treatments-xml/data/03/AF/87/03AF878BFFCAFFB6FF32F8BAC538FE4E.xml
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<mods:title id="DB280CF659295D5AD52F08C1376C1FF1">Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species</mods:title>
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<mods:namePart id="B99DF53C8240CDFE03EA63E865D35070">Lee, Young June</mods:namePart>
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<mods:namePart id="3A8BA997FA7BC576523EA11C579E7E82">Marshall, David C.</mods:namePart>
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<mods:namePart id="02D236BBA3BFE8A23742EADBE533DFDD">Mohagan, Alma</mods:namePart>
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<mods:namePart id="C3C4B0C4845251C58132E64670ED1115">Hill, Kathy B. R.</mods:namePart>
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<paragraph id="8BB9369DFFCAFFB7FF32F8BAC606FBA3" blockId="8.[151,551,1907,1933]" box="[151,551,1907,1933]" pageId="8" pageNumber="195">
<heading id="D0F181F1FFCAFFB7FF32F8BAC606FBA3" bold="true" box="[151,551,1907,1933]" fontSize="11" level="1" pageId="8" pageNumber="195" reason="1">
<emphasis id="B972EA8FFFCAFFB7FF32F8BAC606FBA3" bold="true" box="[151,551,1907,1933]" pageId="8" pageNumber="195">
Tribe
<taxonomicName id="4C064D1EFFCAFFB7FF47F8BAC606FBA3" ID-CoL="8HBWN" authority="Buckton, 1889" authorityName="Buckton" authorityYear="1889" box="[226,551,1907,1933]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini Buckton, 1889</taxonomicName>
</emphasis>
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<paragraph id="8BB9369DFFCAFFB6FF32F870C6DCFDA6" blockId="8.[151,1436,1977,2038]" lastBlockId="9.[151,1437,151,608]" lastPageId="9" lastPageNumber="196" pageId="8" pageNumber="195">
<emphasis id="B972EA8FFFCAFFB7FF32F870C52AFBFC" bold="true" box="[151,267,1977,2002]" pageId="8" pageNumber="195">Remarks.</emphasis>
&quot;Huechysaria&quot; (= Huechysini Distant, 1905) was proposed by
<bibRefCitation id="EF974B6CFFCAFFB7FC70F870C0A7FBFC" author="Distant" box="[981,1158,1977,2002]" pageId="8" pageNumber="201" refString="Distant, W. L. (1905 b) Rhynchotal notes - XXXII. Annals and Magazine of Natural History, (7), 15, 478 - 487. http: // dx. doi. org / 10.1080 / 03745480509442837" type="journal article" year="1905" yearSuffix="b">Distant (1905b)</bibRefCitation>
with
<taxonomicName id="4C064D1EFFCAFFB7FB6DF870C5BAFBD8" authority="Amyot &amp; Audinet-Serville, 1843" authorityName="Amyot &amp; Audinet-Serville" authorityYear="1843" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="195" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCAFFB7FB6DF870C109FBFC" box="[1224,1320,1977,2002]" italics="true" pageId="8" pageNumber="195">Huechys</emphasis>
<bibRefCitation id="EF974B6CFFCAFFB7FA95F873C5BAFBD8" author="Amyot" pageId="8" pageNumber="201" refString="Amyot, C. J. B. &amp; Audinet-Serville, J. G. (1843) Homopteres. Homoptera Latr. pp. 455 - 588 In: Histoire naturelle des insectes, hemipteres. Librairie Encyclopedique de Roret, Paris, 676 pp." type="book chapter" year="1843">Amyot &amp; Audinet-Serville, 1843</bibRefCitation>
</taxonomicName>
as the
<typeStatus id="54BD883FFFCAFFB7FE51F817C605FBD8" box="[500,548,2014,2038]" pageId="8" pageNumber="195">type</typeStatus>
genus mainly by the opaque or semiopaque wings. However, the molecular phylogenetic study by
<bibRefCitation id="EF974B6CFFCBFFB6FE07FF5EC682FC9E" box="[418,675,151,176]" pageId="9" pageNumber="202" refString="Marshall, D. C., Hill, K. B. R., Moulds, M., Vanderpool, D., Cooley, J. R., Mohagan, A. B. &amp; Simon, C. (2015) Inflation of molecular clock rates and dates: molecular phylogenetics, biogeography, and diversification of a global cicada radiation from Australasia (Hemiptera: Cicadidae: Cicadettini). Systematic Biology, 65 (1), 16 - 34. http: // dx. doi. org / 10.1093 / sysbio / syv 069" type="journal article">
