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<document id="C420FA0CA234573306EA7DE51777F05B" ID-DOI="10.11646/zootaxa.5351.5.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8392232" ID-ZooBank="1BD2A231-4D66-47A7-8EE6-4A1C241C4190" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695992926648" checkinUser="plazi" docAuthor="Zhao, Fangyu, Yang, Lan, Li, Shuqiang, Zheng, Guo &amp; Yao, Zhiyuan" docDate="2023" docId="03CA8799E728FFC112C1D475DD78FE0A" docLanguage="en" docName="zootaxa.5351.5.3.pdf" docOrigin="Zootaxa 5351 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.5.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana chuandiani Li, Zheng &amp; Yao 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="549" masterDocId="FFF3FFE1E72BFFC71256D03EDE72FF83" masterDocTitle="A further study on the Belisana spiders (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China" masterLastPageNumber="558" masterPageNumber="543" pageNumber="546" updateTime="1695993610286" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="2ABC32C5AC7C0BA802DD08B9AE70BA7C">A further study on the Belisana spiders (Araneae: Pholcidae) from Xishuangbanna, Yunnan, China</mods:title>
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<mods:namePart id="73389E06E9DD2A69DA70D520436106B7">Zhao, Fangyu</mods:namePart>
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<mods:namePart id="89D184407DF7B050A8C931974467D6C7">Yang, Lan</mods:namePart>
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<mods:namePart id="701467B6227A8AC8EA619EF3C1282131">Li, Shuqiang</mods:namePart>
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<mods:affiliation id="AD5193325F6C3B38703CCA7E4688A332">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
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<mods:namePart id="38D63F601B45274180AB093FE50E0C68">Zheng, Guo</mods:namePart>
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<mods:affiliation id="80101E78751DC509E664EBE9824F9F09">zhengguo @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 9735 - 9700</mods:affiliation>
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<mods:namePart id="9786EF9307B0352523512E54605EFB01">Yao, Zhiyuan</mods:namePart>
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<mods:date id="0AF722C45DE84F37081A5EC13A4423F3">2023</mods:date>
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<paragraph id="8BDC368FE728FFC412C1D475DCBDFBE5" blockId="3.[151,719,1099,1162]" box="[151,719,1099,1126]" pageId="3" pageNumber="546">
<heading id="D09481E3E728FFC412C1D475DCBDFBE5" bold="true" box="[151,719,1099,1126]" fontSize="11" level="1" pageId="3" pageNumber="546" reason="1">
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<taxonomicName id="4C634D0CE728FFC412C1D475DC18FBE5" authority="Li, Zheng &amp; Yao" authorityName="Li, Zheng &amp; Yao" authorityYear="2023" box="[151,618,1099,1126]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="546" phylum="Arthropoda" rank="species" species="chuandiani" status="sp. nov.">
<emphasis id="B917EA9DE728FFC412C1D475DFE1FBE5" bold="true" box="[151,403,1099,1126]" italics="true" pageId="3" pageNumber="546">Belisana chuandiani</emphasis>
Li, Zheng &amp; Yao
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<taxonomicNameLabel id="A22457E6E728FFC41027D472DCBDFBE5" box="[625,719,1100,1126]" pageId="3" pageNumber="546" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="C3796504E728FFC412C1D451DFDBFB09" box="[151,425,1135,1162]" pageId="3" pageNumber="546" type="description">
<paragraph id="8BDC368FE728FFC412C1D451DFDBFB09" blockId="3.[151,719,1099,1162]" box="[151,425,1135,1162]" pageId="3" pageNumber="546">
<figureCitation id="13582A0AE728FFC412C1D451DE89FB09" box="[151,251,1135,1162]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="4.[151,250,1856,1880]" captionStartId-1="5.[151,250,1856,1880]" captionTargetBox-0="[233,1355,181,1832]" captionTargetBox-1="[233,1355,181,1832]" captionTargetId-0="figure-22@4.[233,1355,181,1832]" captionTargetId-1="figure-18@5.[233,1355,181,1832]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." captionText-1="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.8392238" figureDoi-1="http://doi.org/10.5281/zenodo.8392242" httpUri-0="https://zenodo.org/record/8392238/files/figure.png" httpUri-1="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Figs 23</figureCitation>
