</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:5DD9F28C-C72D-4402-B6FD-D3B43A1E4384" httpUri="http://treatment.plazi.org/id/F564A87D66755CB68D71A53BECEB9B0C" lastPageNumber="317" pageId="0" pageNumber="317">
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<subSubSection pageId="0" pageNumber="317" type="nomenclature">
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<paragraph pageId="0" pageNumber="317">
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3.1.2.
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<taxonomicName ID-CoL="634WN" LSID="http://zoobank.org/5DD9F28C-C72D-4402-B6FD-D3B43A1E4384" authority="Maury, 1982" authorityName="Maury" authorityYear="1982" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha" order="Solifugae" pageId="0" pageNumber="317" phylum="Arthropoda" rank="genus">Eutrecha Maury, 1982</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="317" type="description">
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<paragraph pageId="0" pageNumber="317">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Known records of Eutrecha Maury, 1982 (squares, circles and pentagons) and Xenotrecha Maury, 1982 (crosses). Records with question mark correspond to unconfirmed record from Suriname and the unprecise type locality of X. huebneri in the headwaters of the Orinoco River, in Venezuela. See text for details." figureDoi="10.3897/asp.81.e95181.figure1" httpUri="https://binary.pensoft.net/fig/833383" pageId="0" pageNumber="317">Figs 1</figureCitation>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eutrecha Maury, 1982, schematic representation of cheliceral morphology, retrolateral aspect. A Eutrecha longirostris Maury, 1982, holotype ♂ (MAGS 167). B Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia. C Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008). Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MRLC, movable finger, retrolateral carina; MRVC, movable finger, retroventral longitudinal carina; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth. Arrowheads in B indicate extremely reduced FSM, FM, and FD teeth." figureDoi="10.3897/asp.81.e95181.figure2" httpUri="https://binary.pensoft.net/fig/833384" pageId="0" pageNumber="317">, 2</figureCitation>
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), habitus, dorsal aspect (A), sinistral pedipalp basitarsus and telotarsus, prolateral (B) and ventral (C) aspects, anterior opisthosomal sternites, with spiracular sternites centered (D). Scale bars = 2 mm (A, D), 1 mm (B, C). Arrows: spiniform setae (in B and C); ctenidia on 1 st post-genital sternite (in D)." figureDoi="10.3897/asp.81.e95181.figure4" httpUri="https://binary.pensoft.net/fig/833386" pageId="0" pageNumber="317">, 4</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers and stridulatory plate (D). Scale bars = 1 mm." figureDoi="10.3897/asp.81.e95181.figure5" httpUri="https://binary.pensoft.net/fig/833387" pageId="0" pageNumber="317">, 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, fixed finger, ventral aspect (A), schematic representation of fixed and movable fingers dentition, proventral aspect (B). Scale bars = 0.5 mm. Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MPL, movable finger, prolateral tooth; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSM, profondal submedial tooth; PFSP, profondal subproximal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth." figureDoi="10.3897/asp.81.e95181.figure6" httpUri="https://binary.pensoft.net/fig/833388" pageId="0" pageNumber="317">, 6</figureCitation>
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, propeltidium (A), anterior opisthosomal sternites with spiracular sternites centered (B) [arrows indicate paramedian areas with ctenidia on 1 st post-genital sternite], dextral chelicera, stridulatory plate, prolateral aspect (C) [note the modified stridulatory apparatus]. Scale bars = 1 mm (A, C), 0.5 mm (B)." figureDoi="10.3897/asp.81.e95181.figure7" httpUri="https://binary.pensoft.net/fig/833389" pageId="0" pageNumber="317">, 7</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers (D). Scale bars = 1 mm (A, C), 0.5 mm (B, D)." figureDoi="10.3897/asp.81.e95181.figure8" httpUri="https://binary.pensoft.net/fig/833390" pageId="0" pageNumber="317">, 8</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, sinistral chelicera, retrolateral aspect (A), prolateral aspect (B), and ventral aspect of fixed finger (C), dextral chelicera, fixed finger, ventral aspect (D). A - C ♀ (MPUJ ENT 61896), San Jacinto, Bolivar Department, Colombia. D ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia. Scale bars = 1 mm (A, B), 0.5 mm (C, D)." figureDoi="10.3897/asp.81.e95181.figure9" httpUri="https://binary.pensoft.net/fig/833391" pageId="0" pageNumber="317">, 9</figureCitation>
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Landscape (A) and habitat (B) of Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017 at type locality in San Jacinto, Bolivar Department, Colombia." figureDoi="10.3897/asp.81.e95181.figure10" httpUri="https://binary.pensoft.net/fig/833392" pageId="0" pageNumber="317">, 10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="317" type="reference_group">
