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<document id="62E5BC5586D6EFF94C79675768ABD477" ID-CLB-Dataset="306530" ID-DOI="10.11646/zootaxa.5550.1.32" ID-GBIF-Dataset="2a3f736b-5f9f-4f70-a3a3-6ef61142fa1d" ID-ISSN="1175-5326" ID-Zenodo-Dep="14389333" ID-ZooBank="CA43A365-5B9C-44B8-B7FC-BCFEE44150C1" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="valdenar" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="valdenar" checkinTime="1733923103320" checkinUser="plazi" docAuthor="Chen, I-Shiung &amp; Lai, Hong-Thih" docDate="2024" docId="03C4DE500B1FFFF5B993E003FD84BC33" docLanguage="en" docName="zootaxa.5550.1.32.pdf" docOrigin="Zootaxa 5550 (1)" docSource="https://doi.org/10.11646/zootaxa.5550.1.32" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oryzias chenglongensis Chen &amp; Lai, 2024, new species" docType="treatment" docVersion="3" lastPageNumber="326" masterDocId="FFFDA6280B1EFFF3B904E364FFE0B86A" masterDocTitle="A new ricefish of genus Oryzias (Teleostei: Adrianichthyidae) from western Taiwan" masterLastPageNumber="327" masterPageNumber="320" pageNumber="321" updateTime="1734370632196" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B2F0A2DC14DF0670DA90CF7EA2316954">A new ricefish of genus Oryzias (Teleostei: Adrianichthyidae) from western Taiwan</mods:title>
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<mods:namePart id="7F4ACC4ABCD85397DC5F92313E8958F0">Chen, I-Shiung</mods:namePart>
<mods:affiliation id="1E174787A33D0C5A3B1E17D693065965">Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. &amp; Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C.</mods:affiliation>
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<mods:namePart id="3A3C484F118B654DB007E1604B12B207">Lai, Hong-Thih</mods:namePart>
<mods:affiliation id="098B94E294C053DF85DF4E1389907B6A">Department and Graduate Institute of Aquatic Biosciences, National Charyi University, No. 300, Xuefu Rd., Luliao Village, Charyi City, 600355, Taiwan, R. O. C.</mods:affiliation>
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<mods:date id="8997F40C312313AF0070BA9E180B3498">2024</mods:date>
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<mods:number id="6164AA52B475A4BDE43F199BEFAE21D4">2024-12-10</mods:number>
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<treatment id="03C4DE500B1FFFF5B993E003FD84BC33" ID-DOI="http://doi.org/10.5281/zenodo.14502338" ID-Zenodo-Dep="14502338" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF5B993E003FD84BC33" httpUri="http://treatment.plazi.org/id/03C4DE500B1FFFF5B993E003FD84BC33" lastPageId="6" lastPageNumber="326" pageId="1" pageNumber="321">
<subSubSection id="C3773CCD0B1FFFF2B993E003FDACBBEA" box="[151,588,870,897]" pageId="1" pageNumber="321" type="nomenclature">
<paragraph id="8BD26F460B1FFFF2B993E003FDACBBEA" blockId="1.[151,588,870,933]" box="[151,588,870,897]" pageId="1" pageNumber="321">
<heading id="D09AD82A0B1FFFF2B993E003FDACBBEA" bold="true" box="[151,588,870,897]" fontSize="11" level="1" pageId="1" pageNumber="321" reason="1">
<emphasis id="B919B3540B1FFFF2B993E003FDACBBEA" bold="true" box="[151,588,870,897]" pageId="1" pageNumber="321">
<taxonomicName id="4C6D14C50B1FFFF2B993E003FE55BBEB" authorityName="Chen &amp; Lai" authorityYear="2024" box="[151,437,870,897]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1FFFF2B993E003FE55BBEB" bold="true" box="[151,437,870,897]" italics="true" pageId="1" pageNumber="321">Oryzias chenglongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A0E2F0B1FFFF2B8B9E003FDACBBEA" box="[445,588,870,897]" pageId="1" pageNumber="321" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1FFFF2B993E0EFFEF2BBCF" box="[151,274,907,933]" pageId="1" pageNumber="321" type="description">
<paragraph id="8BD26F460B1FFFF2B993E0EFFEF2BBCF" blockId="1.[151,588,870,933]" box="[151,274,907,933]" pageId="1" pageNumber="321">
(
<figureCitation id="135673C30B1FFFF2B99BE0EFFEEABBCF" box="[159,266,907,933]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1944,1968]" captionStartId-1="2.[151,250,1169,1193]" captionStartId-2="4.[151,250,1588,1612]" captionStartId-3="5.[151,250,1548,1572]" captionTargetBox-0="[151,1436,957,1921]" captionTargetBox-1="[151,1436,181,1145]" captionTargetBox-2="[151,1436,742,1565]" captionTargetBox-3="[151,1436,181,1524]" captionTargetId-0="figure-226@1.[151,1436,957,1921]" captionTargetId-1="figure-19@2.[151,1436,181,1145]" captionTargetId-2="figure-95@4.[151,1436,728,1565]" captionTargetId-3="figure-25@5.[151,1436,179,872]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Oryzias chenglongensis n. sp., male, NTOUP-2018-06-310; 24.8 mm SL, Chenglong wetland, Yunlin County, Taiwan." captionText-1="FIGURE 2. Oryzias chenglongensis n. sp., female, NTOUP-2018 -06-311, 26.6 mm SL, Chenglong wetland, Yunlin County, Taiwan." captionText-2="FIGURE 3. Head profile and pigmentation pattern of Oryzias chenglongensis, holotype." captionText-3="FIGURE 4. Dorsal-fin pigmentation pattern of Oryzias chenglongensis, holotype (above) and Oryzias curvinotus (below)." figureDoi-0="http://doi.org/10.5281/zenodo.14389335" figureDoi-1="http://doi.org/10.5281/zenodo.14389337" figureDoi-2="http://doi.org/10.5281/zenodo.14389343" figureDoi-3="http://doi.org/10.5281/zenodo.14389345" httpUri-0="https://zenodo.org/record/14389335/files/figure.png" httpUri-1="https://zenodo.org/record/14389337/files/figure.png" httpUri-2="https://zenodo.org/record/14389343/files/figure.png" httpUri-3="https://zenodo.org/record/14389345/files/figure.png" pageId="1" pageNumber="321">Figs. 1–4</figureCitation>
)
</paragraph>
</subSubSection>
<caption id="DF123FCE0B1FFFF2B993E4FCFF09BFBE" ID-DOI="http://doi.org/10.5281/zenodo.14389335" ID-Zenodo-Dep="14389335" httpUri="https://zenodo.org/record/14389335/files/figure.png" pageId="1" pageNumber="321" startId="1.[151,250,1944,1968]" targetBox="[151,1436,957,1921]" targetPageId="1" targetType="figure">
<paragraph id="8BD26F460B1FFFF2B993E4FCFF09BFBE" blockId="1.[151,1437,1944,2004]" pageId="1" pageNumber="321">
<emphasis id="B919B3540B1FFFF2B993E4FCFEF6BFDA" bold="true" box="[151,278,1944,1968]" pageId="1" pageNumber="321">FIGURE 1.</emphasis>
<taxonomicName id="4C6D14C50B1FFFF2B81BE4FDFDF3BFDA" authorityName="Chen &amp; Lai" authorityYear="2024" box="[287,531,1944,1968]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1FFFF2B81BE4FDFDF3BFDA" box="[287,531,1944,1968]" italics="true" pageId="1" pageNumber="321">Oryzias chenglongensis</emphasis>
</taxonomicName>
<emphasis id="B919B3540B1FFFF2BB18E4FCFDB8BFDA" bold="true" box="[540,600,1944,1968]" pageId="1" pageNumber="321">
<taxonomicNameLabel id="A22A0E2F0B1FFFF2BB18E4FCFDB8BFDA" box="[540,600,1944,1968]" pageId="1" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, NTOUP-2018-06-310; 24.8 mm SL, Chenglong wetland, Yunlin County, Taiwan.
</paragraph>
</caption>
<caption id="DF123FCE0B1CFFF1B993E7F5FF09BCA7" ID-DOI="http://doi.org/10.5281/zenodo.14389337" ID-Zenodo-Dep="14389337" httpUri="https://zenodo.org/record/14389337/files/figure.png" pageId="2" pageNumber="322" startId="2.[151,250,1169,1193]" targetBox="[151,1436,181,1145]" targetPageId="2" targetType="figure">
<paragraph id="8BD26F460B1CFFF1B993E7F5FF09BCA7" blockId="2.[151,1437,1169,1229]" pageId="2" pageNumber="322">
<emphasis id="B919B3540B1CFFF1B993E7F5FEF4BCC3" bold="true" box="[151,276,1169,1193]" pageId="2" pageNumber="322">FIGURE 2.</emphasis>
<taxonomicName id="4C6D14C50B1CFFF1B81FE7F5FDEDBCC3" authorityName="Chen &amp; Lai" authorityYear="2024" box="[283,525,1169,1193]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="2" pageNumber="322" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1CFFF1B81FE7F5FDEDBCC3" box="[283,525,1169,1193]" italics="true" pageId="2" pageNumber="322">Oryzias chenglongensis</emphasis>
</taxonomicName>
<emphasis id="B919B3540B1CFFF1BB11E7F5FDAFBCC3" bold="true" box="[533,591,1169,1193]" pageId="2" pageNumber="322">
<taxonomicNameLabel id="A22A0E2F0B1CFFF1BB11E7F5FDAFBCC3" box="[533,591,1169,1193]" pageId="2" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, NTOUP-2018 -06-311, 26.6 mm SL, Chenglong wetland, Yunlin County, Taiwan.
</paragraph>
</caption>
<subSubSection id="C3773CCD0B1CFFF1B9C3E67BFDAEBDA3" pageId="2" pageNumber="322" type="materials_examined">
<paragraph id="8BD26F460B1CFFF1B9C3E67BFF38BD37" blockId="2.[151,1436,1311,1481]" pageId="2" pageNumber="322">
<materialsCitation id="3B05651B0B1CFFF1B9C3E67BFF38BD37" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="1" stateProvince="Yunlin" typeStatus="holotype">
<emphasis id="B919B3540B1CFFF1B9C3E67BFED4BD53" bold="true" box="[199,308,1311,1337]" pageId="2" pageNumber="322">
<typeStatus id="54D6D1E40B1CFFF1B9C3E67BFED4BD53" box="[199,308,1311,1337]" pageId="2" pageNumber="322" type="holotype">Holotype</typeStatus>
</emphasis>
.—NTOUP-2018-06-310;
<quantity id="4C95C2A30B1CFFF1BB59E67BFD21BD50" box="[605,705,1311,1338]" metricMagnitude="-2" metricUnit="m" metricValue="2.48" pageId="2" pageNumber="322" unit="mm" value="24.8">24.8 mm</quantity>
SL, coll. I-S.
<location id="8EB2399D0B1CFFF1BA64E67BFC7FBD53" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF5B993E003FD84BC33:8EB2399D0B1CFFF1BA64E67BFC7FBD53" box="[864,927,1311,1337]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BAAEE67BFBC5BD53" box="[938,1061,1311,1337]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1BD97E67BFAD9BD53" box="[1171,1337,1311,1337]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1BC47E67BFA78BD53" box="[1347,1432,1311,1337]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC.
</materialsCitation>
</paragraph>
<paragraph id="8BD26F460B1CFFF1B9C3E603FDAEBDA3" blockId="2.[151,1436,1311,1481]" pageId="2" pageNumber="322">
<materialsCitation id="3B05651B0B1CFFF1B9C3E603FE04BDCF" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="3" stateProvince="Yunlin" typeStatus="paratype">
<emphasis id="B919B3540B1CFFF1B9C3E603FEDFBDEB" bold="true" box="[199,319,1383,1409]" pageId="2" pageNumber="322">
<typeStatus id="54D6D1E40B1CFFF1B9C3E603FEDFBDEB" box="[199,319,1383,1409]" pageId="2" pageNumber="322" type="paratype">Paratypes</typeStatus>
</emphasis>
.—NTOUP-2018-06-311;
<specimenCount id="9D6BA4CF0B1CFFF1BB6FE603FD1EBDEB" box="[619,766,1383,1409]" count="3" pageId="2" pageNumber="322" type="generic">3 specimens</specimenCount>
,
<quantity id="4C95C2A30B1CFFF1BA09E603FC55BDE8" box="[781,949,1383,1410]" metricMagnitude="-2" metricUnit="m" metricValue="2.505" metricValueMax="2.66" metricValueMin="2.35" pageId="2" pageNumber="322" unit="mm" value="25.05" valueMax="26.6" valueMin="23.5">23.5–26.6 mm</quantity>
SL, coll. I-S.
<location id="8EB2399D0B1CFFF1BD63E603FB46BDEB" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF5B993E003FD84BC33:8EB2399D0B1CFFF1BD63E603FB46BDEB" box="[1127,1190,1383,1409]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BDB2E603FAD1BDEB" box="[1206,1329,1383,1409]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1B993E6EFFEDDBDCF" box="[151,317,1419,1445]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1B843E6EFFE7CBDCF" box="[327,412,1419,1445]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC
</materialsCitation>
.
<materialsCitation id="3B05651B0B1CFFF1B8EAE6EFFDAABDA3" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="20" stateProvince="Yunlin" typeStatus="paratype">
NTOUP-2018-02-312,
<quantity id="4C95C2A30B1CFFF1BBF7E6EFFC75BDCC" box="[755,917,1419,1446]" metricMagnitude="-2" metricUnit="m" metricValue="1.985" metricValueMax="2.45" metricValueMin="1.52" pageId="2" pageNumber="322" unit="mm" value="19.85" valueMax="24.5" valueMin="15.2">15.2–24.5 mm</quantity>
SL,
<specimenCount id="9D6BA4CF0B1CFFF1BACEE6EFFB85BDCF" box="[970,1125,1419,1445]" count="20" pageId="2" pageNumber="322" type="generic">20 specimens</specimenCount>
, coll. I-S.
<location id="8EB2399D0B1CFFF1BDDCE6EFFAF7BDCF" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF5B993E003FD84BC33:8EB2399D0B1CFFF1BDDCE6EFFAF7BDCF" box="[1240,1303,1419,1445]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BC25E6EFFA7CBDCF" box="[1313,1436,1419,1445]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1B9F9E6CBFE42BDA3" box="[253,418,1455,1481]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1B8A9E6CBFDE2BDA3" box="[429,514,1455,1481]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1CFFF1B993E57FFCF8BFF7" pageId="2" pageNumber="322" type="diagnosis">
<paragraph id="8BD26F460B1CFFF1B993E57FFEF0BE5F" blockId="2.[151,272,1563,1589]" box="[151,272,1563,1589]" pageId="2" pageNumber="322">
<emphasis id="B919B3540B1CFFF1B993E57FFEF0BE5F" bold="true" box="[151,272,1563,1589]" pageId="2" pageNumber="322">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD26F460B1CFFF1B993E507FCF8BFF7" blockId="2.[151,1437,1635,1950]" pageId="2" pageNumber="322">
The new species,
<taxonomicName id="4C6D14C50B1CFFF1B864E507FD8BBE17" authorityName="Chen &amp; Lai" authorityYear="2024" box="[352,619,1635,1661]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="2" pageNumber="322" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1CFFF1B864E507FD8BBE17" box="[352,619,1635,1661]" italics="true" pageId="2" pageNumber="322">Oryzias chenglongensis</emphasis>
</taxonomicName>
can be well distinguished from other congeners by the following unique combination of features: (1) fin-ray counts: dorsal-fin rays 6 (6 seen in
<typeStatus id="54D6D1E40B1CFFF1BAB2E5E3FBFDBECB" box="[950,1053,1671,1697]" pageId="2" pageNumber="322" type="holotype">holotype</typeStatus>
); anal-fin rays 19–20 (19); pelvic-fin rays 6 (6); pectoral-fin rays 10 (10); principle caudal-fin rays i,4/5,i (i,4/5,i); (2) body compressed laterally with body depth at pelvic origin 24.0–24.3 (24.3), body depth at anal fin origin 15.5–20.4 (20.4); (3) the dorsal-fin origin inserted almost at the vertical of 14
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anal-fin rays, membrane between dorsal-fin 5
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and 6
<superScript id="7C18C20E0B1CFFF1BD9AE597FB4BBF6B" attach="left" box="[1182,1195,1779,1793]" fontSize="6" pageId="2" pageNumber="322">th</superScript>
rays without distinct notch in adult male; and (4) body translucent, light yellowish green or brown, scattered with minute melanophores, a narrowly dotted, longitudinal black line along the middle forward to vertical above pelvic fin; T-shape black mark of caudal fin base in male; operculum and pre-pectoral region slivery, no any silvery scales on lateral body, lip color grayish or creamy yellow and eye bluish silvery dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1DFFF6B993E3F3FAD1BF02" lastPageId="5" lastPageNumber="325" pageId="3" pageNumber="323" type="description">
<paragraph id="8BD26F460B1DFFF0B993E3F3FEC8B8DB" blockId="3.[151,296,151,177]" box="[151,296,151,177]" pageId="3" pageNumber="323">
<emphasis id="B919B3540B1DFFF0B993E3F3FEC8B8DB" bold="true" box="[151,296,151,177]" pageId="3" pageNumber="323">Description</emphasis>
</paragraph>
<paragraph id="8BD26F460B1DFFF0B993E3BBFE35B92B" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Body proportions as their detailed morphometry were summarized in
<tableCitation id="C6EF5AFD0B1DFFF0BA9DE3BBFC0CB893" box="[921,1004,223,249]" captionStart="TABLE 1" captionStartId="3.[152,244,1125,1149]" captionTargetPageId="3" captionText="TABLE 1. Morphometry of Oryzias chenglongensis n. sp. from Taiwan." httpUri="http://table.plazi.org/id/DF123FCE0B1DFFF0B99CE701FC44BC17" pageId="3" pageNumber="323" tableUuid="DF123FCE0B1DFFF0B99CE701FC44BC17">Table 1</tableCitation>
. Dorsal-fin rays 6 (6 seen in
<typeStatus id="54D6D1E40B1DFFF0BC2DE3BBFA70B893" box="[1321,1424,223,249]" pageId="3" pageNumber="323" type="holotype">holotype</typeStatus>
); anal-fin rays 19–20 (19); pelvic-fin rays 6 (6); pectoral-fin rays 10 (10); principle caudal-fin rays i,4/5,i (i,4/5,i), and vertebral counts 29–30 (30).
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E22FFEE8BAEF" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Small species, with maximum size of available specimens examined as
<quantity id="4C95C2A30B1DFFF0BD02E22FFB8CB90C" box="[1030,1132,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="3" pageNumber="323" unit="mm" value="26.6">26.6 mm</quantity>
SL. Head moderate large, with head length 22.8–24.5 (22.8) depressed dorsoventrally; eye very large, eye diameter 34.8–41.3 (39.9); snout very short, snout length 22.8–25.9 (22.8). Body compressed laterally, robust anteriorly and slender posteriorly, body depth at pelvic origin 24.0–24.3 (24.3), body depth at anal fin origin 15.5–20.4 (20.4). No greatly pronounced abdominal concavity between pelvic fins and anterior part of anal fin in female. Mouth terminal, up-turned; tip of upper lip about the horizontal line crossing the upper margin of pupil, lower jaw projecting forward beyond upper jaw, each jaws in having minute conical teeth; lips slightly fleshy. Dorsal body profile relatively straight from head to mid-dorsum and curving from mid-dorsum to dorsal-fin origin; ventral profile as a gentle arch from head to anal-fin origin.
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E1EBFAADBAA4" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">Dorsal profile of head concave. Cheek very limited by large eye. Orbit not projecting toward the dorsal profile. Gill slit opening to superior margin of pectoral-fin base; isthmus located inferior to posterior margin of eye.</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E1B3FEACBB53" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">Scales relatively large and deciduous, cycloid. No scales on rostral zone. Cephalic lateral-line system absent on dorsal, lateral, jugular of head. Nostril opens upper jaw posterosuperior, and lachrymal in front of eye opens near anterior margin.</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E027FD4CBC7B" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Dorsal fin situated for posteriorly, the dorsal-fin origin inserted almost at the vertical of 14
<superScript id="7C18C20E0B1DFFF0BDEAE027FB1BBB3B" attach="left" box="[1262,1275,835,849]" fontSize="6" pageId="3" pageNumber="323">th</superScript>
anal-fin rays. Membrane between dorsal-fin 5
<superScript id="7C18C20E0B1DFFF0BB00E003FDF1BB1F" attach="left" box="[516,529,871,885]" fontSize="6" pageId="3" pageNumber="323">th</superScript>
and 6
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rays without distinct notch in adult male. Dorsal fin rays may elongate with growth in male. Pelvic fin abdominal. Anal fin base rather long, 28.3–30.1 (29.8); membrane margin between each anal-fin ray indented in male, somewhat and slightly concave in female. Pectoral fin rays expended medially. Last pelvic-fin ray connected along less than one-half its length to body forming connecting membrane. Caudal fin truncate, median to posterior 1/3 part branched.
