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<document id="78641C430716158D51E354F2BD45BF2C" ID-CLB-Dataset="218234" ID-DOI="10.1206/0003-0090.458.1.1" ID-GBIF-Dataset="9aaee2d0-a89e-47d4-8578-cdd525f7ebfb" ID-ISSN="0003-0090" ID-Zenodo-Dep="7400026" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1670242945939" checkinUser="felipe" docAuthor="Jones, Lance E. &amp; Grimaldi, David A." docDate="2022" docId="3C6787EDFFB9FFE2633CFADAFC21FD44" docLanguage="en" docName="BAmMusNatHist.2022.458.1-177.pdf" docOrigin="Bulletin of the American Museum of Natural History 2022 (458)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-458/issue-1/0003-0090.458.1.1/Revision-of-the-Nearctic-Species-of-the-Genus-Amiota-Loew/10.1206/0003-0090.458.1.1.full" docStyle="DocumentStyle:915933466F796C9C739DF4DB6B8DCFA6.9:BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleId="915933466F796C9C739DF4DB6B8DCFA6" docStyleName="BulAmeMusNatHis.2011-.journal_article.1cover.type1" docStyleVersion="9" docTitle="Amiota onyx Jones &amp; Grimaldi 2022, sp. nov." docType="treatment" docVersion="9" lastPageNumber="78" masterDocId="C05EFF95FFF4FFAC6000FFF0FFEDFF88" masterDocTitle="Revision Of The Nearctic Species Of The Genus Amiota Loew (Diptera: Drosophilidae)" masterLastPageNumber="181" masterPageNumber="1" pageNumber="77" updateTime="1699413003214" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="859B4528C17CF0ABA4FEA40DF98095E6">Revision Of The Nearctic Species Of The Genus Amiota Loew (Diptera: Drosophilidae)</mods:title>
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<mods:namePart id="A9F55A7D8209F7EAAD9BD27457E005EC">Jones, Lance E.</mods:namePart>
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<mods:namePart id="3FAECBB8E4DAE137ED7024D5D8C35255">Grimaldi, David A.</mods:namePart>
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<treatment id="3C6787EDFFB9FFE2633CFADAFC21FD44" ID-DOI="http://doi.org/10.5281/zenodo.7506537" ID-GBIF-Taxon="208565081" ID-Zenodo-Dep="7506537" LSID="urn:lsid:plazi:treatment:3C6787EDFFB9FFE2633CFADAFC21FD44" httpUri="http://treatment.plazi.org/id/3C6787EDFFB9FFE2633CFADAFC21FD44" lastPageId="78" lastPageNumber="78" pageId="77" pageNumber="77">
<subSubSection id="FCD46570FFB9FFE1633CFADAFBC1FAC9" box="[828,1068,1322,1346]" pageId="77" pageNumber="77" type="nomenclature">
<paragraph id="B47136FBFFB9FFE1633CFADAFBC1FAC9" blockId="77.[828,1068,1322,1346]" box="[828,1068,1322,1346]" pageId="77" pageNumber="77">
<heading id="EF398197FFB9FFE1633CFADAFBC1FAC9" box="[828,1068,1322,1346]" centered="true" fontSize="9" level="2" pageId="77" pageNumber="77" reason="2">
<emphasis id="86BAEAE9FFB9FFE1633CFADAFC3EFACA" bold="true" box="[828,979,1322,1346]" pageId="77" pageNumber="77">
<taxonomicName id="73CE4D78FFB9FFE1633CFADAFC21FAC9" ID-CoL="BDJJ9" authority="Jones &amp; Grimaldi, 2022" authorityName="Jones &amp; Grimaldi" authorityYear="2022" box="[828,972,1322,1346]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="77" pageNumber="77" phylum="Arthropoda" rank="species" species="onyx" status="sp. nov.">
<emphasis id="86BAEAE9FFB9FFE1633CFADAFC21FAC9" bold="true" box="[828,972,1322,1346]" italics="true" pageId="77" pageNumber="77">Amiota onyx</emphasis>