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<paragraph pageId="0" pageNumber="317">
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<taxonomicName authorityName="Maury" authorityYear="1982" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha" order="Solifugae" pageId="0" pageNumber="317" phylum="Arthropoda" rank="genus">Eutrecha</taxonomicName>
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Maury, 1982: 125, 138;
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<bibRefCitation author="Harvey, MS" journalOrPublisher="CSIRO Publishing, Collingwood, Victoria, Australia. First Edition" pageId="0" pageNumber="317" refId="B18" refString="Harvey, MS, 2003. Catalogue of the Smaller Arachnid Orders of the World. CSIRO Publishing, Collingwood, Victoria, Australia. First Edition" title="Catalogue of the Smaller Arachnid Orders of the World." year="2003">Harvey 2003</bibRefCitation>
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: 210;
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<bibRefCitation author="Hernandez, HLA" journalOrPublisher="CSIRO Publishing, Collingwood, Victoria, Australia. First Edition" pageId="0" pageNumber="317" publicationUrl="http://sea-entomologia.org/PDF/BSEA42ARACNO/B42447.pdf" refId="B19" refString="Hernandez, HLA, Colmenares, GPA, 2008. Segundo registro de Eutrecha longirostris Maury, 1982 (Solifugae: Ammotrechidae) de Venezuela. Boletin de la Sociedad Entomologica Aragonesa 42: 447-448. http://sea-entomologia.org/PDF/BSEA42ARACNO/B42447.pdf" title="Segundo registro de Eutrecha longirostris Maury, 1982 (Solifugae: Ammotrechidae) de Venezuela. Boletin de la Sociedad Entomologica Aragonesa 42: 447 - 448." url="http://sea-entomologia.org/PDF/BSEA42ARACNO/B42447.pdf" year="2008">
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<normalizedToken originalValue="Hernández">Hernandez</normalizedToken>
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and Colmenares 2008
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</bibRefCitation>
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: 447;
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<bibRefCitation author="Acosta-Berrocal, M" journalOrPublisher="Revista Iberica de Aracnologia" pageId="0" pageNumber="317" pagination="64 - 70" publicationUrl="http://sea-entomologia.org/PDF/BSEA49ARACNO/B49083.pdf" refId="B1" refString="Acosta-Berrocal, M, Bedoya-Roqueme, E, Salleg-Perez, G, Quiros-Rodriguez, JA, 2017. Una especie nueva de Ammotrechella Roewer 1934 (Solifugae: Ammotrechidae), de Colombia. Revista Iberica de Aracnologia 31: 64 - 70, http://sea-entomologia.org/PDF/BSEA49ARACNO/B49083.pdf" title="Una especie nueva de Ammotrechella Roewer 1934 (Solifugae: Ammotrechidae), de Colombia." url="http://sea-entomologia.org/PDF/BSEA49ARACNO/B49083.pdf" volume="31" year="2017">Acosta-Berrocal et al. 2017</bibRefCitation>
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: 64;
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<bibRefCitation author="Villareal-Blanco, E" journalOrPublisher="Revista Iberica de Aracnologia" pageId="0" pageNumber="317" pagination="139 - 143" refId="B40" refString="Villareal-Blanco, E, Armas, LF, Martinez, L, 2017. Una especie nueva del genero Eutrecha Maury, 1982 del Caribe Colombiano (Solifugae: Ammotrechidae). Revista Iberica de Aracnologia 30: 139 - 143" title="Una especie nueva del genero Eutrecha Maury, 1982 del Caribe Colombiano (Solifugae: Ammotrechidae)." volume="30" year="2017">Villareal-Blanco et al. 2017</bibRefCitation>
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: 139, 142, fig. 15;
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<bibRefCitation DOI="https://doi.org/10.1016/j.jcz.2021.10.001" author="Iuri, HA" journalOrPublisher="Zoologischer Anzeiger" pageId="0" pageNumber="317" pagination="126 - 155" refId="B21" refString="Iuri, HA, Ramirez, MJ, Mattoni, CI, Ojanguren-Affilastro, AA, 2021. Revision and cladistic analysis of subfamily Nothopuginae (Solifugae, Ammotrechidae). Zoologischer Anzeiger 295: 126 - 155, DOI: https://doi.org/10.1016/j.jcz.2021.10.001" title="Revision and cladistic analysis of subfamily Nothopuginae (Solifugae, Ammotrechidae)." url="https://doi.org/10.1016/j.jcz.2021.10.001" volume="295" year="2021">Iuri et al. 2021</bibRefCitation>
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: 153.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="317" type="revised diagnosis">
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<paragraph pageId="0" pageNumber="317">Revised diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="317">
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A member of
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<taxonomicName authorityName="Roewer" authorityYear="1934" family="Ammotrechidae" kingdom="Animalia" lsidName="" order="Solifugae" pageId="0" pageNumber="317" rank="subFamily" subFamily="Ammotrechinae">Ammotrechinae</taxonomicName>
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s.str. as herein defined.