,
<figureCitation id="13582A0AE728FFC41350D44EDF3EFB09" box="[262,332,1136,1162]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9AB</figureCitation>
,
<figureCitation id="13582A0AE728FFC41301D44EDFDBFB09" box="[343,425,1136,1162]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[194,1392,184,1916]" captionTargetId="figure-22@14.[191,1395,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Female genitalia in ventral and dorsal views. AB. Belisana chuandiani sp. nov.; CD. Belisana fengzheni sp. nov.; EF. Belisana zhengi Yao, Pham &amp; Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392270" httpUri="https://zenodo.org/record/8392270/files/figure.png" pageId="3" pageNumber="546">10AB</figureCitation>
</paragraph>
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<emphasis id="B917EA9DE728FFC412C1D486DF37FB52" bold="true" box="[151,325,1207,1234]" pageId="3" pageNumber="546">Type material.</emphasis>
<materialsCitation id="3B0B3CD2E728FFC4131DD486DE87FA9A" collectingDate="2012-08-22" collectorName="Guo Zheng &amp; Xue Li &amp; Wenyue Zhu" country="China" county="Xishuangbanna" elevation="736" latitude="22.13075" location="Male" longLatPrecision="1" longitude="100.66248" municipality="Nabanhe Nature Reserve" pageId="3" pageNumber="546" specimenCount="1" stateProvince="Yunnan" typeStatus="holotype">
<emphasis id="B917EA9DE728FFC4131DD486DFB3FB51" bold="true" box="[331,449,1208,1234]" pageId="3" pageNumber="546">
<typeStatus id="54D8882DE728FFC4131DD486DFCEFB51" box="[331,444,1208,1234]" pageId="3" pageNumber="546">Holotype</typeStatus>
:
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<location id="8EBC6054E728FFC4139ED489DC70FB52" LSID="urn:lsid:plazi:treatment:03CA8799E728FFC112C1D475DD78FE0A:8EBC6054E728FFC4139ED489DC70FB52" box="[456,514,1207,1233]" country="China" county="Xishuangbanna" latitude="22.13075" longLatPrecision="1" longitude="100.66248" municipality="Nabanhe Nature Reserve" name="Male" pageId="3" pageNumber="546" stateProvince="Yunnan">Male</location>
(SYNU-Ar00308), near
<location id="8EBC6054E728FFC4114FD489DDCBFB52" LSID="urn:lsid:plazi:treatment:03CA8799E728FFC112C1D475DD78FE0A:8EBC6054E728FFC4114FD489DDCBFB52" box="[793,953,1207,1233]" country="China" county="Xishuangbanna" latitude="22.13075" longLatPrecision="1" longitude="100.66248" municipality="Nabanhe Nature Reserve" name="Mandian Falls" pageId="3" pageNumber="546" stateProvince="Yunnan">Mandian Falls</location>
(
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,
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,
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elevation
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),
<collectingMunicipality id="6BB8ACF5E728FFC41292D4E5DFADFB75" box="[196,479,1243,1270]" pageId="3" pageNumber="546">Nabanhe Nature Reserve</collectingMunicipality>
,
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<collectingRegion id="49A7F86DE728FFC410F0D4E2DD76FB75" box="[678,772,1244,1270]" country="China" name="Yunnan" pageId="3" pageNumber="546">Yunnan</collectingRegion>
</emphasis>
,
<emphasis id="B917EA9DE728FFC41144D4E5DD29FB76" bold="true" box="[786,859,1243,1269]" pageId="3" pageNumber="546">
<collectingCountry id="F374761FE728FFC41144D4E5DD29FB76" box="[786,859,1243,1269]" name="China" pageId="3" pageNumber="546">China</collectingCountry>
</emphasis>
,
<date id="FFDD104FE728FFC4113FD4E2DA6CFB75" box="[873,1054,1244,1270]" pageId="3" pageNumber="546" value="2012-08-22">
<collectingDate id="EF99E9A7E728FFC4113FD4E2DA6CFB75" box="[873,1054,1244,1270]" pageId="3" pageNumber="546" value="2012-08-22">22 August 2012</collectingDate>
</date>
,
<collectorName id="26965359E728FFC4167FD4E2DAD8FB76" box="[1065,1194,1243,1270]" pageId="3" pageNumber="546">Guo Zheng</collectorName>
,
<collectorName id="26965359E728FFC416E0D4E2DB74FB76" box="[1206,1286,1243,1270]" pageId="3" pageNumber="546">Xue Li</collectorName>
, and
<collectorName id="26965359E728FFC41717D4E2DEB6FA9A" pageId="3" pageNumber="546">Wenyue Zhu</collectorName>
leg.
