Cicadas (Hemiptera: Cicadidae) from Camiguin of Mindanao Province and Dinagat Island in the Philippines, with a new genus and three new species
Author
Lee, Young June
Author
Marshall, David C.
Author
Mohagan, Alma
Author
Hill, Kathy B. R.
text
Zootaxa
2016
4097
2
187
202
journal article
51470
10.11646/zootaxa.4097.2.2
fcd803c5-6723-4fbe-8db8-6f39b2168947
1175-5326
262363
732FE827-45BC-4AA6-AA6C-658C02E00FAC
Tribe
Cicadettini Buckton, 1889
Remarks.
"Huechysaria" (= Huechysini Distant, 1905) was proposed by
Distant (1905b)
with
Huechys
Amyot & Audinet-Serville, 1843
as the
type
genus mainly by the opaque or semiopaque wings. However, the molecular phylogenetic study by
Marshall
et al
. (2015)
suggests that
Huechys
should be placed within
Cicadettini
. This suggested placement is well supported morphologically by the presence of a pair of flat upper lobes, a welldeveloped dorsal beak, a duck-bill-shaped median lobe of uncus, and a pair of clasper-like processes, which are commonly seen in the male genitalia of
Huechys
as well as that of the
type
genus of
Cicadettini
,
Cicadetta
Kolenati, 1857
.
Huechys
also has the following characteristics, which were listed as distinguishing characters of
Cicadettini
by
Moulds (2005)
: pygofer upper lobes flat; and pseudoparameres present (see
Fig. 4
). Here, Huechysini
syn. nov.
is synonymized with
Cicadettini
.
Moulds (2005)
also proposed as a diagnostic character for
Cicadettini
the timbals not extending ventrally beyond the wing bases although he later found the opposite character-state in
Uradolichos
Moulds, 2012
, which he placed in
Cicadettini
. Because the timbals slightly extend below the wing bases ventrally in specimens of
Huechys parvula
Haupt, 1924
listed below and some examined specimens of
Kosemia
Matsumura, 1927
and
Tettigetta
Kolenati,
1857
in the first author's care, this character may have to be excluded from the characters for diagnosing
Cicadettini
.