</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="9D895792FFB9FFE163DAFADAFBC1FAC9" box="[986,1068,1322,1345]" pageId="77" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB9FFE162C4FAABFB48FAFC" box="[708,1189,1371,1396]" pageId="77" pageNumber="77" type="description">
<paragraph id="B47136FBFFB9FFE162C4FAABFB48FAFC" blockId="77.[708,1189,1371,1396]" box="[708,1189,1371,1396]" pageId="77" pageNumber="77">
<heading id="EF398197FFB9FFE162C4FAABFB48FAFC" box="[708,1189,1371,1396]" centered="true" fontSize="9" level="2" pageId="77" pageNumber="77" reason="2">
<figureCitation id="2CF52A7EFFB9FFE162C4FAABFC86FAFB" box="[708,875,1371,1395]" captionStart="FIG" captionStartId="83.[108,150,1205,1226]" captionTargetBox="[108,1212,224,1177]" captionTargetId="figure-233@83.[108,1212,224,1177]" captionTargetPageId="83" captionText="FIG. 35. Heads and lateral views, A. avipes species group. AB. A. minor (Malloch) (Am 651). Lack of white spots is natural. CD. A. onyx, sp. nov. (Am 1450)." figureDoi="http://doi.org/10.5281/zenodo.7400096" httpUri="https://zenodo.org/record/7400096/files/figure.png" pageId="77" pageNumber="77">Figures 35CD</figureCitation>
,
<figureCitation id="2CF52A7EFFB9FFE16376FAACFC2FFAFC" box="[886,962,1372,1396]" captionStart="FIG" captionStartId="86.[109,151,1555,1576]" captionTargetBox="[117,1208,224,1529]" captionTargetId="figure-52@86.[107,1211,224,1529]" captionTargetPageId="86" captionText="FIG. 37. Aristae, A. avipes species group. A. A. avipes. B. A. biacuminis. C. A. forceps. D, F. A. minor (Malloch). E, G. A. onyx. HI. A. pseudominor. Not to the same scale." figureDoi="http://doi.org/10.5281/zenodo.7400100" httpUri="https://zenodo.org/record/7400100/files/figure.png" pageId="77" pageNumber="77">37E, G</figureCitation>
,
<figureCitation id="2CF52A7EFFB9FFE163CEFAACFC10FAFC" box="[974,1021,1372,1396]" captionStart="FIG" captionStartId="87.[107,149,1612,1633]" captionTargetBox="[183,1134,230,1569]" captionTargetId="figure-68@87.[183,1138,224,1582]" captionTargetPageId="87" captionText="FIG. 38. Male terminalia, A. onyx. A, CD, F, H. A. Posterior view. C. Aedeagal apodeme. D. Lateral view. F. Outer paraphysis, lateral. H. Surstylus. (Am 1505, holotype). Male terminalia, A. avipes. B, E, G, I. B. Posterior view of genitalia. E. Lateral view. G. Outer paraphysis, lateral. I. Surstyli. (Am 1453, holotype)." figureDoi="http://doi.org/10.5281/zenodo.7400102" httpUri="https://zenodo.org/record/7400102/files/figure.png" pageId="77" pageNumber="77">38A</figureCitation>
, C-D, F, H 88B
</heading>
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFB9FFE162CCFA7DFCB1F926" pageId="77" pageNumber="77" type="diagnosis">
<paragraph id="B47136FBFFB9FFE162CCFA7DFCB1F926" blockId="77.[684,1212,1421,1710]" pageId="77" pageNumber="77">
DIAGNOSIS: Small to medium-sized fly (ThL
<quantity id="73369B1EFFB9FFE162ACFA5EFCBEFA4E" box="[684,851,1454,1478]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.22" metricValueMin="0.98" pageId="77" pageNumber="77" unit="mm" value="1.1" valueMax="1.22" valueMin="0.98">0.981.22 mm</quantity>
), medium brown, darker above grading lighter ventrally, legs yellow; white markings characteristic of genus present, spot under wing base slightly faded; outer paraphyses heavily sclerotized, symmetrical, short, forming stout, strongly curved claw, winglike basal projection, pointing laterally; inner paraphysis lost.