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<emphasis id="B917EA9DE728FFC412ADD53EDF00FA99" bold="true" box="[251,370,1280,1306]" pageId="3" pageNumber="546">
<typeStatus id="54D8882DE728FFC412ADD53EDF1EFA99" box="[251,364,1280,1306]" pageId="3" pageNumber="546">Paratype</typeStatus>
:
</emphasis>
<specimenCount id="9D65FD06E728FFC4132FD53EDFA8FA9A" box="[377,474,1279,1306]" pageId="3" pageNumber="546" type="female">1 female</specimenCount>
(SYNU-Ar00309), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3796504E728FFC41291D51ADFBEFAE1" pageId="3" pageNumber="546" type="etymology">
<paragraph id="8BDC368FE728FFC41291D51ADFBEFAE1" blockId="3.[151,1437,1207,1990]" pageId="3" pageNumber="546">
<emphasis id="B917EA9DE728FFC41291D51ADF3CFABD" bold="true" box="[199,334,1316,1342]" pageId="3" pageNumber="546">Etymology.</emphasis>
The specific name is a patronym in honor of the late Chinese arachnologist Chuandian Zhu; noun (name) in the genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3796504E728FFC41291D555DBFAF995" pageId="3" pageNumber="546" type="diagnosis">
<paragraph id="8BDC368FE728FFC41291D555DBFAF995" blockId="3.[151,1437,1207,1990]" pageId="3" pageNumber="546">
<emphasis id="B917EA9DE728FFC41291D555DF30FA06" bold="true" box="[199,322,1387,1413]" pageId="3" pageNumber="546">Diagnosis.</emphasis>
The new species resembles
<taxonomicName id="4C634D0CE728FFC410D1D552DDC2FA05" authority="Yao &amp; Li, 2018" authorityName="Yao &amp; Li" authorityYear="2018" box="[647,944,1387,1414]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="546" phylum="Arthropoda" rank="species" species="gupian">
<emphasis id="B917EA9DE728FFC410D1D552DC84FA06" box="[647,758,1388,1413]" italics="true" pageId="3" pageNumber="546">B. gupian</emphasis>
<bibRefCitation id="EFF24B7EE728FFC410ABD552DDC2FA05" author="Yao, Z. &amp; Zhu, K. &amp; Du, Z. &amp; Li, S." box="[765,944,1387,1414]" pageId="3" pageNumber="546" pagination="243 - 262" refId="ref10105" refString="Yao, Z., Zhu, K., Du, Z. &amp; Li, S. (2018) The Belisana spiders (Araneae: Pholcidae) from Xishuangbanna Tropical Botanical Garden, Yunnan, China. Zootaxa, 4425 (2), 243 - 262. https: // doi. org / 10.11646 / zootaxa. 4425.2.3" type="journal article" year="2018">Yao &amp; Li, 2018</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFF24B7EE728FFC41196D552DADCFA05" author="Yao, Z. &amp; Zhu, K. &amp; Du, Z. &amp; Li, S." box="[960,1198,1387,1414]" pageId="3" pageNumber="546" pagination="243 - 262" refId="ref10105" refString="Yao, Z., Zhu, K., Du, Z. &amp; Li, S. (2018) The Belisana spiders (Araneae: Pholcidae) from Xishuangbanna Tropical Botanical Garden, Yunnan, China. Zootaxa, 4425 (2), 243 - 262. https: // doi. org / 10.11646 / zootaxa. 4425.2.3" type="journal article" year="2018">
Yao
<emphasis id="B917EA9DE728FFC411A1D552DA43FA06" box="[1015,1073,1387,1413]" italics="true" pageId="3" pageNumber="546">et al.</emphasis>
2018: 247
</bibRefCitation>
, figs 34, 11B, 12B) having similar bulbal apophyses (
<figureCitation id="13582A0AE728FFC41046D5B1DC15FA29" box="[528,615,1423,1450]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Fig. 3C</figureCitation>
), male chelicerae (
<figureCitation id="13582A0AE728FFC4116CD5B1DDE1FA29" box="[826,915,1423,1450]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Fig. 3D</figureCitation>
), and shape of epigyne (
<figureCitation id="13582A0AE728FFC416F3D5B1DA8CFA29" box="[1189,1278,1423,1450]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Fig. 3A</figureCitation>
), but it can be distinguished by prolatero-subdistal sclerite of procursus nearly angular (arrow