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E77FFABBBC5F" blockId="3.[151,1437,223,1078]" box="[199,1371,1051,1078]" pageId="3" pageNumber="323">Males with a short, tubular urogenital papilla; female with an enlarged bilobed fleshy, urogenital papilla.</paragraph>
<caption id="DF123FCE0B1DFFF0B99CE701FC44BC17" ID-Table-UUID="DF123FCE0B1DFFF0B99CE701FC44BC17" box="[152,932,1125,1149]" httpUri="http://table.plazi.org/id/DF123FCE0B1DFFF0B99CE701FC44BC17" pageId="3" pageNumber="323" startId="3.[152,244,1125,1149]" targetBox="[163,1310,1165,1954]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="8BD26F460B1DFFF0B99CE701FC44BC17" blockId="3.[152,932,1125,1149]" box="[152,932,1125,1149]" pageId="3" pageNumber="323">
<emphasis id="B919B3540B1DFFF0B99CE701FEEFBC17" bold="true" box="[152,271,1125,1149]" pageId="3" pageNumber="323">TABLE 1.</emphasis>
Morphometry of
<taxonomicName id="4C6D14C50B1DFFF0B8C9E701FD2BBC17" authorityName="Chen &amp; Lai" authorityYear="2024" box="[461,715,1125,1149]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="3" pageNumber="323" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1DFFF0B8C9E701FD2BBC17" box="[461,715,1125,1149]" italics="true" pageId="3" pageNumber="323">Oryzias chenglongensis</emphasis>
</taxonomicName>
<emphasis id="B919B3540B1DFFF0BBD6E701FCEEBC17" bold="true" box="[722,782,1125,1149]" pageId="3" pageNumber="323">
<taxonomicNameLabel id="A22A0E2F0B1DFFF0BBD6E701FCEEBC17" box="[722,782,1125,1149]" pageId="3" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Taiwan.
</paragraph>
</caption>
<paragraph id="8BD26F460B1DFFF0BBFCE7E9FAE6BFC8" pageId="3" pageNumber="323">
<table id="F96D9DE60B1D000CB9A7E7E9FAFEBFC8" box="[163,1310,1165,1954]" colsContinueIn="4.[163,1310,202,669]" gridcols="4" gridrows="20" pageId="3" pageNumber="323">
<tr id="355D6D040B1D000CB9A7E7E9FAFEBCCC" box="[163,1310,1165,1190]" gridrow="0" pageId="3" pageNumber="323" rowspan-0="1">
<th id="768C04780B1D000CBBFCE7E9FCB6BCCC" box="[760,854,1165,1190]" gridcol="1" gridrow="0" pageId="3" pageNumber="323">Holotype</th>
<th id="768C04780B1D000CBADAE7E9FBD7BCCC" box="[990,1079,1165,1190]" gridcol="2" gridrow="0" pageId="3" pageNumber="323">Paratype</th>
<th id="768C04780B1D000CBDC1E7E9FAFEBCCC" box="[1221,1310,1165,1190]" gridcol="3" gridrow="0" pageId="3" pageNumber="323">Paratype</th>
</tr>
<tr id="355D6D040B1D000CB9A7E7D1FAFEBCA7" box="[163,1310,1205,1229]" gridrow="1" pageId="3" pageNumber="323" rowspan-0="1">
<td id="768C04780B1D000CBBFCE7D1FCB6BCA7" box="[760,854,1205,1229]" gridcol="1" gridrow="1" pageId="3" pageNumber="323">Male</td>
<td id="768C04780B1D000CBADAE7D1FBD7BCA7" box="[990,1079,1205,1229]" gridcol="2" gridrow="1" pageId="3" pageNumber="323">Female</td>
<td id="768C04780B1D000CBDC1E7D1FAFEBCA7" box="[1221,1310,1205,1229]" gridcol="3" gridrow="1" pageId="3" pageNumber="323">Female</td>
</tr>
<tr id="355D6D040B1D000CB9A7E7BAFAFEBC9C" box="[163,1310,1246,1270]" gridrow="2" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E7BAFD70BC9C" box="[163,656,1246,1270]" gridcol="0" gridrow="2" pageId="3" pageNumber="323">Standard length (mm)</th>
<td id="768C04780B1D000CBBFCE7BAFCB6BC9C" box="[760,854,1246,1270]" gridcol="1" gridrow="2" pageId="3" pageNumber="323">24.8</td>
<td id="768C04780B1D000CBADAE7BAFBD7BC9C" box="[990,1079,1246,1270]" gridcol="2" gridrow="2" pageId="3" pageNumber="323">26.6</td>
<td id="768C04780B1D000CBDC1E7BAFAFEBC9C" box="[1221,1310,1246,1270]" gridcol="3" gridrow="2" pageId="3" pageNumber="323">23.8</td>
</tr>
<tr id="355D6D040B1D000CB9A7E662FAFEBD74" box="[163,1310,1286,1310]" gridrow="3" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E662FAFEBD74" box="[163,1310,1286,1310]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="3" pageNumber="323">% in Standard length</th>
</tr>
<tr id="355D6D040B1D000CB9A7E64AFAFEBD2C" box="[163,1310,1326,1350]" gridrow="4" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E64AFD70BD2C" box="[163,656,1326,1350]" gridcol="0" gridrow="4" pageId="3" pageNumber="323">Body depth at pelvic fin origin</th>
<td id="768C04780B1D000CBBFCE64AFCB6BD2C" box="[760,854,1326,1350]" gridcol="1" gridrow="4" pageId="3" pageNumber="323">24.3%</td>
<td id="768C04780B1D000CBADAE64AFBD7BD2C" box="[990,1079,1326,1350]" gridcol="2" gridrow="4" pageId="3" pageNumber="323">24.0%</td>
<td id="768C04780B1D000CBDC1E64AFAFEBD2C" box="[1221,1310,1326,1350]" gridcol="3" gridrow="4" pageId="3" pageNumber="323">24.2%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E632FAFEBD04" box="[163,1310,1366,1390]" gridrow="5" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E632FD70BD04" box="[163,656,1366,1390]" gridcol="0" gridrow="5" pageId="3" pageNumber="323">Body depth at anal fin origin</th>
<td id="768C04780B1D000CBBFCE632FCB6BD04" box="[760,854,1366,1390]" gridcol="1" gridrow="5" pageId="3" pageNumber="323">20.4%</td>
<td id="768C04780B1D000CBADAE632FBD7BD04" box="[990,1079,1366,1390]" gridcol="2" gridrow="5" pageId="3" pageNumber="323">19.2%</td>
<td id="768C04780B1D000CBDC1E632FAFEBD04" box="[1221,1310,1366,1390]" gridcol="3" gridrow="5" pageId="3" pageNumber="323">15.5%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E61AFAFEBDFD" box="[163,1310,1406,1431]" gridrow="6" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E61AFD70BDFD" box="[163,656,1406,1431]" gridcol="0" gridrow="6" pageId="3" pageNumber="323">Body width</th>
<td id="768C04780B1D000CBBFCE61AFCB6BDFD" box="[760,854,1406,1431]" gridcol="1" gridrow="6" pageId="3" pageNumber="323">16.2%</td>
<td id="768C04780B1D000CBADAE61AFBD7BDFD" box="[990,1079,1406,1431]" gridcol="2" gridrow="6" pageId="3" pageNumber="323">16.6%</td>
<td id="768C04780B1D000CBDC1E61AFAFEBDFD" box="[1221,1310,1406,1431]" gridcol="3" gridrow="6" pageId="3" pageNumber="323">18.3%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E6C3FAFEBDD5" box="[163,1310,1447,1471]" gridrow="7" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E6C3FD70BDD5" box="[163,656,1447,1471]" gridcol="0" gridrow="7" pageId="3" pageNumber="323">Body depth</th>
<td id="768C04780B1D000CBBFCE6C3FCB6BDD5" box="[760,854,1447,1471]" gridcol="1" gridrow="7" pageId="3" pageNumber="323">22.8%</td>
<td id="768C04780B1D000CBADAE6C3FBD7BDD5" box="[990,1079,1447,1471]" gridcol="2" gridrow="7" pageId="3" pageNumber="323">24.5%</td>
<td id="768C04780B1D000CBDC1E6C3FAFEBDD5" box="[1221,1310,1447,1471]" gridcol="3" gridrow="7" pageId="3" pageNumber="323">23.8</td>
</tr>
<tr id="355D6D040B1D000CB9A7E6ABFAFEBD8D" box="[163,1310,1487,1511]" gridrow="8" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E6ABFD70BD8D" box="[163,656,1487,1511]" gridcol="0" gridrow="8" pageId="3" pageNumber="323">Caudal peduncle depth</th>
<td id="768C04780B1D000CBBFCE6ABFCB6BD8D" box="[760,854,1487,1511]" gridcol="1" gridrow="8" pageId="3" pageNumber="323">9.1%</td>
<td id="768C04780B1D000CBADAE6ABFBD7BD8D" box="[990,1079,1487,1511]" gridcol="2" gridrow="8" pageId="3" pageNumber="323">8.8%</td>
<td id="768C04780B1D000CBDC1E6ABFAFEBD8D" box="[1221,1310,1487,1511]" gridcol="3" gridrow="8" pageId="3" pageNumber="323">8.2%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E693FAFEBE65" box="[163,1310,1527,1551]" gridrow="9" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E693FD70BE65" box="[163,656,1527,1551]" gridcol="0" gridrow="9" pageId="3" pageNumber="323">Caudal peduncle length</th>
<td id="768C04780B1D000CBBFCE693FCB6BE65" box="[760,854,1527,1551]" gridcol="1" gridrow="9" pageId="3" pageNumber="323">13.9%</td>
<td id="768C04780B1D000CBADAE693FBD7BE65" box="[990,1079,1527,1551]" gridcol="2" gridrow="9" pageId="3" pageNumber="323">13.5%</td>
<td id="768C04780B1D000CBDC1E693FAFEBE65" box="[1221,1310,1527,1551]" gridcol="3" gridrow="9" pageId="3" pageNumber="323">13.8%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E57BFAFEBE52" box="[163,1310,1567,1592]" gridrow="10" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E57BFD70BE52" box="[163,656,1567,1592]" gridcol="0" gridrow="10" pageId="3" pageNumber="323">Height of dorsal fin</th>
<td id="768C04780B1D000CBBFCE57BFCB6BE52" box="[760,854,1567,1592]" gridcol="1" gridrow="10" pageId="3" pageNumber="323">18.8%</td>
<td id="768C04780B1D000CBADAE57BFBD7BE52" box="[990,1079,1567,1592]" gridcol="2" gridrow="10" pageId="3" pageNumber="323">12.7%</td>
<td id="768C04780B1D000CBDC1E57BFAFEBE52" box="[1221,1310,1567,1592]" gridcol="3" gridrow="10" pageId="3" pageNumber="323">15.6%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E52CFAFEBE0A" box="[163,1310,1608,1632]" gridrow="11" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E52CFD70BE0A" box="[163,656,1608,1632]" gridcol="0" gridrow="11" pageId="3" pageNumber="323">Dorsal-fin base</th>
<td id="768C04780B1D000CBBFCE52CFCB6BE0A" box="[760,854,1608,1632]" gridcol="1" gridrow="11" pageId="3" pageNumber="323">8.6%</td>
<td id="768C04780B1D000CBADAE52CFBD7BE0A" box="[990,1079,1608,1632]" gridcol="2" gridrow="11" pageId="3" pageNumber="323">7.6%</td>
<td id="768C04780B1D000CBDC1E52CFAFEBE0A" box="[1221,1310,1608,1632]" gridcol="3" gridrow="11" pageId="3" pageNumber="323">8.7%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E514FAFEBEE2" box="[163,1310,1648,1672]" gridrow="12" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E514FD70BEE2" box="[163,656,1648,1672]" gridcol="0" gridrow="12" pageId="3" pageNumber="323">Height of anal fin</th>
<td id="768C04780B1D000CBBFCE514FCB6BEE2" box="[760,854,1648,1672]" gridcol="1" gridrow="12" pageId="3" pageNumber="323">14.6%</td>
<td id="768C04780B1D000CBADAE514FBD7BEE2" box="[990,1079,1648,1672]" gridcol="2" gridrow="12" pageId="3" pageNumber="323">13.3%</td>
<td id="768C04780B1D000CBDC1E514FAFEBEE2" box="[1221,1310,1648,1672]" gridcol="3" gridrow="12" pageId="3" pageNumber="323">14.3%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E5FCFAFEBEDA" box="[163,1310,1688,1712]" gridrow="13" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E5FCFD70BEDA" box="[163,656,1688,1712]" gridcol="0" gridrow="13" pageId="3" pageNumber="323">Anal-fin base</th>
<td id="768C04780B1D000CBBFCE5FCFCB6BEDA" box="[760,854,1688,1712]" gridcol="1" gridrow="13" pageId="3" pageNumber="323">29.8%</td>
<td id="768C04780B1D000CBADAE5FCFBD7BEDA" box="[990,1079,1688,1712]" gridcol="2" gridrow="13" pageId="3" pageNumber="323">28.3%</td>
<td id="768C04780B1D000CBDC1E5FCFAFEBEDA" box="[1221,1310,1688,1712]" gridcol="3" gridrow="13" pageId="3" pageNumber="323">30.1%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E5A4FAFEBEB3" box="[163,1310,1728,1753]" gridrow="14" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E5A4FD70BEB3" box="[163,656,1728,1753]" gridcol="0" gridrow="14" pageId="3" pageNumber="323">Pelvic fin length</th>
<td id="768C04780B1D000CBBFCE5A4FCB6BEB3" box="[760,854,1728,1753]" gridcol="1" gridrow="14" pageId="3" pageNumber="323">11.9%</td>
<td id="768C04780B1D000CBADAE5A4FBD7BEB3" box="[990,1079,1728,1753]" gridcol="2" gridrow="14" pageId="3" pageNumber="323">13.7%</td>
<td id="768C04780B1D000CBDC1E5A4FAFEBEB3" box="[1221,1310,1728,1753]" gridcol="3" gridrow="14" pageId="3" pageNumber="323">14.2%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E58DFAFEBF6B" box="[163,1310,1769,1793]" gridrow="15" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E58DFD70BF6B" box="[163,656,1769,1793]" gridcol="0" gridrow="15" pageId="3" pageNumber="323">Pectoral-fin lrngth</th>
<td id="768C04780B1D000CBBFCE58DFCB6BF6B" box="[760,854,1769,1793]" gridcol="1" gridrow="15" pageId="3" pageNumber="323">18.8%</td>
<td id="768C04780B1D000CBADAE58DFBD7BF6B" box="[990,1079,1769,1793]" gridcol="2" gridrow="15" pageId="3" pageNumber="323">17.5%</td>
<td id="768C04780B1D000CBDC1E58DFAFEBF6B" box="[1221,1310,1769,1793]" gridcol="3" gridrow="15" pageId="3" pageNumber="323">17.7%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E475FAFEBF43" box="[163,1310,1809,1833]" gridrow="16" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E475FD70BF43" box="[163,656,1809,1833]" gridcol="0" gridrow="16" pageId="3" pageNumber="323">Predorsal length</th>
<td id="768C04780B1D000CBBFCE475FCB6BF43" box="[760,854,1809,1833]" gridcol="1" gridrow="16" pageId="3" pageNumber="323">79.2%</td>
<td id="768C04780B1D000CBADAE475FBD7BF43" box="[990,1079,1809,1833]" gridcol="2" gridrow="16" pageId="3" pageNumber="323">79.2%</td>
<td id="768C04780B1D000CBDC1E475FAFEBF43" box="[1221,1310,1809,1833]" gridcol="3" gridrow="16" pageId="3" pageNumber="323">77.0%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E45DFAFEBF3B" box="[163,1310,1849,1873]" gridrow="17" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E45DFD70BF3B" box="[163,656,1849,1873]" gridcol="0" gridrow="17" pageId="3" pageNumber="323">Preanal length</th>
<td id="768C04780B1D000CBBFCE45DFCB6BF3B" box="[760,854,1849,1873]" gridcol="1" gridrow="17" pageId="3" pageNumber="323">59.0%</td>
<td id="768C04780B1D000CBADAE45DFBD7BF3B" box="[990,1079,1849,1873]" gridcol="2" gridrow="17" pageId="3" pageNumber="323">60.3%</td>
<td id="768C04780B1D000CBDC1E45DFAFEBF3B" box="[1221,1310,1849,1873]" gridcol="3" gridrow="17" pageId="3" pageNumber="323">59.5%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E405FAFEBF10" box="[163,1310,1889,1914]" gridrow="18" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E405FD70BF10" box="[163,656,1889,1914]" gridcol="0" gridrow="18" pageId="3" pageNumber="323">Prepelvic length</th>
<td id="768C04780B1D000CBBFCE405FCB6BF10" box="[760,854,1889,1914]" gridcol="1" gridrow="18" pageId="3" pageNumber="323">44.9%</td>
<td id="768C04780B1D000CBADAE405FBD7BF10" box="[990,1079,1889,1914]" gridcol="2" gridrow="18" pageId="3" pageNumber="323">46.8%</td>
<td id="768C04780B1D000CBDC1E405FAFEBF10" box="[1221,1310,1889,1914]" gridcol="3" gridrow="18" pageId="3" pageNumber="323">45.5%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E4EEFAFEBFC8" box="[163,1310,1930,1954]" gridrow="19" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E4EEFD70BFC8" box="[163,656,1930,1954]" gridcol="0" gridrow="19" pageId="3" pageNumber="323">Length from Pectoral fin basee to pelvic fin base</th>
<td id="768C04780B1D000CBBFCE4EEFCB6BFC8" box="[760,854,1930,1954]" gridcol="1" gridrow="19" pageId="3" pageNumber="323">16.8%</td>
<td id="768C04780B1D000CBADAE4EEFBD7BFC8" box="[990,1079,1930,1954]" gridcol="2" gridrow="19" pageId="3" pageNumber="323">17.2%</td>
<td id="768C04780B1D000CBDC1E4EEFAFEBFC8" box="[1221,1310,1930,1954]" gridcol="3" gridrow="19" pageId="3" pageNumber="323">16.6%</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD26F460B1DFFF0BD5DE4D7FA70BFA0" blockId="3.[1113,1424,1971,1995]" box="[1113,1424,1971,1995]" pageId="3" pageNumber="323">
<tableNote id="768B6EC80B1DFFF0BD5DE4D7FA70BFA0" box="[1113,1424,1971,1995]" pageId="3" pageNumber="323" targetBox="[163,1310,1165,1954]" targetPageId="3">......continued on the next page</tableNote>
</paragraph>
<caption id="DF123FCE0B1AFFF7B9A7E3C6FE71B8D0" box="[163,401,162,186]" isContinuationCaption="true" pageId="4" pageNumber="324" startId="4.[163,247,162,186]" targetBox="[163,1310,202,669]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="8BD26F460B1AFFF7B9A7E3C6FE71B8D0" blockId="4.[163,401,162,186]" box="[163,401,162,186]" pageId="4" pageNumber="324">
<emphasis id="B919B3540B1AFFF7B9A7E3C6FEF0B8D0" bold="true" box="[163,272,162,186]" pageId="4" pageNumber="324">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BD26F460B1AFFF7BBFCE3AEFAE6BAF7" pageId="4" pageNumber="324">
<table id="F96D9DE60B1A000CB9A7E3AEFAFEBAF7" box="[163,1310,202,669]" colsContinueFrom="3.[163,1310,1165,1954]" gridcols="4" gridrows="5" pageId="4" pageNumber="324">
<tr id="355D6D040B1A000CB9A7E3AEFAFEB889" box="[163,1310,202,227]" gridrow="0" pageId="4" pageNumber="324" rowspan-0="1">
<th id="768C04780B1A000CBBFCE3AEFCB6B889" box="[760,854,202,227]" gridcol="1" gridrow="0" pageId="4" pageNumber="324">Holotype</th>
<th id="768C04780B1A000CBADAE3AEFBD7B889" box="[990,1079,202,227]" gridcol="2" gridrow="0" pageId="4" pageNumber="324">Paratype</th>
<th id="768C04780B1A000CBDC1E3AEFAFEB889" box="[1221,1310,202,227]" gridcol="3" gridrow="0" pageId="4" pageNumber="324">Paratype</th>
</tr>
<tr id="355D6D040B1A000CB9A7E396FAFEB960" box="[163,1310,242,266]" gridrow="1" pageId="4" pageNumber="324" rowspan-0="1">
<td id="768C04780B1A000CBBFCE396FCB6B960" box="[760,854,242,266]" gridcol="1" gridrow="1" pageId="4" pageNumber="324">Male</td>
<td id="768C04780B1A000CBADAE396FBD7B960" box="[990,1079,242,266]" gridcol="2" gridrow="1" pageId="4" pageNumber="324">Female</td>
<td id="768C04780B1A000CBDC1E396FAFEB960" box="[1221,1310,242,266]" gridcol="3" gridrow="1" pageId="4" pageNumber="324">Female</td>
</tr>
<tr id="355D6D040B1A000CB9A7E27FFAFEB959" box="[163,1310,283,307]" gridrow="2" pageId="4" pageNumber="324">
<th id="768C04780B1A000CB9A7E27FFD60B959" box="[163,640,283,307]" gridcol="0" gridrow="2" pageId="4" pageNumber="324">Length from dorsal fin basee to caudal fin base</th>