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFBAFFE2608CFF12FE22F9EF" pageId="78" pageNumber="78" type="description">
<paragraph id="B47136FBFFBAFFE2608CFF12FE22F9EF" blockId="78.[108,637,226,1738]" pageId="78" pageNumber="78">
DESCRIPTION: Small to medium-sized fly (ThL
<quantity id="73369B1EFFBAFFE260A9FEF3FEA4FE93" box="[169,329,259,283]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.22" metricValueMin="0.98" pageId="78" pageNumber="78" unit="mm" value="1.1" valueMax="1.22" valueMin="0.98">0.981.22 mm</quantity>
), medium brown body, fading ventrally; legs yellow. Characteristic markings of genus present, marking under wing base rather faint. Frons medium brown, darkening to black ventrally. Facial marking of moderate width (FML/FMW 0.260.29). Palp brown. Cheek yellowish. Tergites 1 and 2 lightly colored. Arista: Medium, plumose; longest branch D2; A.R. 0.32; 2 longer and 3 shorter dorsal, 3 short ventral branches; branches D35 pointed mediad, V13 pointed laterad; arista trunk with medium-length microtrichia nearly to apex. Male genitalia: Epandrium dorsally complete but with ventral margin in this area graded into membrane below; row of ~6 setae leading near posterior margin. Cercus long, pendulous, dorsally grading into membrane and epandrium; lateral margins of cercus discrete; cerci large, occupying virtually all space surrounded by epandrium. Surstylus with 11 prensisetae, apices blunt, evenly and densely spaced, comblike; middle region and ventral margin with setulae. Subepandrial sclerite pyramidal in posterior view, not projecting beyond epandrium. Outer paraphysis heavily sclerotized, short, forming stout hooked claw; base with small row of minute sensilla; distinctive low carina basal to sensilla; basal winglike projection, pointing laterally, the distal end with faint striations. Inner paraphysis lost. Aedeagal apodeme with all 4 margins constricted, bent nearly 90° in lateral view; basal margin slightly wider than distal end; distal end hardly expanded; marginal constrictions very shallow. Hypandrium thick, roughly square in terminal view; lateral arms very thick, bulging, with a prominent ostium. Ejaculatory apodeme 0.60× length of epandrium; stalk slightly sinuous. Head and thorax measurements: (n = 2; Am 1450, 1505) FL/FW 0.55 (0.550.56), EL/EW 1.31 (1.271.36), EL/CW 11.72 (11.2012.25), FML/FMW 0.27 (0.260.29),
<collectingRegion id="760AF819FFBAFFE261B0F9DEFE3DF9CE" box="[432,464,1582,1606]" country="Brazil" name="Para" pageId="78" pageNumber="78">PR</collectingRegion>
/RR 0.49 (0.44 0.55), ThL 1.10 (0.981.22 mm).
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFBAFFE2608CF980FD1EFE4C" pageId="78" pageNumber="78" type="materials_examined">
<paragraph id="B47136FBFFBAFFE2608CF980FB51FEB4" blockId="78.[108,637,226,1738]" lastBlockId="78.[684,1212,226,717]" pageId="78" pageNumber="78">
<collectionCode id="D2DFAE3EFFBAFFE2608CF980FEAEF900" box="[140,323,1648,1672]" pageId="78" pageNumber="78">TYPE MATERIAL</collectionCode>
:
<materialsCitation id="04A63CA6FFBAFFE2614FF980FB51FEB4" ID-GBIF-Occurrence="4010926508" collectingDate="1969-06-04" collectionCode="CNC" collectorName="B. V. Peterson" country="Mexico" determinerName="D. Grimaldi" latitude="15.537566" location="Huixtla" longLatPrecision="1" longitude="-92.47319" pageId="78" pageNumber="78" specimenCount="1" stateProvince="Chiapas" typeStatus="holotype">
<emphasis id="86BAEAE9FFBAFFE2614FF980FE2DF900" bold="true" box="[335,448,1648,1672]" pageId="78" pageNumber="78">
<typeStatus id="6B758859FFBAFFE2614FF980FE56F900" box="[335,443,1648,1672]" pageId="78" pageNumber="78" type="holotype">Holotype</typeStatus>
:
</emphasis>
male:
<collectingCountry id="CCD9766BFFBAFFE2620EF980FD95F900" box="[526,632,1648,1672]" name="Mexico" pageId="78" pageNumber="78">MEXICO</collectingCountry>
, Chis. [
<collectingRegion id="760AF819FFBAFFE260B0F961FEE0F921" box="[176,269,1681,1705]" country="Mexico" name="Chiapas" pageId="78" pageNumber="78">Chiapas</collectingRegion>
]
<quantity id="73369B1EFFBAFFE26118F961FE69F921" box="[280,388,1681,1705]" metricMagnitude="4" metricUnit="m" metricValue="3.621024" metricValueMax="4.02336" metricValueMin="3.218688" pageId="78" pageNumber="78" unit="mi" value="22.5" valueMax="25.0" valueMin="20.0">2025 mi.</quantity>
N
<location id="B1116020FFBAFFE261A3F961FE1BF921" LSID="urn:lsid:plazi:treatment:3C6787EDFFB9FFE2633CFADAFC21FD44:B1116020FFBAFFE261A3F961FE1BF921" box="[419,502,1681,1705]" country="Mexico" latitude="15.537566" longLatPrecision="1" longitude="-92.47319" name="Huixtla" pageId="78" pageNumber="78" stateProvince="Chiapas">Huixtla</location>
, [
<geoCoordinate id="D1FA503CFFBAFFE26205F961FD95F921" box="[517,632,1681,1705]" degrees="15.537566" direction="north" orientation="latitude" pageId="78" pageNumber="78" precision="1" value="15.537566">15.537566</geoCoordinate>
,
<geoCoordinate id="D1FA503CFFBAFFE2606CF942FF00F942" box="[108,237,1714,1738]" degrees="92.473191" direction="west" orientation="longitude" pageId="78" pageNumber="78" precision="1" value="-92.47319">-92.473191</geoCoordinate>
], 3000,
<date id="C070103BFFBAFFE26159F942FE25F942" box="[345,456,1714,1738]" pageId="78" pageNumber="78" value="1969-06-04">
<collectingDate id="D034E9D3FFBAFFE26159F942FE25F942" box="[345,456,1714,1738]" pageId="78" pageNumber="78" value="1969-06-04">4.VI.1969</collectingDate>
</date>
, B.
<collectionCode id="D2DFAE3EFFBAFFE261EFF942FDEEF941" box="[495,515,1714,1737]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="78" pageNumber="78" type="Museum">V</collectionCode>
.
<collectorName id="193B532DFFBAFFE26215F942FD95F941" box="[533,632,1714,1737]" pageId="78" pageNumber="78">Peterson</collectorName>
, “
<taxonomicName id="73CE4D78FFBAFFE262B5FF12FC9AFF72" box="[693,887,226,250]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="subGenus" subGenus="Amiota">Amiota (Amiota)</taxonomicName>
” det.
<determinerName id="41ADDA3FFFBAFFE263BDFF12FBA8FF72" box="[957,1093,226,250]" pageId="78" pageNumber="78">D. Grimaldi</determinerName>
1991,
<collectorName id="193B532DFFBAFFE26493FF12FB51FF72" box="[1171,1212,226,250]" pageId="78" pageNumber="78">Am</collectorName>
1505, [glued directly to pin, dissected].