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<taxonomicName authorityName="Maury" authorityYear="1982" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha" order="Pararthropoda" pageId="0" pageNumber="317" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="317">Eutrecha</emphasis>
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</taxonomicName>
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is presumed to be a monophyletic group of solifuges, whose males exhibit the following combination of features that allows to readily distinguish the genus from all other genera in
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<taxonomicName authorityName="Roewer" authorityYear="1934" family="Ammotrechidae" kingdom="Animalia" lsidName="" order="Solifugae" pageId="0" pageNumber="317" rank="subFamily" subFamily="Ammotrechinae">Ammotrechinae</taxonomicName>
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s.l.
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<emphasis bold="true" pageId="0" pageNumber="317">i)</emphasis>
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Cheliceral fixed finger with FSM, FM, and FD teeth moderately reduced (i.e., evidently visible but smaller than MSM tooth of movable finger, in
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<taxonomicName lsidName="E. belenensis" pageId="0" pageNumber="317" rank="species" species="belenensis">
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<emphasis italics="true" pageId="0" pageNumber="317">E. belenensis</emphasis>
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</taxonomicName>
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) (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eutrecha Maury, 1982, schematic representation of cheliceral morphology, retrolateral aspect. A Eutrecha longirostris Maury, 1982, holotype ♂ (MAGS 167). B Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia. C Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008). Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MRLC, movable finger, retrolateral carina; MRVC, movable finger, retroventral longitudinal carina; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth. Arrowheads in B indicate extremely reduced FSM, FM, and FD teeth." figureDoi="10.3897/asp.81.e95181.figure2" httpUri="https://binary.pensoft.net/fig/833384" pageId="0" pageNumber="317">2C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers and stridulatory plate (D). Scale bars = 1 mm." figureDoi="10.3897/asp.81.e95181.figure5" httpUri="https://binary.pensoft.net/fig/833387" pageId="0" pageNumber="317">5B</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, fixed finger, ventral aspect (A), schematic representation of fixed and movable fingers dentition, proventral aspect (B). Scale bars = 0.5 mm. Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MPL, movable finger, prolateral tooth; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSM, profondal submedial tooth; PFSP, profondal subproximal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth." figureDoi="10.3897/asp.81.e95181.figure6" httpUri="https://binary.pensoft.net/fig/833388" pageId="0" pageNumber="317">6</figureCitation>
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) to much reduced or undistinguishable (
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<taxonomicName lsidName="E. florezi" pageId="0" pageNumber="317" rank="species" species="florezi">
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<emphasis italics="true" pageId="0" pageNumber="317">E. florezi</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="E. longirostris" pageId="0" pageNumber="317" rank="species" species="longirostris">
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<emphasis italics="true" pageId="0" pageNumber="317">E. longirostris</emphasis>
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</taxonomicName>
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) (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eutrecha Maury, 1982, schematic representation of cheliceral morphology, retrolateral aspect. A Eutrecha longirostris Maury, 1982, holotype ♂ (MAGS 167). B Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia. C Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008). Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MRLC, movable finger, retrolateral carina; MRVC, movable finger, retroventral longitudinal carina; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth. Arrowheads in B indicate extremely reduced FSM, FM, and FD teeth." figureDoi="10.3897/asp.81.e95181.figure2" httpUri="https://binary.pensoft.net/fig/833384" pageId="0" pageNumber="317">2A, B</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers (D). Scale bars = 1 mm (A, C), 0.5 mm (B, D)." figureDoi="10.3897/asp.81.e95181.figure8" httpUri="https://binary.pensoft.net/fig/833390" pageId="0" pageNumber="317">8B</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="317">ii)</emphasis>
