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<document ID-DOI="http://dx.doi.org/10.3897/phytokeys.198.79514" ID-Pensoft-Pub="1314-2003-198-1" ID-Pensoft-UUID="486F1F1B4F5854D2831AAA341B9A322C" ModsDocID="1314-2003-198-1" checkinTime="1654168732655" checkinUser="pensoft" docAuthor="Aubriot, Xavier &amp; Knapp, Sandra" docDate="2022" docId="49333F06B6534328EC583216A79EEFF9" docLanguage="en" docName="PhytoKeys 198: 1-270" docOrigin="PhytoKeys 198" docPubDate="2022-06-01" docSource="http://dx.doi.org/10.3897/phytokeys.198.79514" docTitle="Solanum camranhense Dy Phon &amp; Hul, Fl. Cambodge, Laos &amp; Vietnam 35: 16. 2014." docType="treatment" docVersion="1" id="486F1F1B4F5854D2831AAA341B9A322C" lastPageNumber="1" masterDocId="486F1F1B4F5854D2831AAA341B9A322C" masterDocTitle="A revision of the &quot; spiny solanums &quot; of Tropical Asia (Solanum, the Leptostemonum Clade, Solanaceae)" masterLastPageNumber="270" masterPageNumber="1" pageNumber="1" updateTime="1654168732655" updateUser="pensoft">
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<mods:title>A revision of the &quot; spiny solanums &quot; of Tropical Asia (Solanum, the Leptostemonum Clade, Solanaceae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Aubriot, Xavier</mods:namePart>
<mods:affiliation>Universite Paris-Saclay, CNRS, AgroParisTech, Ecologie Systematique et Evolution, 91190, Gif-sur-Yvette, France &amp; The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Knapp, Sandra</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7698-3945</mods:nameIdentifier>
<mods:affiliation>The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK</mods:affiliation>
<mods:nameIdentifier type="email">s.knapp@nhm.ac.uk</mods:nameIdentifier>
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<mods:title>PhytoKeys</mods:title>
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<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-06-01</mods:number>
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<mods:detail type="volume">
<mods:number>198</mods:number>
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<mods:url>http://dx.doi.org/10.3897/phytokeys.198.79514</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/phytokeys.198.79514</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2003-198-1</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:49333F06B6534328EC583216A79EEFF9" httpUri="http://treatment.plazi.org/id/49333F06B6534328EC583216A79EEFF9" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
5.
<taxonomicName authority="Dy Phon &amp; Hul, Fl. Cambodge, Laos &amp; Vietnam 35: 16. 2014." authorityName="Dy Phon &amp; Hul, Fl. Cambodge, Laos &amp; Vietnam 35: 16. 2014." class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">Solanum camranhense Dy Phon &amp; Hul, Fl. Cambodge, Laos &amp; Vietnam 35: 16. 2014.</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Diversity of habits and habitats for Solanum subgenus Leptostemonum in tropical Asia A Solanum giganteum, a small tree of the tropical forest understory in India and Sri Lanka (field photograph, unvouchered, India) B S. trilobatum, a scandent species in its typical habitat, along the road in mangrove area (Meeboonya et al. RM 245, Thailand) C S. lasiocarpum, an erect shrub found in disturbed and human inhabited areas where it is often cultivated (Meeboonya et al. RM 272, Thailand) D S. cyanocarphium, a creeper with decumbent stems in forest understory (field photograph, unvouchered, Vietnam) E S. camranhense, a scandent shrub endemic to the coastal dunes of South Vietnam (field photograph, unvouchered, Vietnam) F S. insanum, the wild progenitor of the brinjal eggplant usually found in degraded areas (Meeboonya et al. RM 305, Thailand). Photograph credits: A S. More B, C, F X. Aubriot; D M. Nuraliev E S. Hul." figureDoi="10.3897/phytokeys.198.79514.figure1" httpUri="https://binary.pensoft.net/fig/689694" pageId="0" pageNumber="1">Figs 1E</figureCitation>
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Solanum camranhense Dy Phon &amp; Hul A herbarium specimen (holotype) collected in Vietnam in 1961 (Le Cong Kiet 94, P 00055921) B inflorescence (field photograph, unvouchered, Vietnam) C detail view of the fruits (field photograph, unvouchered, Vietnam). Photograph credits: A CC-BY, Museum national d'Histoire naturelle, Paris B, C S. Hul." figureDoi="10.3897/phytokeys.198.79514.figure10" httpUri="https://binary.pensoft.net/fig/689703" pageId="0" pageNumber="1">, 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">
<typeStatus>Type</typeStatus>
.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation collectingDate="1961-03-07" collectorName="Le Cong Kiet" country="Vietnam" location="My La" municipality="Province Nha Trang" specimenCount="1" stateProvince="Khanh Hoa" typeStatus="holotype">
<collectingCountry name="Vietnam">Vietnam</collectingCountry>
.