<collectorName id="193B532DFFBAFFE2646CFEF4FD38FEB4" pageId="78" pageNumber="78">Deposited</collectorName>
in the
<collectorName id="193B532DFFBAFFE26320FED4FB89FEB4" box="[800,1124,292,316]" pageId="78" pageNumber="78">Canadian National Collection</collectorName>
(
<collectionCode id="D2DFAE3EFFBAFFE26472FED4FB5DFEB4" box="[1138,1200,292,316]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="78" pageNumber="78">CNC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="B47136FBFFBAFFE262DCFEB6FD1EFE4C" blockId="78.[684,1212,226,717]" pageId="78" pageNumber="78">
<collectionCode id="D2DFAE3EFFBAFFE262DCFEB6FBA5FED6" box="[732,1096,326,350]" pageId="78" pageNumber="78">OTHER MATERIAL EXAMINED</collectionCode>
:
<emphasis id="86BAEAE9FFBAFFE26460FEB6FCF5FE08" bold="true" pageId="78" pageNumber="78">
<collectingCountry id="CCD9766BFFBAFFE26460FEB6FB5AFED6" box="[1120,1207,326,350]" name="Mexico" pageId="78" pageNumber="78">Mexico</collectingCountry>
:
<collectingRegion id="760AF819FFBAFFE262ACFE98FCFEFE08" box="[684,787,360,384]" country="Mexico" name="Durango" pageId="78" pageNumber="78">Durango</collectingRegion>
:
</emphasis>
<quantity id="73369B1EFFBAFFE2631EFE98FC8CFE08" box="[798,865,360,384]" metricMagnitude="4" metricUnit="m" metricValue="2.2530816" pageId="78" pageNumber="78" unit="mi" value="14.0">14 mi.</quantity>
SW. El Salto, 8000,
<date id="C070103BFFBAFFE2643DFE98FB55FE08" box="[1085,1208,360,384]" pageId="78" pageNumber="78" value="1964-06-26">1964-06-26</date>
, leg. J.F. McAlpine, attracted to man (Am 1450*,
<collectionCode id="D2DFAE3EFFBAFFE262ACFE5CFD0AFE4C" box="[684,743,428,452]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="78" pageNumber="78">CNC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFBAFFE262CCFE3DFC90FDAF" pageId="78" pageNumber="78" type="etymology">
<paragraph id="B47136FBFFBAFFE262CCFE3DFC90FDAF" blockId="78.[684,1212,226,717]" pageId="78" pageNumber="78">ETYMOLOGY: Greek for “claw,” in reference to the paraphyses that in lateral view resemble a dark, curving claw.</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFBAFFE262CCFDC1FBB9FDE2" pageId="78" pageNumber="78" type="distribution">
<paragraph id="B47136FBFFBAFFE262CCFDC1FBB9FDE2" blockId="78.[684,1212,226,717]" pageId="78" pageNumber="78">
DISTRIBUTION:
<taxonomicName id="73CE4D78FFBAFFE2637DFDC1FBE6FDC0" authorityName="Jones &amp; Grimaldi" authorityYear="2022" box="[893,1035,561,584]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="species" species="onyx">
<emphasis id="86BAEAE9FFBAFFE2637DFDC1FBE6FDC0" box="[893,1035,561,584]" italics="true" pageId="78" pageNumber="78">Amiota onyx</emphasis>
</taxonomicName>
is known from
<collectingRegion id="760AF819FFBAFFE262ACFDA2FCE0FDE2" box="[684,781,594,618]" country="Mexico" name="Durango" pageId="78" pageNumber="78">Durango</collectingRegion>
and
<collectingRegion id="760AF819FFBAFFE26345FDA2FC35FDE2" box="[837,984,594,618]" country="Mexico" name="Chiapas" pageId="78" pageNumber="78">Chiapas State</collectingRegion>
in
<collectingRegion id="760AF819FFBAFFE263FDFDA2FBBDFDE2" box="[1021,1104,594,618]" country="Mexico" name="Mexico" pageId="78" pageNumber="78">Mexico</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="FCD46570FFBAFFE262CCFD83FC21FD44" pageId="78" pageNumber="78" type="discussion">
<paragraph id="B47136FBFFBAFFE262CCFD83FC21FD44" blockId="78.[684,1212,226,717]" pageId="78" pageNumber="78">
COMMENTS: This species exhibits the characteristic behavior of attraction to the eyes and face common to many
<taxonomicName id="73CE4D78FFBAFFE26377FD45FC2BFD44" box="[887,966,693,716]" class="Insecta" family="Drosophilidae" genus="Amiota" kingdom="Animalia" order="Diptera" pageId="78" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="86BAEAE9FFBAFFE26377FD45FC2BFD44" box="[887,966,693,716]" italics="true" pageId="78" pageNumber="78">Amiota</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>