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Cheliceral fixed finger with medial notch (MN) between FM and FSM teeth, when these teeth are present (even if greatly reduced) (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eutrecha Maury, 1982, schematic representation of cheliceral morphology, retrolateral aspect. A Eutrecha longirostris Maury, 1982, holotype ♂ (MAGS 167). B Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia. C Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008). Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MRLC, movable finger, retrolateral carina; MRVC, movable finger, retroventral longitudinal carina; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth. Arrowheads in B indicate extremely reduced FSM, FM, and FD teeth." figureDoi="10.3897/asp.81.e95181.figure2" httpUri="https://binary.pensoft.net/fig/833384" pageId="0" pageNumber="317">2B, C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers and stridulatory plate (D). Scale bars = 1 mm." figureDoi="10.3897/asp.81.e95181.figure5" httpUri="https://binary.pensoft.net/fig/833387" pageId="0" pageNumber="317">5B</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, fixed finger, ventral aspect (A), schematic representation of fixed and movable fingers dentition, proventral aspect (B). Scale bars = 0.5 mm. Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MPL, movable finger, prolateral tooth; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; PFM, profondal medial tooth; PFP, profondal proximal tooth; PFSM, profondal submedial tooth; PFSP, profondal subproximal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth." figureDoi="10.3897/asp.81.e95181.figure6" httpUri="https://binary.pensoft.net/fig/833388" pageId="0" pageNumber="317">6</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers (D). Scale bars = 1 mm (A, C), 0.5 mm (B, D)." figureDoi="10.3897/asp.81.e95181.figure8" httpUri="https://binary.pensoft.net/fig/833390" pageId="0" pageNumber="317">8B</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="317">iii)</emphasis>
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Fixed finger median apical diastema (FMAD) present between FM and FD teeth, when these are present (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Eutrecha Maury, 1982, schematic representation of cheliceral morphology, retrolateral aspect. A Eutrecha longirostris Maury, 1982, holotype ♂ (MAGS 167). B Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia. C Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008). Abbreviations: FD, fixed finger, distal tooth; FM, fixed finger, medial tooth; FMAD, fixed finger, median apical diastema; FP, fixed finger, proximal tooth; FSM, fixed finger, submedial tooth; FT, fixed finger, terminal tooth; MM, movable finger, medial tooth; MN, movable finger, medial notch; MP, movable finger, proximal tooth; MRLC, movable finger, retrolateral carina; MRVC, movable finger, retroventral longitudinal carina; MSM, movable finger, submedial tooth; MT, movable finger, terminal tooth; RFA, retrofondal apical tooth; RFM, retrofondal medial tooth; RFP, retrofondal proximal tooth; RFSP, retrofondal subproximal tooth. Arrowheads in B indicate extremely reduced FSM, FM, and FD teeth." figureDoi="10.3897/asp.81.e95181.figure2" httpUri="https://binary.pensoft.net/fig/833384" pageId="0" pageNumber="317">2B, C</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers and stridulatory plate (D). Scale bars = 1 mm." figureDoi="10.3897/asp.81.e95181.figure5" httpUri="https://binary.pensoft.net/fig/833387" pageId="0" pageNumber="317">5B</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers (D). Scale bars = 1 mm (A, C), 0.5 mm (B, D)." figureDoi="10.3897/asp.81.e95181.figure8" httpUri="https://binary.pensoft.net/fig/833390" pageId="0" pageNumber="317">8B</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="317">iv)</emphasis>
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Flagellum without prolateral plumose setiform organ (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers and stridulatory plate (D). Scale bars = 1 mm." figureDoi="10.3897/asp.81.e95181.figure5" httpUri="https://binary.pensoft.net/fig/833387" pageId="0" pageNumber="317">5D</figureCitation>
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,
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, dextral chelicera, retrolateral aspect (A) and close-up of fingers (B), prolateral aspect (C) and close-up of fingers (D). Scale bars = 1 mm (A, C), 0.5 mm (B, D)." figureDoi="10.3897/asp.81.e95181.figure8" httpUri="https://binary.pensoft.net/fig/833390" pageId="0" pageNumber="317">8D</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="317">v)</emphasis>
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Ctenidia, in the form of short
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<normalizedToken originalValue="“fleshy”">"fleshy"</normalizedToken>
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setae, present only on 1st post-genital sternite (spiracular sternite I) (Figs