<collectingRegion country="Vietnam" name="Khanh Hoa">
<normalizedToken originalValue="Khánh">Khanh</normalizedToken>
<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
</collectingRegion>
: &quot;
<collectingMunicipality>Province Nha Trang</collectingMunicipality>
, dunes littorales de Cam Ranh,
<location LSID="urn:lsid:plazi:treatment:49333F06B6534328EC583216A79EEFF9:EC773ED7554E3AB6D6F6A60E3F7352EB" country="Vietnam" municipality="Province Nha Trang" name="My La" stateProvince="Khanh Hoa">My La</location>
face
<normalizedToken originalValue="à">a</normalizedToken>
la lagune de Bau Ro&quot;,
<collectingDate value="1961-03-07">7 Mar 1961</collectingDate>
,
<emphasis italics="true" pageId="0" pageNumber="1">
<collectorName>
<normalizedToken originalValue="Lê">Le</normalizedToken>
<normalizedToken originalValue="Công">Cong</normalizedToken>
<normalizedToken originalValue="Kiêt">Kiet</normalizedToken>
</collectorName>
94
</emphasis>
(
<typeStatus>holotype</typeStatus>
: P [P00055921]; isotype: P [P00055922])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Scandent shrubs, to 1.5 m tall, unarmed or sparsely prickly. Stems more or less erect, terete, unarmed or occasionally with a few scattered tiny prickles, sparsely to densely stellate-pubescent; prickles to 1.25 mm long, to 1 mm in diameter at the base, straight or curved, acicular to deltate, orange, glabrous; pubescence of mixed sessile and very short-stalked porrect-stellate trichomes, the stalks to 0.1 mm long, the rays 6-9, 0.1-0.4 mm long, the midpoints absent or up to 0.1 mm long; new growth densely stellate-pubescent, light brownish in dry material; bark of older stems brownish grey, glabrescent. Sympodial units plurifoliate, the leaves not geminate. Leaves simple, not lobed, the blades 2-3.5 cm long, 1.5-2.5 cm wide, ca. 1-1.5 times longer than wide, broadly ovate to suborbicular, chartaceous, slightly discolorous, unarmed; adaxial surface densely stellate-pubescent, the stellate trichomes porrect, sessile to stalked, the stalks to 0.1 mm, the rays 6-8, 0.1-0.4 mm long, the midpoints to 0.1 mm; abaxial surface densely stellate-pubescent with trichomes like those of the adaxial surface; major veins 3-5 pairs, drying yellowish light-green; base truncate to subcordate; margins entire or shallowly sinuate; apex obtuse; petioles 0.4-1 cm long, 1/5-1/3 of the leaf blade length, unarmed and densely stellate-pubescent, the pubescence of sessile and short-stalked stellate-trichomes like those of the blades. Inflorescences 1.5-2 cm long, internodal and lateral, sometimes appearing terminal, unbranched, with ca. 4-7 flowers, only 1 or 2 flowers open at any one time, densely stellate-pubescent, with a mix of sessile and short-stalked porrect trichomes like those of the stems, unarmed; peduncle 0.5-2 cm long, unarmed; pedicels 5-9 mm long, ca. 0.5 mm in diameter at the base, ca. 1 mm in diameter at the apex, spreading to erect, unarmed, densely stellate-pubescent with porrect trichomes like those of the inflorescence axes, articulated at the base; pedicel scars irregularly spaced 0.5-4 mm apart. Buds elongate ellipsoid, more or less strongly exserted from the calyx before anthesis. Flowers 5-merous, apparently all perfect. Calyx with the tube ca. 3 mm long, campanulate, the lobes 2-3 mm long, ca. 1 mm wide, deltate and constricting to a short acumen, the acumen 1/4-1/3 the total lobe length, unarmed and densely stellate-pubescent abaxially with porrect-stellate trichomes like those of the pedicels. Corolla 0.8-1 cm in diameter, blue to light purple, stellate, lobed ca. 3/4 of the way to the base, the lobes 5-7 mm long, 1.5-2 mm wide, deltate, spreading at anthesis, densely stellate-pubescent abaxially on parts exposed in bud. Stamens equal; anthers 4-6 mm long, ca. 0.75 mm wide, tapering, orange, glabrous, poricidal at the tips, the pores not lengthening to slits with age; filament tube &lt;0.5 mm long, glabrous; free portion of the filaments 0.5-1 mm long, glabrous. Ovary globose, with minute glandular hairs; style 7-8.5 mm long, slender, curved at the apex, glabrous; stigma capitate, minutely papillate. Fruit a globose berry, several per infructescence, 0.5-0.8 cm in diameter, the pericarp thin and smooth, red when mature, glabrous; fruiting pedicels 0.7-1 cm long, 0.5-1 mm in diameter at the base, 1.6-2 mm in diameter at the apex, woody, erect to spreading, straight to slightly deflexed, unarmed; fruiting calyx lobes not expanding, 1/2-2/3 the length of the mature fruit, broadly deltate, reflexed, unarmed. Seeds 8-10 per berry, 2.5-3 mm long, 2-2.5 mm wide, flattened-reniform, dull yellow, the surface minutely pitted, the testal cells pentagonal in outline. Chromosome number: not known.</paragraph>
<caption doi="10.3897/phytokeys.198.79514.figure10" httpUri="https://binary.pensoft.net/fig/689703" pageId="0" pageNumber="1" start="Figure 10" startId="F10">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 10.</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum camranhense</emphasis>