<quantity id="4C9B9B6AE728FFC41641D58ADA30FA4E" box="[1047,1090,1459,1486]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="546" unit="in" value="1.0">1 in</quantity>
<figureCitation id="13582A0AE728FFC4161FD58DDAD6FA4D" box="[1097,1188,1459,1486]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2C</figureCitation>
,
<figureCitation id="13582A0AE728FFC416E6D58ADAA7FA4D" box="[1200,1237,1460,1486]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9A</figureCitation>
; vs. half-round in
<taxonomicName id="4C634D0CE728FFC412C1D5E6DF71FA72" authorityName="Yao &amp; Li" authorityYear="2018" box="[151,259,1496,1521]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="546" phylum="Arthropoda" rank="species" species="gupian">
<emphasis id="B917EA9DE728FFC412C1D5E6DF71FA72" box="[151,259,1496,1521]" italics="true" pageId="3" pageNumber="546">B. gupian</emphasis>
</taxonomicName>
), by procursus straight (
<figureCitation id="13582A0AE728FFC41047D5E9DCFCFA71" box="[529,654,1495,1522]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2AB</figureCitation>
,
<figureCitation id="13582A0AE728FFC410CFD5E6DCADFA71" box="[665,735,1496,1522]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9AB</figureCitation>
; vs.distal part of procursus directed towards antero-ventrally in
<taxonomicName id="4C634D0CE728FFC412C1D5C2DF76F996" authorityName="Yao &amp; Li" authorityYear="2018" box="[151,260,1532,1557]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="546" phylum="Arthropoda" rank="species" species="gupian">
<emphasis id="B917EA9DE728FFC412C1D5C2DF76F996" box="[151,260,1532,1557]" italics="true" pageId="3" pageNumber="546">B. gupian</emphasis>
</taxonomicName>
), and by vulval pore plates nearly elliptic (
<figureCitation id="13582A0AE728FFC410B3D5C5DD33F995" box="[741,833,1531,1558]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Figs 3B</figureCitation>
,
<figureCitation id="13582A0AE728FFC4111BD5C2DD0DF995" box="[845,895,1532,1558]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[194,1392,184,1916]" captionTargetId="figure-22@14.[191,1395,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Female genitalia in ventral and dorsal views. AB. Belisana chuandiani sp. nov.; CD. Belisana fengzheni sp. nov.; EF. Belisana zhengi Yao, Pham &amp; Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392270" httpUri="https://zenodo.org/record/8392270/files/figure.png" pageId="3" pageNumber="546">10B</figureCitation>
; vs. anteriorly strongly widened in
<taxonomicName id="4C634D0CE728FFC4175CD5C2DB0AF996" authorityName="Yao &amp; Li" authorityYear="2018" box="[1290,1400,1532,1557]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="546" phylum="Arthropoda" rank="species" species="gupian">
<emphasis id="B917EA9DE728FFC4175CD5C2DB0AF996" box="[1290,1400,1532,1557]" italics="true" pageId="3" pageNumber="546">B. gupian</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C3796504E728FFC11291D621DAE4FEE6" lastPageId="6" lastPageNumber="549" pageId="3" pageNumber="546" type="description">
<paragraph id="8BDC368FE728FFC41291D621DB3CF846" blockId="3.[151,1437,1207,1990]" pageId="3" pageNumber="546">
<emphasis id="B917EA9DE728FFC41291D621DFE9F9B9" bold="true" box="[199,411,1567,1594]" pageId="3" pageNumber="546">Description. Male</emphasis>
(
<emphasis id="B917EA9DE728FFC413FCD61EDC63F9B9" bold="true" box="[426,529,1568,1594]" pageId="3" pageNumber="546">
<typeStatus id="54D8882DE728FFC413FCD61EDC63F9B9" box="[426,529,1568,1594]" pageId="3" pageNumber="546">holotype</typeStatus>
</emphasis>
, SYNU-Ar00308): Total length 1.47 (1.56 with clypeus), prosoma 0.61 long, 0.55 wide, opisthosoma 0.86 long, 0.75 wide. Leg I: (4.08 + 0.30 + + + ), leg II missing, leg III: 6.34 (1.74 + 0.23 + 1.54 + 2.16 + 0.67), leg IV: (2.49 + 0.23 + 2.28 + + ). Eye interdistances and diameters: PME-PME 0.06, PME 0.06, PME-ALE 0.03, AME absent. Sternum width/length: 0.46/0.43. Habitus as in
<figureCitation id="13582A0AE728FFC416D1D6B5DA8EF925" box="[1159,1276,1675,1702]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Figs 3EF</figureCitation>
. Dorsal shield of prosoma and sternum yellowish, without marks. Legs whitish, without darker rings. Opisthosoma yellowish, with dorsal and lateral black spots. Thoracic furrow absent. Clypeus unmodified. Chelicerae (
<figureCitation id="13582A0AE728FFC416E4D6EDDB7BF96D" box="[1202,1289,1747,1774]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Fig. 3D</figureCitation>
) with pair of proximo-lateral apophyses and pair of curved distal apophyses (distance between tips: 0.46). Pedipalp as in
<figureCitation id="13582A0AE728FFC4173AD6C9DEAFF8B5" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2AB</figureCitation>
; trochanter two times longer than wide, with small retrolateral apophysis (arrow
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<figureCitation id="13582A0AE728FFC416E6D725DB7AF8B5" box="[1200,1288,1819,1846]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Fig. 2B</figureCitation>
); femur with retrolatero-proximal apophysis (arrow
<quantity id="4C9B9B6AE728FFC4101FD77EDC06F8DA" box="[585,628,1855,1882]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="546" unit="in" value="2.0">2 in</quantity>
<figureCitation id="13582A0AE728FFC4102CD701DCA3F8D9" box="[634,721,1855,1882]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Fig. 2B</figureCitation>
); procursus (
<figureCitation id="13582A0AE728FFC41137D701DD93F8D9" box="[865,993,1855,1882]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2AD</figureCitation>
,
<figureCitation id="13582A0AE728FFC411BAD77EDA42F8D9" box="[1004,1072,1856,1882]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9AB</figureCitation>
) simple proximally but complex distally, with nearly angular prolatero-subdistal sclerite (arrow
<quantity id="4C9B9B6AE728FFC41105D75ADD0FF8FE" box="[851,893,1891,1918]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="546" unit="in" value="1.0">1 in</quantity>
<figureCitation id="13582A0AE728FFC411D5D75DDDAEF8FD" box="[899,988,1891,1918]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2C</figureCitation>
,
<figureCitation id="13582A0AE728FFC411B1D75ADA7FF8FD" box="[999,1037,1892,1918]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9A</figureCitation>
), nearly half-round, distally pointed distal membranous lamella (arrow
<quantity id="4C9B9B6AE728FFC4104DD7B6DC37F822" box="[539,581,1927,1954]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="3" pageNumber="546" unit="in" value="2.0">2 in</quantity>
<figureCitation id="13582A0AE728FFC4101DD7B9DCD6F821" box="[587,676,1927,1954]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2C</figureCitation>
,
<figureCitation id="13582A0AE728FFC410E6D7B6DCA7F821" box="[688,725,1928,1954]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9A</figureCitation>
), and two retrolateral membranous lamellae (arrows in
<figureCitation id="13582A0AE728FFC4176BD7B9DBEAF821" box="[1341,1432,1927,1954]" captionStart="FIGURE 2" captionStartId="4.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-22@4.[233,1355,181,1832]" captionTargetPageId="4" captionText="FIGURE 2. Belisana chuandiani sp. nov., holotype male (SYNU-Ar00308).AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="3" pageNumber="546">Figs 2D</figureCitation>
