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<mods:title id="6C82D989C67FB565AE3D7B516826CF82">Revision of F. R. C. Reeds Ordovician trilobite types from Myanmar (Burma) and western Yunnan Province, China</mods:title>
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<treatment id="03CB87846E761D2BFF3AC6BB48E8F003" ID-DOI="http://doi.org/10.5281/zenodo.6810348" ID-GBIF-Taxon="197214828" ID-Zenodo-Dep="6810348" LSID="urn:lsid:plazi:treatment:03CB87846E761D2BFF3AC6BB48E8F003" httpUri="http://treatment.plazi.org/id/03CB87846E761D2BFF3AC6BB48E8F003" lastPageNumber="336" pageId="35" pageNumber="336">
<subSubSection id="C37865196E761D2BFF3AC6BB4BECF658" box="[151,503,762,789]" pageId="35" pageNumber="336" type="nomenclature">
<paragraph id="8BDD36926E761D2BFF3AC6BB4BECF658" blockId="35.[151,503,762,789]" box="[151,503,762,789]" pageId="35" pageNumber="336">
<heading id="D09581FE6E761D2BFF3AC6BB4BECF658" bold="true" box="[151,503,762,789]" fontSize="11" level="1" pageId="35" pageNumber="336" reason="1">
<taxonomicName id="4C624D116E761D2BFF3AC6BB4BECF658" ID-CoL="6VRX" authority="Rouault, 1847" authorityName="Rouault" authorityYear="1847" box="[151,503,762,789]" class="Trilobita" family="Calymenidae" genus="Prionocheilus" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFF3AC6BB4BECF658" bold="true" box="[151,503,762,789]" pageId="35" pageNumber="336">
<emphasis id="B916EA806E761D2BFF3AC6BB4B5AF659" bold="true" box="[151,321,762,789]" italics="true" pageId="35" pageNumber="336">Prionocheilus</emphasis>
<bibRefCitation id="EFF34B636E761D2BFEE5C6BA4BECF658" author="Rouault, M." box="[328,503,762,789]" pageId="35" pageNumber="336" pagination="309 - 328" refId="ref46935" refString="Rouault, M. (1847) Extrait du Memoire sur les Trilobites du Departement d'Ille-et-Villaine. Bulletin de la Societe Geologique du France, 4, 309 - 328." type="journal article" year="1847">Rouault, 1847</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37865196E761D2BFF3AC77E495BF614" box="[151,832,831,856]" pageId="35" pageNumber="336" type="type_taxon">
<paragraph id="8BDD36926E761D2BFF3AC77E495BF614" blockId="35.[151,832,831,856]" box="[151,832,831,856]" pageId="35" pageNumber="336">
<emphasis id="B916EA806E761D2BFF3AC77E4B05F61B" bold="true" box="[151,286,831,855]" pageId="35" pageNumber="336">
<typeStatus id="54D988306E761D2BFF3AC77E4AD7F61B" box="[151,204,831,855]" pageId="35" pageNumber="336">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C624D116E761D2BFE87C77E48B2F61B" authority="Rouault, 1847" authorityName="Rouault" authorityYear="1847" box="[298,681,831,855]" class="Trilobita" family="Calymenidae" genus="Prionocheilus" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="verneuli">
<emphasis id="B916EA806E761D2BFE87C77E480AF61B" box="[298,529,831,855]" italics="true" pageId="35" pageNumber="336">Prionocheilus verneuli</emphasis>
<bibRefCitation id="EFF34B636E761D2BFDB5C77E48B2F61B" author="Rouault, M." box="[536,681,831,855]" pageId="35" pageNumber="336" pagination="309 - 328" refId="ref46935" refString="Rouault, M. (1847) Extrait du Memoire sur les Trilobites du Departement d'Ille-et-Villaine. Bulletin de la Societe Geologique du France, 4, 309 - 328." type="journal article" year="1847">Rouault, 1847</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C37865196E761D2BFF3AC7C448E8F003" pageId="35" pageNumber="336" type="discussion">
<paragraph id="8BDD36926E761D2BFF3AC7C448E8F003" blockId="35.[151,1437,901,1360]" pageId="35" pageNumber="336">
<emphasis id="B916EA806E761D2BFF3AC7C44B0EF6D3" bold="true" box="[151,277,901,927]" pageId="35" pageNumber="336">Discussion</emphasis>
. We accept the priority of the generic name
<taxonomicName id="4C624D116E761D2BFC8DC7C449A6F6D3" authorityName="Rouault" authorityYear="1847" box="[800,957,901,927]" class="Trilobita" family="Calymenidae" genus="Prionocheilus" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFC8DC7C449A6F6D3" box="[800,957,901,927]" italics="true" pageId="35" pageNumber="336">Prionocheilus</emphasis>
</taxonomicName>
over
<taxonomicName id="4C624D116E761D2BFBAEC7C44E90F6D3" authorityName="Hawle &amp; Corda" authorityYear="1847" box="[1027,1163,901,927]" class="Trilobita" family="Calymenidae" genus="Pharostoma" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFBAEC7C44E90F6D3" box="[1027,1163,901,927]" italics="true" pageId="35" pageNumber="336">Pharostoma</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFF34B636E761D2BFB66C7C74F57F6D3" author="Dean, W. T." box="[1227,1356,901,928]" pageId="35" pageNumber="336" pagination="1 - 60" refId="ref43521" refString="Dean, W. T. (1971) The trilobites of the Chair of Kildare Limestone (upper Ordovician) of eastern Ireland. Part 1. Monographs of the Palaeontographical Society, 531, 1 - 60." type="journal article" year="1971">Dean 1971</bibRefCitation>
).
<bibRefCitation id="EFF34B636E761D2BFACEC7C44AC0F688" author="Reed, F. R. C." pageId="35" pageNumber="336" pagination="1 - 98" refId="ref46807" refString="Reed, F. R. C. (1915) Supplementary Memoir on new Ordovician and Silurian fossils from the Northern Shan States. Palaeontologia Indica, New Series 6, 1 - 98." type="book chapter" year="1915">Reed (1915</bibRefCitation>
, p. 46) erected a species based on a cranidium,
<taxonomicName id="4C624D116E761D2BFD45C7E84E47F68F" authorityName="Reed" authorityYear="1915" box="[744,1116,937,963]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="liluensis" subGenus="Pharostoma">
<emphasis id="B916EA806E761D2BFD45C7E84943F68F" box="[744,856,937,963]" italics="true" pageId="35" pageNumber="336">Calymene</emphasis>
(
<emphasis id="B916EA806E761D2BFCCAC7E849F4F68F" box="[871,1007,937,963]" italics="true" pageId="35" pageNumber="336">Pharostoma</emphasis>
)
<emphasis id="B916EA806E761D2BFC52C7E84E47F68F" box="[1023,1116,937,963]" italics="true" pageId="35" pageNumber="336">liluensis</emphasis>
</taxonomicName>
, which might be regarded as a
<taxonomicName id="4C624D116E761D2BFF04C78C4BBFF6AB" box="[169,420,973,999]" class="Trilobita" family="Calymenidae" genus="Prionocheilus" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B916EA806E761D2BFF04C78C4B5DF6AB" box="[169,326,973,999]" italics="true" pageId="35" pageNumber="336">Prionocheilus</emphasis>
species.
</taxonomicName>
<bibRefCitation id="EFF34B636E761D2BFE04C78C486AF6AB" author="Zhou, T. - M. &amp; Lui, Y. - R. &amp; Meng, X. - S. &amp; Sun, Z. - H." box="[425,625,973,999]" pageId="35" pageNumber="336" pagination="104 - 226" refId="ref49276" refString="Zhou, T. - M., Lui, Y. - R., Meng, X. - S. &amp; Sun, Z. - H. (1977) Trilobita. In: Geological Institute of Hubei (Ed.), Palaeontological Atlas of Central and Southern China. Geological Publishing House, Beijing, pp. 104 - 226. [in Chinese]" type="book chapter" year="1977">
Zhou
<emphasis id="B916EA806E761D2BFE47C78F483AF6AB" box="[490,545,973,999]" italics="true" pageId="35" pageNumber="336">et al.</emphasis>
(1977)
</bibRefCitation>
erected the genus
<taxonomicName id="4C624D116E761D2BFC91C78F49BCF6AB" authorityName="Zhou" authorityYear="1977" box="[828,935,974,999]" class="Trilobita" family="Calymenidae" genus="Xuanenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFC91C78F49BCF6AB" box="[828,935,974,999]" italics="true" pageId="35" pageNumber="336">Xuanenia</emphasis>
</taxonomicName>
,
<typeStatus id="54D988306E761D2BFC19C78F49FFF6A4" box="[948,996,974,1000]" pageId="35" pageNumber="336">type</typeStatus>
species
<taxonomicName id="4C624D116E761D2BFBEDC78F4F47F6AB" authority="Zhou, 1977" authorityName="Zhou" authorityYear="1977" box="[1088,1372,973,1000]" class="Trilobita" family="Calymenidae" genus="Xuanenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="pustulosus">
<emphasis id="B916EA806E761D2BFBEDC78F4E4AF6AB" box="[1088,1105,974,999]" italics="true" pageId="35" pageNumber="336">X</emphasis>
.
<emphasis id="B916EA806E761D2BFBF3C78C4ECFF6AB" box="[1118,1236,973,999]" italics="true" pageId="35" pageNumber="336">pustulosus</emphasis>
<bibRefCitation id="EFF34B636E761D2BFB74C78C4F47F6AB" author="Zhou, T. - M. &amp; Lui, Y. - R. &amp; Meng, X. - S. &amp; Sun, Z. - H." box="[1241,1372,973,999]" pageId="35" pageNumber="336" pagination="104 - 226" refId="ref49276" refString="Zhou, T. - M., Lui, Y. - R., Meng, X. - S. &amp; Sun, Z. - H. (1977) Trilobita. In: Geological Institute of Hubei (Ed.), Palaeontological Atlas of Central and Southern China. Geological Publishing House, Beijing, pp. 104 - 226. [in Chinese]" type="book chapter" year="1977">Zhou, 1977</bibRefCitation>
</taxonomicName>
, from central South
<collectingCountry id="F37576026E761D2BFE9EC7B04B62F147" box="[307,377,1009,1035]" name="China" pageId="35" pageNumber="336">China</collectingCountry>
, based on an imperfectly preserved cranidium. The same genus and species were subsequently described by
<bibRefCitation id="EFF34B636E761D2BFE81C0544BE0F163" author="Tripp, R. P. &amp; Zhou, Z. - Y. &amp; Pan, Z. - Q." box="[300,507,1045,1071]" pageId="35" pageNumber="336" pagination="25 - 68" refId="ref47815" refString="Tripp, R. P., Zhou, Z. - Y. &amp; Pan, Z. - Q. (1989) Trilobites from the Upper Ordovician Tangtou Formation, Jiangsu Province, China. Transactions of the Royal Society of Edinburgh, Earth Sciences, 80, 25 - 68. https: // doi. org / 10.1017 / S 0263593300012256" type="journal article" year="1989">
Tripp
<emphasis id="B916EA806E761D2BFEDCC0574BB1F163" box="[369,426,1045,1071]" italics="true" pageId="35" pageNumber="336">et al.</emphasis>
(1989)
</bibRefCitation>
for material from the Upper Ordovican Tangtou Formation,
<collectingRegion id="49A6F8706E761D2BFB09C0544F7DF163" box="[1188,1382,1045,1071]" country="China" name="Jiangsu" pageId="35" pageNumber="336">Jiangsu Province</collectingRegion>
. The similarity of some of this material to
<taxonomicName id="4C624D116E761D2BFDE7C07848C9F11F" authorityName="Hawle &amp; Corda" authorityYear="1847" box="[586,722,1081,1107]" class="Trilobita" family="Calymenidae" genus="Pharostoma" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFDE7C07848C9F11F" box="[586,722,1081,1107]" italics="true" pageId="35" pageNumber="336">Pharostoma</emphasis>
</taxonomicName>
<taxonomicName id="4C624D116E761D2BFD4FC0784924F11F" authorityName="Reed" authorityYear="1915" box="[738,831,1081,1107]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="liluensis">
<emphasis id="B916EA806E761D2BFD4FC0784924F11F" box="[738,831,1081,1107]" italics="true" pageId="35" pageNumber="336">liluensis</emphasis>
</taxonomicName>
was noted already by
<bibRefCitation id="EFF34B636E761D2BFBECC07B4F50F11F" author="Fortey, R. A. &amp; Cocks, L. R. M." box="[1089,1355,1081,1108]" pageId="35" pageNumber="336" pagination="43 - 56" refId="ref43940" refString="Fortey, R. A. &amp; Cocks, L. R. M. (1998) Biogeography and palaeogeography of the Sibumasu terrane in the Ordovician: a review. In: Hall, R. &amp; Holloway, J. D. (Eds.), Biogeography and Geological Evolution of SE Asia. Backhuys Publishers, Leiden, pp. 43 - 56." type="book chapter" year="1998">Fortey &amp; Cocks (1998)</bibRefCitation>
, and is confirmed by the present study.
<bibRefCitation id="EFF34B636E761D2BFDAAC01C48C0F13B" author="Tripp, R. P. &amp; Zhou, Z. - Y. &amp; Pan, Z. - Q." box="[519,731,1117,1143]" pageId="35" pageNumber="336" pagination="25 - 68" refId="ref47815" refString="Tripp, R. P., Zhou, Z. - Y. &amp; Pan, Z. - Q. (1989) Trilobites from the Upper Ordovician Tangtou Formation, Jiangsu Province, China. Transactions of the Royal Society of Edinburgh, Earth Sciences, 80, 25 - 68. https: // doi. org / 10.1017 / S 0263593300012256" type="journal article" year="1989">
Tripp
<emphasis id="B916EA806E761D2BFDE0C01F489CF13B" box="[589,647,1117,1143]" italics="true" pageId="35" pageNumber="336">et al.</emphasis>
(1989)
</bibRefCitation>
remarked that “
<taxonomicName id="4C624D116E761D2BFC38C01F4E1BF13B" authorityName="Zhou" authorityYear="1977" box="[917,1024,1118,1143]" class="Trilobita" family="Calymenidae" genus="Xuanenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFC38C01F4E1BF13B" box="[917,1024,1118,1143]" italics="true" pageId="35" pageNumber="336">Xuanenia</emphasis>
</taxonomicName>
is extremely close to
<taxonomicName id="4C624D116E761D2BFB52C01C4B31F1D7" authority="Roualt, 1847" authorityName="Rouault" authorityYear="1847" class="Trilobita" family="Calymenidae" genus="Prionocheilus" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFB52C01C4F87F13B" box="[1279,1436,1117,1143]" italics="true" pageId="35" pageNumber="336">Prionocheilus</emphasis>
Roualt, 1847
</taxonomicName>
(sic)[and] differs only in the effaced lateral glabellar lobes and furrows and smaller pygidium with fewer segments.” There is little difference with regard to glabellar lobes and furrows between the cranidium figured by
<bibRefCitation id="EFF34B636E761D2BFF16C0884B9EF1AF" author="Tripp, R. P. &amp; Zhou, Z. - Y. &amp; Pan, Z. - Q." box="[187,389,1225,1251]" pageId="35" pageNumber="336" pagination="25 - 68" refId="ref47815" refString="Tripp, R. P., Zhou, Z. - Y. &amp; Pan, Z. - Q. (1989) Trilobites from the Upper Ordovician Tangtou Formation, Jiangsu Province, China. Transactions of the Royal Society of Edinburgh, Earth Sciences, 80, 25 - 68. https: // doi. org / 10.1017 / S 0263593300012256" type="journal article" year="1989">
Tripp
<emphasis id="B916EA806E761D2BFEADC08B4B22F1AF" box="[256,313,1225,1251]" italics="true" pageId="35" pageNumber="336">et al.</emphasis>
(1989
</bibRefCitation>
, fig. 17k) and the
<typeStatus id="54D988306E761D2BFDF8C08B489EF1A8" box="[597,645,1226,1252]" pageId="35" pageNumber="336">type</typeStatus>
specimen of
<taxonomicName id="4C624D116E761D2BFC8AC08849B4F1AF" authorityName="Hawle &amp; Corda" authorityYear="1847" box="[807,943,1225,1251]" class="Trilobita" family="Calymenidae" genus="Pharostoma" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFC8AC08849B4F1AF" box="[807,943,1225,1251]" italics="true" pageId="35" pageNumber="336">Pharostoma</emphasis>
</taxonomicName>
<taxonomicName id="4C624D116E761D2BFC13C0884E00F1AF" authorityName="Reed" authorityYear="1915" box="[958,1051,1225,1251]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="liluensis">
<emphasis id="B916EA806E761D2BFC13C0884E00F1AF" box="[958,1051,1225,1251]" italics="true" pageId="35" pageNumber="336">liluensis</emphasis>
</taxonomicName>
, which in turn is not significantly different from cranidia of
<taxonomicName id="4C624D116E761D2BFE6FC0AC4882F044" box="[450,665,1261,1288]" class="Trilobita" family="Calymenidae" genus="Prionocheilus" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B916EA806E761D2BFE6FC0AC4844F04B" box="[450,607,1261,1287]" italics="true" pageId="35" pageNumber="336">Prionocheilus</emphasis>
spp.
</taxonomicName>
; we prefer to use that generic name pending further clarification of
<taxonomicName id="4C624D116E761D2BFF3AC1534B19F067" authorityName="Zhou" authorityYear="1977" box="[151,258,1298,1323]" class="Trilobita" family="Calymenidae" genus="Xuanenia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="genus">
<emphasis id="B916EA806E761D2BFF3AC1534B19F067" box="[151,258,1298,1323]" italics="true" pageId="35" pageNumber="336">Xuanenia</emphasis>
</taxonomicName>
. It is possible that
<taxonomicName id="4C624D116E761D2BFE7BC1534854F067" authorityName="Reed" authorityYear="1915" box="[470,591,1297,1323]" class="Trilobita" family="Calymenidae" genus="Calymene" kingdom="Animalia" order="Phacopida" pageId="35" pageNumber="336" phylum="Arthropoda" rank="species" species="liluensis">
<emphasis id="B916EA806E761D2BFE7BC1534854F067" box="[470,591,1297,1323]" italics="true" pageId="35" pageNumber="336">P. liluensis</emphasis>
</taxonomicName>
has priority over subsequently named species in this group, but a comparative study of adjacent species is needed to clarify this.
</paragraph>
</subSubSection>
</treatment>
</document>