</taxonomicName>
Dy Phon &amp; Hul
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
herbarium specimen (holotype) collected in Vietnam in 1961 (
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="Lê">Le</normalizedToken>
<normalizedToken originalValue="Công">Cong</normalizedToken>
<normalizedToken originalValue="Kiêt">Kiet</normalizedToken>
94
</emphasis>
, P00055921)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
inflorescence (field photograph, unvouchered, Vietnam)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
detail view of the fruits (field photograph, unvouchered, Vietnam). Photograph credits:
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
CC-BY,
<normalizedToken originalValue="Muséum">Museum</normalizedToken>
national
<normalizedToken originalValue="dHistoire">d'Histoire</normalizedToken>
naturelle, Paris
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
S. Hul.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(Fig.
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of S. camranhense." figureDoi="10.3897/phytokeys.198.79514.figure11" httpUri="https://binary.pensoft.net/fig/689704" pageId="0" pageNumber="1">11</figureCitation>
).
</emphasis>
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum camranhense</emphasis>
</taxonomicName>
is endemic to Vietnam; the few known collections are restricted to
<normalizedToken originalValue="Khánh">Khanh</normalizedToken>
<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
and
<normalizedToken originalValue="Bình">Binh</normalizedToken>
Thuận provinces of South Vietnam.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum camranhense</emphasis>
</taxonomicName>
has only been collected on coastal dunes of stabilized red sands; 15-20 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="common names and uses">
<paragraph pageId="0" pageNumber="1">Common names and uses.</paragraph>
<paragraph pageId="0" pageNumber="1">
Vietnam.
<normalizedToken originalValue="Khánh">Khanh</normalizedToken>
<normalizedToken originalValue="Hòa">Hoa</normalizedToken>
: củ
<normalizedToken originalValue="vè">ve</normalizedToken>
[Vietnamese] (
<emphasis italics="true" pageId="0" pageNumber="1">Chevalier 38932</emphasis>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="preliminary conservation status">
<paragraph pageId="0" pageNumber="1">Preliminary conservation status</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">
(
<bibRefCitation author="Hutchings, A" journalOrPublisher="AGRIVITA Journal of Agricultural Science" pageId="0" pageNumber="1" publicationUrl="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" refId="B105" refString="2019. . http://www.iucnredlist.org/documents/RedListGuidelines.pdf" url="http://www.iucnredlist.org/documents/RedListGuidelines.pdf" year="2019">IUCN 2019</bibRefCitation>
).
</emphasis>
Endangered (EN [B2ab(i,ii,iii)]); EOO (555 km2), AOO 16 km2). Due to the paucity of collections, it is difficult to document the range of
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum camranhense</emphasis>
</taxonomicName>
with confidence. However, its occurrence in the fragmented and anthropogenically disturbed habitats of coastal southern Vietnam (
<bibRefCitation author="Wikramanayake, E" journalOrPublisher="Island Press, London" pageId="0" pageNumber="1" refId="B271" refString="Wikramanayake, E, Dinerstein, E, Loucks, CJ, Olson, DM, Morrison, J, Lamoreux, J, McKnight, M, Hedao, P, 2002. Terrestrial ecoregions of the Indo-Pacific: a conservation assessment. Island Press, London" title="Terrestrial ecoregions of the Indo-Pacific: a conservation assessment." year="2002">Wikramanayake et al. 2002</bibRefCitation>
) suggests it is of conservation concern.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum camranhense</emphasis>
</taxonomicName>
is morphologically similar to the sympatric
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. robinsonii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="robinsonii">
<emphasis italics="true" pageId="0" pageNumber="1">S. robinsonii</emphasis>
</taxonomicName>
, but differs from that species in its scandent (versus erect) habit, the denser pubescence that dries yellowish brown, shorter leaves (2-3.5 cm long versus 3-8 cm long in
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. robinsonii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="robinsonii">
<emphasis italics="true" pageId="0" pageNumber="1">S. robinsonii</emphasis>
</taxonomicName>
), and smaller, more deeply lobed corollas (usually less than 1 cm in diameter and lobed 3/4 of the way to the base versus 1-2 cm in diameter and only lobed halfway to the base in
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. robinsonii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="robinsonii">
<emphasis italics="true" pageId="0" pageNumber="1">S. robinsonii</emphasis>
</taxonomicName>
).