<td id="768C04780B1A000CBBFCE27FFCB6B959" box="[760,854,283,307]" gridcol="1" gridrow="2" pageId="4" pageNumber="324">14.3%</td>
<td id="768C04780B1A000CBADAE27FFBD7B959" box="[990,1079,283,307]" gridcol="2" gridrow="2" pageId="4" pageNumber="324">15.8%</td>
<td id="768C04780B1A000CBDC1E27FFAFEB959" box="[1221,1310,283,307]" gridcol="3" gridrow="2" pageId="4" pageNumber="324">16.9%</td>
</tr>
<tr id="355D6D040B1A000CB9A7E208FAFEBA27" box="[163,1310,364,589]" gridrow="3" pageId="4" pageNumber="324">
<th id="768C04780B1A000CB9A7E208FD60BA27" box="[163,640,364,589]" gridcol="0" gridrow="3" pageId="4" pageNumber="324">% in Head length Head width Eye diameter Postorbital length Snout length Interorbital width</th>
<td id="768C04780B1A000CBBFCE208FCB6BA27" box="[760,854,364,589]" gridcol="1" gridrow="3" pageId="4" pageNumber="324">66.6% 36.9% 39.9% 22.8% 33.1%</td>
<td id="768C04780B1A000CBADAE208FBD7BA27" box="[990,1079,364,589]" gridcol="2" gridrow="3" pageId="4" pageNumber="324">54.8% 35.8% 41.3% 23.9% 35.2%</td>
<td id="768C04780B1A000CBDC1E208FAFEBA27" box="[1221,1310,364,589]" gridcol="3" gridrow="3" pageId="4" pageNumber="324">70.2% 34.8% 40.0% 25.9% 39.1%</td>
</tr>
<tr id="355D6D040B1A000CB9A7E1E1FAFEBAF7" box="[163,1310,645,669]" gridrow="4" pageId="4" pageNumber="324">
<th id="768C04780B1A000CB9A7E1E1FD60BAF7" box="[163,640,645,669]" gridcol="0" gridrow="4" pageId="4" pageNumber="324">Caudal peduncle depth/length</th>
<td id="768C04780B1A000CBBFCE1E1FCB6BAF7" box="[760,854,645,669]" gridcol="1" gridrow="4" pageId="4" pageNumber="324">65.5%</td>
<td id="768C04780B1A000CBADAE1E1FBD7BAF7" box="[990,1079,645,669]" gridcol="2" gridrow="4" pageId="4" pageNumber="324">65.5%</td>
<td id="768C04780B1A000CBDC1E1E1FAFEBAF7" box="[1221,1310,645,669]" gridcol="3" gridrow="4" pageId="4" pageNumber="324">59.4%</td>
</tr>
</table>
</paragraph>
<caption id="DF123FCE0B1AFFF7B993E550FBFBBE26" ID-DOI="http://doi.org/10.5281/zenodo.14389343" ID-Zenodo-Dep="14389343" box="[151,1051,1588,1613]" httpUri="https://zenodo.org/record/14389343/files/figure.png" pageId="4" pageNumber="324" startId="4.[151,250,1588,1612]" targetBox="[151,1436,742,1565]" targetPageId="4" targetType="figure">
<paragraph id="8BD26F460B1AFFF7B993E550FBFBBE26" blockId="4.[151,1051,1588,1613]" box="[151,1051,1588,1613]" pageId="4" pageNumber="324">
<emphasis id="B919B3540B1AFFF7B993E550FEF3BE26" bold="true" box="[151,275,1588,1612]" pageId="4" pageNumber="324">FIGURE 3.</emphasis>
Head profile and pigmentation pattern of
<taxonomicName id="4C6D14C50B1AFFF7BBBAE551FC4FBE27" authorityName="Chen &amp; Lai" authorityYear="2024" box="[702,943,1589,1613]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="4" pageNumber="324" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1AFFF7BBBAE551FC4FBE27" box="[702,943,1589,1613]" italics="true" pageId="4" pageNumber="324">Oryzias chenglongensis</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
<paragraph id="8BD26F460B1AFFF7B993E5FBFE8DBED3" blockId="4.[151,365,1695,1721]" box="[151,365,1695,1721]" pageId="4" pageNumber="324">
<heading id="D09AD82A0B1AFFF7B993E5FBFE8DBED3" bold="true" box="[151,365,1695,1721]" fontSize="11" level="1" pageId="4" pageNumber="324" reason="1">
<emphasis id="B919B3540B1AFFF7B993E5FBFE8DBED3" bold="true" box="[151,365,1695,1721]" pageId="4" pageNumber="324">Coloration in life</emphasis>
</heading>
</paragraph>
<paragraph id="8BD26F460B1AFFF7B993E583FAFBBFFB" blockId="4.[151,1436,1767,1937]" pageId="4" pageNumber="324">Body translucent, light yellowish green or brown, scattered with minute melanophores can be seen alive and preservative, a conspicuous, narrowly dotted, longitudinal black line along the middle forward to vertical above pelvic fin. T-shape black mark of caudal fin base in male. Belly wall, peritoneum slivery, somewhat subrectangular in both sexes. Operculum and pre-pectoral region slivery. No any silvery scales on lateral body. Lip color grayish or somewhat creamy yellow. Eye bluish silvery dorsally, silvery ventrally with some brilliant yellow dots.</paragraph>
<caption id="DF123FCE0B1BFFF6B993E568FA99BE4E" ID-DOI="http://doi.org/10.5281/zenodo.14389345" ID-Zenodo-Dep="14389345" box="[151,1401,1548,1572]" httpUri="https://zenodo.org/record/14389345/files/figure.png" pageId="5" pageNumber="325" startId="5.[151,250,1548,1572]" targetBox="[151,1436,181,1524]" targetPageId="5" targetType="figure">
<paragraph id="8BD26F460B1BFFF6B993E568FA99BE4E" blockId="5.[151,1401,1548,1572]" box="[151,1401,1548,1572]" pageId="5" pageNumber="325">
<emphasis id="B919B3540B1BFFF6B993E568FEF3BE4E" bold="true" box="[151,275,1548,1572]" pageId="5" pageNumber="325">FIGURE 4.</emphasis>
Dorsal-fin pigmentation pattern of
<taxonomicName id="4C6D14C50B1BFFF6BB78E569FC8DBE4E" authorityName="Chen &amp; Lai" authorityYear="2024" box="[636,877,1548,1572]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="5" pageNumber="325" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1BFFF6BB78E569FC8DBE4E" box="[636,877,1548,1572]" italics="true" pageId="5" pageNumber="325">Oryzias chenglongensis</emphasis>
</taxonomicName>
, holotype (above) and
<taxonomicName id="4C6D14C50B1BFFF6BD5AE569FAFEBE4E" baseAuthorityName="Nichols &amp; Pope" baseAuthorityYear="1927" box="[1118,1310,1549,1572]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="5" pageNumber="325" phylum="Chordata" rank="species" species="curvinotus">
<emphasis id="B919B3540B1BFFF6BD5AE569FAFEBE4E" box="[1118,1310,1549,1572]" italics="true" pageId="5" pageNumber="325">Oryzias curvinotus</emphasis>
</taxonomicName>
(below).