<collectingCountry id="F311760DFFCBFFB6FE07FF5EC627FC9E" box="[418,518,151,176]" name="Marshall Islands" pageId="9" pageNumber="196">Marshall</collectingCountry>
<emphasis id="B972EA8FFFCBFFB6FDB4FF50C667FC9E" box="[529,582,151,176]" italics="true" pageId="9" pageNumber="196">et al</emphasis>
. (2015)
</bibRefCitation>
suggests that
<taxonomicName id="4C064D1EFFCBFFB6FCEBFF5EC78FFC9E" box="[846,942,151,176]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FCEBFF5EC78FFC9E" box="[846,942,151,176]" italics="true" pageId="9" pageNumber="196">Huechys</emphasis>
</taxonomicName>
should be placed within
<taxonomicName id="4C064D1EFFCBFFB6FB78FF5EC17DFC9E" box="[1245,1372,151,176]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
. This suggested placement is well supported morphologically by the presence of a pair of flat upper lobes, a welldeveloped dorsal beak, a duck-bill-shaped median lobe of uncus, and a pair of clasper-like processes, which are commonly seen in the male genitalia of
<taxonomicName id="4C064D1EFFCBFFB6FDD9FECDC6FCFD33" box="[636,733,260,285]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FDD9FECDC6FCFD33" box="[636,733,260,285]" italics="true" pageId="9" pageNumber="196">Huechys</emphasis>
</taxonomicName>
as well as that of the
<typeStatus id="54BD883FFFCBFFB6FC56FECCC002FD33" box="[1011,1059,261,285]" pageId="9" pageNumber="196">type</typeStatus>
genus of
<taxonomicName id="4C064D1EFFCBFFB6FB04FECDC13EFD33" box="[1185,1311,260,285]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
,
<taxonomicName id="4C064D1EFFCBFFB6FA8AFECDC563FD6E" authority="Kolenati, 1857" authorityName="Kolenati" authorityYear="1857" class="Insecta" family="Cicadidae" genus="Cicadetta" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FA8AFECDC1BDFD33" box="[1327,1436,260,285]" italics="true" pageId="9" pageNumber="196">Cicadetta</emphasis>
Kolenati, 1857
</taxonomicName>
.
<taxonomicName id="4C064D1EFFCBFFB6FEEAFEEEC58EFD6E" box="[335,431,295,320]" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FEEAFEEEC58EFD6E" box="[335,431,295,320]" italics="true" pageId="9" pageNumber="196">Huechys</emphasis>
</taxonomicName>
also has the following characteristics, which were listed as distinguishing characters of
<taxonomicName id="4C064D1EFFCBFFB6FF32FE85C533FD4B" box="[151,274,332,357]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
by
<bibRefCitation id="EF974B6CFFCBFFB6FEE9FE85C621FD4B" author="Moulds" box="[332,512,332,357]" pageId="9" pageNumber="202" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds (2005)</bibRefCitation>
: pygofer upper lobes flat; and pseudoparameres present (see
<figureCitation id="133D2A18FFCBFFB6FB50FE85C163FD4A" box="[1269,1346,332,357]" captionStart="FIGURE 4" captionStartId="9.[151,250,1338,1360]" captionTargetBox="[313,1275,669,1307]" captionTargetId="figure@9.[305,1282,663,1316]" captionTargetPageId="9" captionText="FIGURE 4. Male pygofer of Huechys funebris Lee, 2011, holotype, male, Leyte, Philippines. A, Ventral view. B, Llateral view (adapted from Lee, 2011)." httpUri="https://zenodo.org/record/262367/files/figure.png" pageId="9" pageNumber="196">Fig. 4</figureCitation>
). Here, Huechysini
<emphasis id="B972EA8FFFCBFFB6FEB8FEA6C5A4FDA6" bold="true" box="[285,389,367,392]" pageId="9" pageNumber="196">syn. nov.</emphasis>
is synonymized with
<taxonomicName id="4C064D1EFFCBFFB6FDDEFEA6C6DBFDA6" box="[635,762,367,392]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
.