,
<figureCitation id="13582A0AE728FFC412C1D792DEC8F845" box="[151,186,1964,1990]" captionStart="FIGURE 9" captionStartId="13.[151,250,1797,1821]" captionTargetBox="[180,1408,385,1771]" captionTargetId="figure-115@13.[177,1411,381,1774]" captionTargetPageId="13" captionText="FIGURE 9. Procursus in prolateral and retrolateral views.AB. Belisana chuandiani sp. nov. (A. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; B. Retrolateral view, arrows point at two retrolateral membranous lamellae); CD. Belisana daxiangi sp. nov. (C. Prolateral view, arrow 1 points at prolatero-distal membranous lamella, arrows 2 and 3 point at two spine-shaped dorso-distal apophyses, arrow 4 points at ventral membranous lamella; D. Retrolateral view, arrow points at retrolatero-distal membranous lamella); EF. Belisana fengzheni sp. nov. (E. Prolateral view, arrows 1 and 2 point at two prolatero-distal membranous lamellae, arrow 3 points at spine-shaped dorso-distal apophysis; F. Retrolateral view, arrow points at retrolatero-distal membranous lamella). f = flap. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392266" httpUri="https://zenodo.org/record/8392266/files/figure.png" pageId="3" pageNumber="546">9B</figureCitation>
); bulb (
<figureCitation id="13582A0AE728FFC41347D795DF15F845" box="[273,359,1963,1990]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="3" pageNumber="546">Fig. 3C</figureCitation>
) with hooked apophysis and simple embolus. Legs with short vertical setae on metatarsi.
</paragraph>
<caption id="DF1C6607E72FFFC312C1D77EDDD2F86B" ID-DOI="http://doi.org/10.5281/zenodo.8392238" ID-Zenodo-Dep="8392238" httpUri="https://zenodo.org/record/8392238/files/figure.png" pageId="4" pageNumber="547" startId="4.[151,250,1856,1880]" targetBox="[233,1355,181,1832]" targetPageId="4" targetType="figure">
<paragraph id="8BDC368FE72FFFC312C1D77EDDD2F86B" blockId="4.[151,1437,1856,2024]" pageId="4" pageNumber="547">
<emphasis id="B917EA9DE72FFFC312C1D77EDF60F8DB" bold="true" box="[151,274,1856,1880]" pageId="4" pageNumber="547">FIGURE 2.</emphasis>
<taxonomicName id="4C634D0CE72FFFC3134ED77EDF9AF8DB" authority="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao, 2023" authorityName="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao" authorityYear="2023" box="[280,488,1856,1880]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="547" phylum="Arthropoda" rank="species" species="chuandiani" status="sp. nov.">
<emphasis id="B917EA9DE72FFFC3134ED77EDF9AF8DB" box="[280,488,1856,1880]" italics="true" pageId="4" pageNumber="547">Belisana chuandiani</emphasis>
</taxonomicName>
<emphasis id="B917EA9DE72FFFC313B8D77EDC4FF8DB" bold="true" box="[494,573,1856,1880]" pageId="4" pageNumber="547">
<taxonomicNameLabel id="A22457E6E72FFFC313B8D77EDC4FF8DB" box="[494,573,1856,1880]" pageId="4" pageNumber="547" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (SYNU-Ar00308). AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow 1 points at small retrolateral apophysis, arrow 2 points at retrolatero-proximal apophysis); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at nearly angular prolatero-subdistal sclerite, arrow 2 points at distally pointed distal membranous lamella; D. Retrolateral view, arrows point at two retrolateral membranous lamellae). b = bulb, ba = bulbal apophysis, e = embolus, pr = procursus. Scale bars: 0.10 (AB), 0.02 (CD).
</paragraph>
</caption>
<caption id="DF1C6607E72EFFC212C1D77EDAD1F847" ID-DOI="http://doi.org/10.5281/zenodo.8392242" ID-Zenodo-Dep="8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="5" pageNumber="548" startId="5.[151,250,1856,1880]" targetBox="[233,1355,181,1832]" targetPageId="5" targetType="figure">
<paragraph id="8BDC368FE72EFFC212C1D77EDAD1F847" blockId="5.[151,1436,1856,1988]" pageId="5" pageNumber="548">
<emphasis id="B917EA9DE72EFFC212C1D77EDF67F8DB" bold="true" box="[151,277,1856,1880]" pageId="5" pageNumber="548">FIGURE 3.</emphasis>
<taxonomicName id="4C634D0CE72EFFC2134BD77EDF82F8DB" authority="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao, 2023" authorityName="Zhao &amp; Yang &amp; Li &amp; Zheng &amp; Yao" authorityYear="2023" box="[285,496,1856,1880]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="548" phylum="Arthropoda" rank="species" species="chuandiani" status="sp. nov.">
<emphasis id="B917EA9DE72EFFC2134BD77EDF82F8DB" box="[285,496,1856,1880]" italics="true" pageId="5" pageNumber="548">Belisana chuandiani</emphasis>
</taxonomicName>
<emphasis id="B917EA9DE72EFFC213AFD77EDC39F8DB" bold="true" box="[505,587,1856,1880]" pageId="5" pageNumber="548">
<taxonomicNameLabel id="A22457E6E72EFFC213AFD77EDC39F8DB" box="[505,587,1856,1880]" pageId="5" pageNumber="548" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNUAr00309). A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH).
</paragraph>
</caption>
<paragraph id="8BDC368FE72DFFC11291D0A9DDDCFEC2" blockId="6.[151,1436,151,393]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D0A9DF6CFF32" bold="true" box="[199,286,151,177]" pageId="6" pageNumber="549">Female</emphasis>
(
<emphasis id="B917EA9DE72DFFC11366D0A9DFE9FF32" bold="true" box="[304,411,151,177]" pageId="6" pageNumber="549">
<typeStatus id="54D8882DE72DFFC11366D0A9DFE9FF32" box="[304,411,151,177]" pageId="6" pageNumber="549">paratype</typeStatus>
</emphasis>
, SYNU-Ar00309): Similar to male, habitus as in
<figureCitation id="13582A0AE72DFFC11185D0A9DA21FF32" box="[979,1107,151,177]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="6" pageNumber="549">Figs 3GH</figureCitation>
. Total length 1.63 (1.71 with clypeus), prosoma 0.60 long, 0.58 wide, opisthosoma 1.03 long, 0.70 wide; leg I missing. Eye interdistances and diameters: PME-PME 0.03, PME 0.08, PME-ALE 0.03. Sternum width/length: 0.44/0.47. Epigyne (
<figureCitation id="13582A0AE72DFFC116ACD0E1DB25FF7A" box="[1274,1367,223,249]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="6" pageNumber="549">Figs 3A</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11735D0E1DBE6FF7A" box="[1379,1428,223,249]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[194,1392,184,1916]" captionTargetId="figure-22@14.[191,1395,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Female genitalia in ventral and dorsal views. AB. Belisana chuandiani sp. nov.; CD. Belisana fengzheni sp. nov.; EF. Belisana zhengi Yao, Pham &amp; Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392270" httpUri="https://zenodo.org/record/8392270/files/figure.png" pageId="6" pageNumber="549">10A</figureCitation>
) simple and flat, with pair of postero-lateral pockets 0.41 apart (ep in
<figureCitation id="13582A0AE72DFFC111DDD13DDA75FE9E" box="[907,1031,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="6" pageNumber="549">Figs 3AB</figureCitation>
,
<figureCitation id="13582A0AE72DFFC11647D13DDA14FE9E" box="[1041,1126,259,285]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[194,1392,184,1916]" captionTargetId="figure-22@14.[191,1395,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Female genitalia in ventral and dorsal views. AB. Belisana chuandiani sp. nov.; CD. Belisana fengzheni sp. nov.; EF. Belisana zhengi Yao, Pham &amp; Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392270" httpUri="https://zenodo.org/record/8392270/files/figure.png" pageId="6" pageNumber="549">10AB</figureCitation>
). Vulva (
<figureCitation id="13582A0AE72DFFC1169ED13DDB50FE9E" box="[1224,1314,259,285]" captionStart="FIGURE 3" captionStartId="5.[151,250,1856,1880]" captionTargetBox="[233,1355,181,1832]" captionTargetId="figure-18@5.[233,1355,181,1832]" captionTargetPageId="5" captionText="FIGURE 3. Belisana chuandiani sp. nov., holotype male (CF, SYNU-Ar00308) and paratype female (AB, GH; SYNU- Ar00309).A. Epigyne, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.20 (EH)." figureDoi="http://doi.org/10.5281/zenodo.8392242" httpUri="https://zenodo.org/record/8392242/files/figure.png" pageId="6" pageNumber="549">Figs 3B</figureCitation>
,
<figureCitation id="13582A0AE72DFFC1177BD13DDB2EFE9E" box="[1325,1372,259,285]" captionStart="FIGURE 10" captionStartId="14.[151,250,1943,1967]" captionTargetBox="[194,1392,184,1916]" captionTargetId="figure-22@14.[191,1395,181,1919]" captionTargetPageId="14" captionText="FIGURE 10. Female genitalia in ventral and dorsal views. AB. Belisana chuandiani sp. nov.; CD. Belisana fengzheni sp. nov.; EF. Belisana zhengi Yao, Pham &amp; Li, 2015. ep = epigynal pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.8392270" httpUri="https://zenodo.org/record/8392270/files/figure.png" pageId="6" pageNumber="549">10B</figureCitation>
) with ridge-shaped anterior arch and pair of curved, long elliptic pore plates.
</paragraph>
<paragraph id="8BDC368FE72DFFC11291D175DAE4FEE6" blockId="6.[151,1436,151,393]" box="[199,1174,330,357]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D175DFF7FEE7" bold="true" box="[199,389,330,357]" pageId="6" pageNumber="549">Natural history.</emphasis>
The species was collected in a seasonal rainforest by canopy fogging.
</paragraph>
</subSubSection>
<subSubSection id="C3796504E72DFFC11291D150DD78FE0A" box="[199,778,366,393]" pageId="6" pageNumber="549" type="distribution">
<paragraph id="8BDC368FE72DFFC11291D150DD78FE0A" blockId="6.[151,1436,151,393]" box="[199,778,366,393]" pageId="6" pageNumber="549">
<emphasis id="B917EA9DE72DFFC11291D150DF13FE0B" bold="true" box="[199,353,366,392]" pageId="6" pageNumber="549">Distribution.</emphasis>
<collectingCountry id="F374761FE72DFFC11331D151DFD8FE0A" box="[359,426,367,393]" name="China" pageId="6" pageNumber="549">China</collectingCountry>
(
<collectingRegion id="49A7F86DE72DFFC113EFD151DC61FE0A" box="[441,531,367,393]" country="China" name="Yunnan" pageId="6" pageNumber="549">Yunnan</collectingRegion>
,
<typeStatus id="54D8882DE72DFFC11048D151DC3CFE0A" box="[542,590,367,393]" pageId="6" pageNumber="549">type</typeStatus>
locality;
<figureCitation id="13582A0AE72DFFC110EED151DC8BFE0A" box="[696,761,367,393]" captionStart="FIGURE 1" captionStartId="3.[151,250,957,981]" captionTargetBox="[151,1436,179,931]" captionTargetId="figure-18@3.[151,1436,179,931]" captionTargetPageId="3" captionText="FIGURE 1. Distribution records of the Belisana species discussed in this paper. 1, B. chuandiani sp. nov.; 2, B. daxiangi sp. nov.; 3, B. fengzheni sp. nov.; 45, B. zhengi." figureDoi="http://doi.org/10.5281/zenodo.8392236" httpUri="https://zenodo.org/record/8392236/files/figure.png" pageId="6" pageNumber="549">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>