</paragraph>
</caption>
<paragraph id="8BD26F460B1BFFF6B9C3E512FE96BF4A" blockId="5.[151,1437,1654,1897]" pageId="5" pageNumber="325">Generally, all fins and belly colored pale white to pale yellow. Caudal fin with yellowish dorsal and ventral submarginal stripes and the stripes darker with melanophores as well as distal yellowish margin in male. Dorsal edge of dorsal and pectoral fins, distal region of pelvic and anal fins with yellow bands. Entire pelvic fin yellowish in male. Pelvic and anal fin with widely scattered tiny melanophores. Caudal fin membrane with about 4 hairline longitudinal stripes.</paragraph>
<paragraph id="8BD26F460B1BFFF6B9C3E44EFAD1BF02" blockId="5.[151,1437,1654,1897]" pageId="5" pageNumber="325">Coloration while specimens in preservative similar above and all brilliant coloration and translucent part entirely faded, body generally light yellowish brown except the melanophores and some larger deep black marks.</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B18FFF5B993E3F3FEA5BA73" pageId="6" pageNumber="326" type="distribution">
<paragraph id="8BD26F460B18FFF5B993E3F3FE33B8DB" blockId="6.[151,467,151,177]" box="[151,467,151,177]" pageId="6" pageNumber="326">
<emphasis id="B919B3540B18FFF5B993E3F3FE33B8DB" bold="true" box="[151,467,151,177]" pageId="6" pageNumber="326">Distribution and habitats</emphasis>
</paragraph>
<paragraph id="8BD26F460B18FFF5B993E3BBFAFAB92B" blockId="6.[151,1437,223,537]" pageId="6" pageNumber="326">
Although the
<typeStatus id="54D6D1E40B18FFF5B834E384FE80B890" box="[304,352,224,250]" pageId="6" pageNumber="326">type</typeStatus>
specimens of
<taxonomicName id="4C6D14C50B18FFF5B8FAE384FCE9B893" authorityName="Chen &amp; Lai" authorityYear="2024" box="[510,777,223,250]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="6" pageNumber="326" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B18FFF5B8FAE384FCE9B893" box="[510,777,223,250]" italics="true" pageId="6" pageNumber="326">Oryzias chenglongensis</emphasis>
</taxonomicName>
eventually described and collected from the
<collectingRegion id="49A9A1A40B18FFF5BDFDE3BBFA7CB893" box="[1273,1436,223,249]" country="Taiwan" name="Yunlin" pageId="6" pageNumber="326">Yunlin County</collectingRegion>
of western
<collectingCountry id="F37A2FD60B18FFF5B812E267FE8BB977" box="[278,363,259,285]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
, it can be also found from Chairyi and
<collectingRegion id="49A9A1A40B18FFF5BA28E267FC98B977" box="[812,888,259,285]" country="Taiwan" name="Tainan" pageId="6" pageNumber="326">Tainan</collectingRegion>
by some photo image evidences of local people without well preserved specimens. It is highly possible to be an endemic species around
<collectingCountry id="F37A2FD60B18FFF5BD72E243FB28B92B" box="[1142,1224,295,321]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
island.
</paragraph>
<paragraph id="8BD26F460B18FFF5B9C3E228FEA5BA73" blockId="6.[151,1437,223,537]" pageId="6" pageNumber="326">
It can be considered that the brackish ricefish might be existed in the wetlands of western
<collectingCountry id="F37A2FD60B18FFF5BDE2E22FFAD8B90F" box="[1254,1336,331,357]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
(at least ranging from
<collectingRegion id="49A9A1A40B18FFF5B831E20BFE60B9E3" box="[309,384,367,393]" country="Taiwan" name="Yunlin" pageId="6" pageNumber="326">Yunlin</collectingRegion>
southward to Taiwan) although the current population may be very low and rare. This new species merely collected from brackish habitat around 20–30 for salinity. The recorded localities from both Chiaryi and
<collectingRegion id="49A9A1A40B18FFF5B9C3E2D3FEF3B9BB" box="[199,275,439,465]" country="Taiwan" name="Tainan" pageId="6" pageNumber="326">Tainan</collectingRegion>
are also the brackish habitats of wetlands and estuaries. In our breeding experiment, this fish can also survive well in fresh water, it might be more widely distributed around the lowland freshwater to brackish region around
<collectingCountry id="F37A2FD60B18FFF5B9EFE29BFEA1BA73" box="[235,321,511,537]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B18FFF5B993E10FFEAEBB7F" pageId="6" pageNumber="326" type="etymology">
<paragraph id="8BD26F460B18FFF5B993E10FFEFEBAEF" blockId="6.[151,286,619,645]" box="[151,286,619,645]" pageId="6" pageNumber="326">
<emphasis id="B919B3540B18FFF5B993E10FFEFEBAEF" bold="true" box="[151,286,619,645]" pageId="6" pageNumber="326">Etymology</emphasis>
</paragraph>
<paragraph id="8BD26F460B18FFF5B993E1D7FEAEBB7F" blockId="6.[151,1436,691,790]" pageId="6" pageNumber="326">
The specific name,
<taxonomicName id="4C6D14C50B18FFF5B882E1D7FDD3BAA7" authorityName="Chen &amp; Lai" authorityYear="2024" box="[390,563,691,717]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="6" pageNumber="326" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B18FFF5B882E1D7FDD3BAA7" box="[390,563,691,717]" italics="true" pageId="6" pageNumber="326">chenglongensis</emphasis>
</taxonomicName>
, is referred to names of the fish
<typeStatus id="54D6D1E40B18FFF5BAD4E1D7FBD2BAA7" box="[976,1074,691,717]" pageId="6" pageNumber="326" type="holotype">holotype</typeStatus>
locality collected “Chonglong wetland”, western
<collectingCountry id="F37A2FD60B18FFF5B874E1B3FE22BA9B" box="[368,450,727,753]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
although its available geographical range may from Yun-lin, Chiar-yi, to Tai-nan of western
<collectingCountry id="F37A2FD60B18FFF5B9F1E19FFEABBB7F" box="[245,331,763,789]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B18FFF5B993E003FD84BC33" pageId="6" pageNumber="326" type="conservation">
<paragraph id="8BD26F460B18FFF5B993E003FEDFBBEB" blockId="6.[151,319,871,897]" box="[151,319,871,897]" pageId="6" pageNumber="326">
<emphasis id="B919B3540B18FFF5B993E003FEDFBBEB" bold="true" box="[151,319,871,897]" pageId="6" pageNumber="326">Conservation</emphasis>
</paragraph>
<paragraph id="8BD26F460B18FFF5B993E0D4FD84BC33" blockId="6.[151,1436,943,1113]" pageId="6" pageNumber="326">
Due the very rare of current new species in wild survey in western
<collectingCountry id="F37A2FD60B18FFF5BA97E0CBFC08BBA3" box="[915,1000,943,969]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
, we only kept very limited individuals into
<typeStatus id="54D6D1E40B18FFF5B9CEE0B0FF1ABB84" box="[202,250,980,1006]" pageId="6" pageNumber="326">type</typeStatus>
series. Its habitat has also faced the great ecological threat from at least two invasive species of American killifishes. Almost other fish individuals of new species have been kept for conservation reason and reproduce their further off spring and save its limited stock at NTOU,
<collectingRegion id="49A9A1A40B18FFF5BBEBE77FFCAEBC5F" box="[751,846,1051,1077]" country="Taiwan" name="Keelung" pageId="6" pageNumber="326">Keelung</collectingRegion>
for avoiding great aquatic ecological threat from this endangered species becoming extinction.
</paragraph>
</subSubSection>
</treatment>
</document>
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