</paragraph>
<paragraph id="8BB9369DFFCBFFB6FF62FE5DC538FE4E" blockId="9.[151,1437,151,608]" pageId="9" pageNumber="196">
<bibRefCitation id="EF974B6CFFCBFFB6FF62FE5DC553FD83" author="Moulds" box="[199,370,404,429]" pageId="9" pageNumber="202" refString="Moulds, M. S. (2005) An appraisal of the higher classification of cicadas (Hemiptera: Cicadoidea) with special reference to the Australian fauna. Records of the Australian Museum, 57, 375 - 446. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1447" type="journal article" year="2005">Moulds (2005)</bibRefCitation>
also proposed as a diagnostic character for
<taxonomicName id="4C064D1EFFCBFFB6FCDBFE5DC7D8FD83" box="[894,1017,404,429]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
the timbals not extending ventrally beyond the wing bases although he later found the opposite character-state in
<taxonomicName id="4C064D1EFFCBFFB6FC5FFE7EC10AFDFE" authority="Moulds, 2012" authorityName="Moulds" authorityYear="2012" box="[1018,1323,439,464]" class="Insecta" family="Cicadidae" genus="Uradolichos" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FC5FFE7EC0A7FDFE" box="[1018,1158,439,464]" italics="true" pageId="9" pageNumber="196">Uradolichos</emphasis>
Moulds, 2012
</taxonomicName>
, which he placed in
<taxonomicName id="4C064D1EFFCBFFB6FEA1FE15C5A2FDDB" box="[260,387,476,501]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
. Because the timbals slightly extend below the wing bases ventrally in specimens of
<taxonomicName id="4C064D1EFFCBFFB6FA99FE15C5A6FE36" authority="Haupt, 1924" authorityName="Haupt" authorityYear="1924" class="Insecta" family="Cicadidae" genus="Huechys" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="B972EA8FFFCBFFB6FA99FE15C4CFFE36" italics="true" pageId="9" pageNumber="196">Huechys parvula</emphasis>
<bibRefCitation id="EF974B6CFFCBFFB6FF5CFDC9C5A6FE36" author="Haupt" box="[249,391,512,536]" pageId="9" pageNumber="201" refString="Haupt, H. (1924) Die Gattung Huechys Am. et Serv. (Homopt.) Beitrage zu einer Monographie. Deutsche entomologische Zeitschrift, 1924, 201 - 225." type="journal article" year="1924">Haupt, 1924</bibRefCitation>
</taxonomicName>
listed below and some examined specimens of
<taxonomicName id="4C064D1EFFCBFFB6FC1BFDC8C0D4FE36" authority="Matsumura, 1927" authorityName="Matsumura" authorityYear="1927" box="[958,1269,512,536]" class="Insecta" family="Cicadidae" genus="Kosemia" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FC1BFDC8C001FE36" box="[958,1056,513,536]" italics="true" pageId="9" pageNumber="196">Kosemia</emphasis>
Matsumura, 1927
</taxonomicName>
and
<taxonomicName id="4C064D1EFFCBFFB6FA96FDC8C51CFE13" authority="Kolenati, 1857" authorityName="Kolenati" authorityYear="1857" class="Insecta" family="Cicadidae" genus="Tettigetta" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B972EA8FFFCBFFB6FA96FDC8C1BDFE36" box="[1331,1436,513,536]" italics="true" pageId="9" pageNumber="196">Tettigetta</emphasis>
Kolenati,
<quantity id="4CFE9B78FFCBFFB6FEA0FDEDC51CFE13" box="[261,317,548,573]" metricMagnitude="1" metricUnit="m" metricValue="4.71678" pageId="9" pageNumber="196" unit="in" value="1857.0">1857</quantity>
</taxonomicName>
in the first author's care, this character may have to be excluded from the characters for diagnosing
<taxonomicName id="4C064D1EFFCBFFB6FF32FD8EC537FE4E" box="[151,278,583,608]" class="Insecta" family="Cicadidae" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="196" phylum="Arthropoda" rank="tribe" tribe="Cicadettini">Cicadettini</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
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