<bibRefCitation author="Hul, S" editor="Aubreville, A" journalOrPublisher="Museum National d'Histoire Naturelle &amp; Royal Botanic Garden Edinburgh, Paris &amp; Edinburgh" pageId="0" pageNumber="1" pagination="1 - 93" refId="B101" refString="Hul, S, Dy Phon, P, 2014. Solanaceae. In: Aubreville, A, Leroy, J, Morat, P, Eds., Flore du Cambodge, du Laos et du Vietnam. Museum National d'Histoire Naturelle &amp; Royal Botanic Garden Edinburgh, Paris &amp; Edinburgh: 1 - 93" title="Solanaceae." volumeTitle="Flore du Cambodge, du Laos et du Vietnam." year="2014">Hul and Dy Phon (2014)</bibRefCitation>
described
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. camranhense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">S. camranhense</emphasis>
</taxonomicName>
as unarmed, but we have seen specimens with a few scattered prickles on the scandent stems. The two species differ in habitat;
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. camranhense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">S. camranhense</emphasis>
</taxonomicName>
is a plant of coastal dunes, while
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. robinsonii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="robinsonii">
<emphasis italics="true" pageId="0" pageNumber="1">S. robinsonii</emphasis>
</taxonomicName>
occurs in coastal forests.
</paragraph>
<caption doi="10.3897/phytokeys.198.79514.figure11" httpUri="https://binary.pensoft.net/fig/689704" pageId="0" pageNumber="1" start="Figure 11" startId="F11">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 11.</emphasis>
Distribution of
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. camranhense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">S. camranhense</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Solanum camranhense" order="Solanales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="camranhense">
<emphasis italics="true" pageId="0" pageNumber="1">Solanum camranhense</emphasis>
</taxonomicName>
is a member of the clade '
<emphasis italics="true" pageId="0" pageNumber="1">
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. camranhense" order="Solanales" pageId="0" pageNumber="1" rank="species" species="camranhense">S. camranhense</taxonomicName>
and relatives
</emphasis>
' of
<bibRefCitation DOI="https://doi.org/10.1111/boj.12412" author="Aubriot, X" journalOrPublisher="Botanical Journal of the Linnean Society" pageId="0" pageNumber="1" pagination="199 - 223" refId="B8" refString="Aubriot, X, Singh, P, Knapp, S, 2016a. Tropical Asian species show the Old World clade of &quot;spiny solanums&quot; (subgenus Leptostemonum pro parte: Solanaceae) is not monophyletic. Botanical Journal of the Linnean Society 181 (2): 199 - 223, DOI: https://doi.org/10.1111/boj.12412" title="Tropical Asian species show the Old World clade of &quot; spiny solanums &quot; (subgenus Leptostemonum pro parte: Solanaceae) is not monophyletic." url="https://doi.org/10.1111/boj.12412" volume="181" year="2016 a">Aubriot et al. (2016a)</bibRefCitation>
with
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. nienkui" order="Solanales" pageId="0" pageNumber="1" rank="species" species="nienkui">
<emphasis italics="true" pageId="0" pageNumber="1">S. nienkui</emphasis>
</taxonomicName>
and
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. putii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="putii">
<emphasis italics="true" pageId="0" pageNumber="1">S. putii</emphasis>
</taxonomicName>
;
<taxonomicName family="Solanaceae" genus="S." kingdom="Plantae" lsidName="S. robinsonii" order="Solanales" pageId="0" pageNumber="1" rank="species" species="robinsonii">
<emphasis italics="true" pageId="0" pageNumber="1">S. robinsonii</emphasis>
</taxonomicName>
has not been included in any molecular analyses to date, but we expect it to be closely related to these taxa.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph pageId="0" pageNumber="1">See Suppl. materials 1-3.</paragraph>
</subSubSection>
</treatment>
</document>