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Eutrecha belenensis sp. nov., holotype ♂ (ICN Aso 008), habitus, dorsal aspect (A), sinistral pedipalp basitarsus and telotarsus, prolateral (B) and ventral (C) aspects, anterior opisthosomal sternites, with spiracular sternites centered (D). Scale bars = 2 mm (A, D), 1 mm (B, C). Arrows: spiniform setae (in B and C); ctenidia on 1 st post-genital sternite (in D)." figureDoi="10.3897/asp.81.e95181.figure4" httpUri="https://binary.pensoft.net/fig/833386" pageId="0" pageNumber="317">4D</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Eutrecha florezi Villareal-Blanco, Armas and Martinez, 2017, ♂ (IAvH I 472), Santuario de Flora y Fauna Los Colorados, Bolivar Department, Colombia, propeltidium (A), anterior opisthosomal sternites with spiracular sternites centered (B) [arrows indicate paramedian areas with ctenidia on 1 st post-genital sternite], dextral chelicera, stridulatory plate, prolateral aspect (C) [note the modified stridulatory apparatus]. Scale bars = 1 mm (A, C), 0.5 mm (B)." figureDoi="10.3897/asp.81.e95181.figure7" httpUri="https://binary.pensoft.net/fig/833389" pageId="0" pageNumber="317">7B</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="317" type="affinities">
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<paragraph pageId="0" pageNumber="317">Affinities.</paragraph>
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<paragraph pageId="0" pageNumber="317">
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In having the same or similar pattern of spiniform setae on the basitarsi and telotarsi of the walking legs and the same segmentation pattern of the leg telotarsi,
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<taxonomicName authorityName="Maury" authorityYear="1982" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha" order="Pararthropoda" pageId="0" pageNumber="317" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="317">Eutrecha</emphasis>
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</taxonomicName>
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most closely resembles
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<taxonomicName authorityName="Maury" authorityYear="1982" class="Arachnida" family="Ammotrechidae" genus="Xenotrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Xenotrecha" order="Solifugae" pageId="0" pageNumber="317" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="317">Xenotrecha</emphasis>
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</taxonomicName>
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than it does resemble
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<taxonomicName authorityName="Banks" authorityYear="1900" class="Arachnida" family="Ammotrechidae" genus="Ammotrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Ammotrecha" order="Solifugae" pageId="0" pageNumber="317" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="317">Ammotrecha</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="317" type="species composition">
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<paragraph pageId="0" pageNumber="317">Species composition.</paragraph>
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<paragraph pageId="0" pageNumber="317">
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<taxonomicName authorityName="Botero-Trujillo & Martínez & Iuri & Ojanguren-Affilastro & Carvalho" authorityYear="2023" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha belenensis" order="Pararthropoda" pageId="0" pageNumber="317" phylum="Arthropoda" rank="species" species="belenensis">
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<emphasis italics="true" pageId="0" pageNumber="317">Eutrecha belenensis</emphasis>
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</taxonomicName>
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<emphasis bold="true" pageId="0" pageNumber="317">sp. nov.</emphasis>
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;
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<taxonomicName authorityName="Villareal-Blanco, Armas and Martinez" authorityYear="2017" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha florezi" order="Pararthropoda" pageId="0" pageNumber="317" phylum="Arthropoda" rank="species" species="florezi">
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<emphasis italics="true" pageId="0" pageNumber="317">Eutrecha florezi</emphasis>
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</taxonomicName>
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Villareal-Blanco, Armas and
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<normalizedToken originalValue="Martínez">Martinez</normalizedToken>
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2017;
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<taxonomicName authorityName="Maury" authorityYear="1982" class="Arachnida" family="Ammotrechidae" genus="Eutrecha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eutrecha longirostris" order="Pararthropoda" pageId="0" pageNumber="317" phylum="Arthropoda" rank="species" species="longirostris">
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<emphasis italics="true" pageId="0" pageNumber="317">Eutrecha longirostris</emphasis>
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</taxonomicName>
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Maury, 1982.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |