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<document id="20D38825D74703A83CB93310F6FEA379" ID-DOI="10.11646/zootaxa.5550.1.32" ID-ISSN="1175-5326" ID-Zenodo-Dep="14389333" ID-ZooBank="CA43A365-5B9C-44B8-B7FC-BCFEE44150C1" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733923103320" checkinUser="plazi" docAuthor="Chen, I-Shiung &amp; Lai, Hong-Thih" docDate="2024" docId="03C4DE500B1FFFF0B993E003FABBBC5F" docLanguage="en" docName="zootaxa.5550.1.32.pdf" docOrigin="Zootaxa 5550 (1)" docSource="https://doi.org/10.11646/zootaxa.5550.1.32" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oryzias chenglongensis Chen &amp; Lai, 2024, new species" docType="treatment" docVersion="2" lastPageNumber="323" masterDocId="FFFDA6280B1EFFF3B904E364FFE0B86A" masterDocTitle="A new ricefish of genus Oryzias (Teleostei: Adrianichthyidae) from western Taiwan" masterLastPageNumber="327" masterPageNumber="320" pageNumber="321" updateTime="1733924007468" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7CBD0F1D0B8CDA849A2AC0D4AA91456D">A new ricefish of genus Oryzias (Teleostei: Adrianichthyidae) from western Taiwan</mods:title>
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<mods:namePart id="233C057360AA32AF3C96612D2B223315">Chen, I-Shiung</mods:namePart>
<mods:affiliation id="74FBDF7614D268D7ADCB7BE4C61F10E6">Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. &amp; Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C.</mods:affiliation>
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<mods:namePart id="5DDF32DCEB932006C69761339E09140D">Lai, Hong-Thih</mods:namePart>
<mods:affiliation id="53D9C9E7EFA2EADF65C51E419EE372BD">Department and Graduate Institute of Aquatic Biosciences, National Charyi University, No. 300, Xuefu Rd., Luliao Village, Charyi City, 600355, Taiwan, R. O. C.</mods:affiliation>
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<mods:date id="3F7B0F32DD8FAF826F6FC3C7BE74297E">2024</mods:date>
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<treatment id="03C4DE500B1FFFF0B993E003FABBBC5F" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF0B993E003FABBBC5F" httpUri="http://treatment.plazi.org/id/03C4DE500B1FFFF0B993E003FABBBC5F" lastPageId="3" lastPageNumber="323" pageId="1" pageNumber="321">
<subSubSection id="C3773CCD0B1FFFF2B993E003FDACBBEA" box="[151,588,870,897]" pageId="1" pageNumber="321" type="nomenclature">
<paragraph id="8BD26F460B1FFFF2B993E003FDACBBEA" blockId="1.[151,588,870,933]" box="[151,588,870,897]" pageId="1" pageNumber="321">
<heading id="D09AD82A0B1FFFF2B993E003FDACBBEA" bold="true" box="[151,588,870,897]" fontSize="11" level="1" pageId="1" pageNumber="321" reason="1">
<emphasis id="B919B3540B1FFFF2B993E003FDACBBEA" bold="true" box="[151,588,870,897]" pageId="1" pageNumber="321">
<taxonomicName id="4C6D14C50B1FFFF2B993E003FE55BBEB" authorityName="Chen &amp; Lai" authorityYear="2024" box="[151,437,870,897]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1FFFF2B993E003FE55BBEB" bold="true" box="[151,437,870,897]" italics="true" pageId="1" pageNumber="321">Oryzias chenglongensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22A0E2F0B1FFFF2B8B9E003FDACBBEA" box="[445,588,870,897]" pageId="1" pageNumber="321" rank="species">new species</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C3773CCD0B1FFFF2B993E0EFFEF2BBCF" box="[151,274,907,933]" pageId="1" pageNumber="321" type="description">
<paragraph id="8BD26F460B1FFFF2B993E0EFFEF2BBCF" blockId="1.[151,588,870,933]" box="[151,274,907,933]" pageId="1" pageNumber="321">
(
<figureCitation id="135673C30B1FFFF2B99BE0EFFEEABBCF" box="[159,266,907,933]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1944,1968]" captionStartId-1="2.[151,250,1169,1193]" captionStartId-2="4.[151,250,1588,1612]" captionStartId-3="5.[151,250,1548,1572]" captionTargetBox-0="[151,1436,957,1921]" captionTargetBox-1="[151,1436,181,1145]" captionTargetBox-2="[151,1436,742,1565]" captionTargetBox-3="[151,1436,181,1524]" captionTargetId-0="figure-226@1.[151,1436,957,1921]" captionTargetId-1="figure-19@2.[151,1436,181,1145]" captionTargetId-2="figure-95@4.[151,1436,728,1565]" captionTargetId-3="figure-25@5.[151,1436,179,872]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Oryzias chenglongensis n. sp., male, NTOUP-2018-06-310; 24.8 mm SL, Chenglong wetland, Yunlin County, Taiwan." captionText-1="FIGURE 2. Oryzias chenglongensis n. sp., female, NTOUP-2018 -06-311, 26.6 mm SL, Chenglong wetland, Yunlin County, Taiwan." captionText-2="FIGURE 3. Head profile and pigmentation pattern of Oryzias chenglongensis, holotype." captionText-3="FIGURE 4. Dorsal-fin pigmentation pattern of Oryzias chenglongensis, holotype (above) and Oryzias curvinotus (below)." figureDoi-0="http://doi.org/10.5281/zenodo.14389335" figureDoi-1="http://doi.org/10.5281/zenodo.14389337" figureDoi-2="http://doi.org/10.5281/zenodo.14389343" figureDoi-3="http://doi.org/10.5281/zenodo.14389345" httpUri-0="https://zenodo.org/record/14389335/files/figure.png" httpUri-1="https://zenodo.org/record/14389337/files/figure.png" httpUri-2="https://zenodo.org/record/14389343/files/figure.png" httpUri-3="https://zenodo.org/record/14389345/files/figure.png" pageId="1" pageNumber="321">Figs. 14</figureCitation>
)
</paragraph>
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<caption id="DF123FCE0B1FFFF2B993E4FCFF09BFBE" ID-DOI="http://doi.org/10.5281/zenodo.14389335" ID-Zenodo-Dep="14389335" httpUri="https://zenodo.org/record/14389335/files/figure.png" pageId="1" pageNumber="321" startId="1.[151,250,1944,1968]" targetBox="[151,1436,957,1921]" targetPageId="1" targetType="figure">
<paragraph id="8BD26F460B1FFFF2B993E4FCFF09BFBE" blockId="1.[151,1437,1944,2004]" pageId="1" pageNumber="321">
<emphasis id="B919B3540B1FFFF2B993E4FCFEF6BFDA" bold="true" box="[151,278,1944,1968]" pageId="1" pageNumber="321">FIGURE 1.</emphasis>
<taxonomicName id="4C6D14C50B1FFFF2B81BE4FDFDF3BFDA" authorityName="Chen &amp; Lai" authorityYear="2024" box="[287,531,1944,1968]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1FFFF2B81BE4FDFDF3BFDA" box="[287,531,1944,1968]" italics="true" pageId="1" pageNumber="321">Oryzias chenglongensis</emphasis>
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<emphasis id="B919B3540B1FFFF2BB18E4FCFDB8BFDA" bold="true" box="[540,600,1944,1968]" pageId="1" pageNumber="321">
<taxonomicNameLabel id="A22A0E2F0B1FFFF2BB18E4FCFDB8BFDA" box="[540,600,1944,1968]" pageId="1" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, male, NTOUP-2018-06-310; 24.8 mm SL, Chenglong wetland, Yunlin County, Taiwan.
</paragraph>
</caption>
<caption id="DF123FCE0B1CFFF1B993E7F5FF09BCA7" ID-DOI="http://doi.org/10.5281/zenodo.14389337" ID-Zenodo-Dep="14389337" httpUri="https://zenodo.org/record/14389337/files/figure.png" pageId="2" pageNumber="322" startId="2.[151,250,1169,1193]" targetBox="[151,1436,181,1145]" targetPageId="2" targetType="figure">
<paragraph id="8BD26F460B1CFFF1B993E7F5FF09BCA7" blockId="2.[151,1437,1169,1229]" pageId="2" pageNumber="322">
<emphasis id="B919B3540B1CFFF1B993E7F5FEF4BCC3" bold="true" box="[151,276,1169,1193]" pageId="2" pageNumber="322">FIGURE 2.</emphasis>
<taxonomicName id="4C6D14C50B1CFFF1B81FE7F5FDEDBCC3" authorityName="Chen &amp; Lai" authorityYear="2024" box="[283,525,1169,1193]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="2" pageNumber="322" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1CFFF1B81FE7F5FDEDBCC3" box="[283,525,1169,1193]" italics="true" pageId="2" pageNumber="322">Oryzias chenglongensis</emphasis>
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<emphasis id="B919B3540B1CFFF1BB11E7F5FDAFBCC3" bold="true" box="[533,591,1169,1193]" pageId="2" pageNumber="322">
<taxonomicNameLabel id="A22A0E2F0B1CFFF1BB11E7F5FDAFBCC3" box="[533,591,1169,1193]" pageId="2" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, NTOUP-2018 -06-311, 26.6 mm SL, Chenglong wetland, Yunlin County, Taiwan.
</paragraph>
</caption>
<subSubSection id="C3773CCD0B1CFFF1B9C3E67BFDAEBDA3" pageId="2" pageNumber="322" type="materials_examined">
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<materialsCitation id="3B05651B0B1CFFF1B9C3E67BFF38BD37" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="1" stateProvince="Yunlin" typeStatus="holotype">
<emphasis id="B919B3540B1CFFF1B9C3E67BFED4BD53" bold="true" box="[199,308,1311,1337]" pageId="2" pageNumber="322">
<typeStatus id="54D6D1E40B1CFFF1B9C3E67BFED4BD53" box="[199,308,1311,1337]" pageId="2" pageNumber="322" type="holotype">Holotype</typeStatus>
</emphasis>
.—NTOUP-2018-06-310;
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SL, coll. I-S.
<location id="8EB2399D0B1CFFF1BA64E67BFC7FBD53" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF0B993E003FABBBC5F:8EB2399D0B1CFFF1BA64E67BFC7FBD53" box="[864,927,1311,1337]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BAAEE67BFBC5BD53" box="[938,1061,1311,1337]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1BD97E67BFAD9BD53" box="[1171,1337,1311,1337]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1BC47E67BFA78BD53" box="[1347,1432,1311,1337]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC.
</materialsCitation>
</paragraph>
<paragraph id="8BD26F460B1CFFF1B9C3E603FDAEBDA3" blockId="2.[151,1436,1311,1481]" pageId="2" pageNumber="322">
<materialsCitation id="3B05651B0B1CFFF1B9C3E603FE04BDCF" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="3" stateProvince="Yunlin" typeStatus="paratype">
<emphasis id="B919B3540B1CFFF1B9C3E603FEDFBDEB" bold="true" box="[199,319,1383,1409]" pageId="2" pageNumber="322">
<typeStatus id="54D6D1E40B1CFFF1B9C3E603FEDFBDEB" box="[199,319,1383,1409]" pageId="2" pageNumber="322" type="paratype">Paratypes</typeStatus>
</emphasis>
.—NTOUP-2018-06-311;
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,
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SL, coll. I-S.
<location id="8EB2399D0B1CFFF1BD63E603FB46BDEB" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF0B993E003FABBBC5F:8EB2399D0B1CFFF1BD63E603FB46BDEB" box="[1127,1190,1383,1409]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BDB2E603FAD1BDEB" box="[1206,1329,1383,1409]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1B993E6EFFEDDBDCF" box="[151,317,1419,1445]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1B843E6EFFE7CBDCF" box="[327,412,1419,1445]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC
</materialsCitation>
.
<materialsCitation id="3B05651B0B1CFFF1B8EAE6EFFDAABDA3" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="20" stateProvince="Yunlin" typeStatus="paratype">
NTOUP-2018-02-312,
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SL,
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, coll. I-S.
<location id="8EB2399D0B1CFFF1BDDCE6EFFAF7BDCF" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF0B993E003FABBBC5F:8EB2399D0B1CFFF1BDDCE6EFFAF7BDCF" box="[1240,1303,1419,1445]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BC25E6EFFA7CBDCF" box="[1313,1436,1419,1445]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1B9F9E6CBFE42BDA3" box="[253,418,1455,1481]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1B8A9E6CBFDE2BDA3" box="[429,514,1455,1481]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1CFFF1B993E57FFCF8BFF7" pageId="2" pageNumber="322" type="diagnosis">
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<emphasis id="B919B3540B1CFFF1B993E57FFEF0BE5F" bold="true" box="[151,272,1563,1589]" pageId="2" pageNumber="322">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD26F460B1CFFF1B993E507FCF8BFF7" blockId="2.[151,1437,1635,1950]" pageId="2" pageNumber="322">
The new species,
<taxonomicName id="4C6D14C50B1CFFF1B864E507FD8BBE17" authorityName="Chen &amp; Lai" authorityYear="2024" box="[352,619,1635,1661]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="2" pageNumber="322" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1CFFF1B864E507FD8BBE17" box="[352,619,1635,1661]" italics="true" pageId="2" pageNumber="322">Oryzias chenglongensis</emphasis>
</taxonomicName>
can be well distinguished from other congeners by the following unique combination of features: (1) fin-ray counts: dorsal-fin rays 6 (6 seen in
<typeStatus id="54D6D1E40B1CFFF1BAB2E5E3FBFDBECB" box="[950,1053,1671,1697]" pageId="2" pageNumber="322" type="holotype">holotype</typeStatus>
); anal-fin rays 1920 (19); pelvic-fin rays 6 (6); pectoral-fin rays 10 (10); principle caudal-fin rays i,4/5,i (i,4/5,i); (2) body compressed laterally with body depth at pelvic origin 24.024.3 (24.3), body depth at anal fin origin 15.520.4 (20.4); (3) the dorsal-fin origin inserted almost at the vertical of 14
<superScript id="7C18C20E0B1CFFF1BB29E597FDDABF6B" attach="left" box="[557,570,1779,1793]" fontSize="6" pageId="2" pageNumber="322">th</superScript>
anal-fin rays, membrane between dorsal-fin 5
<superScript id="7C18C20E0B1CFFF1BD4EE597FBB7BF6B" attach="left" box="[1098,1111,1779,1793]" fontSize="6" pageId="2" pageNumber="322">th</superScript>
and 6
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rays without distinct notch in adult male; and (4) body translucent, light yellowish green or brown, scattered with minute melanophores, a narrowly dotted, longitudinal black line along the middle forward to vertical above pelvic fin; T-shape black mark of caudal fin base in male; operculum and pre-pectoral region slivery, no any silvery scales on lateral body, lip color grayish or creamy yellow and eye bluish silvery dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1DFFF0B993E3F3FABBBC5F" pageId="3" pageNumber="323" type="description">
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<emphasis id="B919B3540B1DFFF0B993E3F3FEC8B8DB" bold="true" box="[151,296,151,177]" pageId="3" pageNumber="323">Description</emphasis>
</paragraph>
<paragraph id="8BD26F460B1DFFF0B993E3BBFE35B92B" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Body proportions as their detailed morphometry were summarized in
<tableCitation id="C6EF5AFD0B1DFFF0BA9DE3BBFC0CB893" box="[921,1004,223,249]" captionStart="TABLE 1" captionStartId="3.[152,244,1125,1149]" captionText="TABLE 1. Morphometry of Oryzias chenglongensis n. sp. from Taiwan." pageId="3" pageNumber="323">Table 1</tableCitation>
. Dorsal-fin rays 6 (6 seen in
<typeStatus id="54D6D1E40B1DFFF0BC2DE3BBFA70B893" box="[1321,1424,223,249]" pageId="3" pageNumber="323" type="holotype">holotype</typeStatus>
); anal-fin rays 1920 (19); pelvic-fin rays 6 (6); pectoral-fin rays 10 (10); principle caudal-fin rays i,4/5,i (i,4/5,i), and vertebral counts 2930 (30).
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E22FFEE8BAEF" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Small species, with maximum size of available specimens examined as
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SL. Head moderate large, with head length 22.824.5 (22.8) depressed dorsoventrally; eye very large, eye diameter 34.841.3 (39.9); snout very short, snout length 22.825.9 (22.8). Body compressed laterally, robust anteriorly and slender posteriorly, body depth at pelvic origin 24.024.3 (24.3), body depth at anal fin origin 15.520.4 (20.4). No greatly pronounced abdominal concavity between pelvic fins and anterior part of anal fin in female. Mouth terminal, up-turned; tip of upper lip about the horizontal line crossing the upper margin of pupil, lower jaw projecting forward beyond upper jaw, each jaws in having minute conical teeth; lips slightly fleshy. Dorsal body profile relatively straight from head to mid-dorsum and curving from mid-dorsum to dorsal-fin origin; ventral profile as a gentle arch from head to anal-fin origin.
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E1EBFAADBAA4" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">Dorsal profile of head concave. Cheek very limited by large eye. Orbit not projecting toward the dorsal profile. Gill slit opening to superior margin of pectoral-fin base; isthmus located inferior to posterior margin of eye.</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E1B3FEACBB53" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">Scales relatively large and deciduous, cycloid. No scales on rostral zone. Cephalic lateral-line system absent on dorsal, lateral, jugular of head. Nostril opens upper jaw posterosuperior, and lachrymal in front of eye opens near anterior margin.</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E027FD4CBC7B" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Dorsal fin situated for posteriorly, the dorsal-fin origin inserted almost at the vertical of 14
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anal-fin rays. Membrane between dorsal-fin 5
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and 6
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rays without distinct notch in adult male. Dorsal fin rays may elongate with growth in male. Pelvic fin abdominal. Anal fin base rather long, 28.330.1 (29.8); membrane margin between each anal-fin ray indented in male, somewhat and slightly concave in female. Pectoral fin rays expended medially. Last pelvic-fin ray connected along less than one-half its length to body forming connecting membrane. Caudal fin truncate, median to posterior 1/3 part branched.
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E77FFABBBC5F" blockId="3.[151,1437,223,1078]" box="[199,1371,1051,1078]" pageId="3" pageNumber="323">Males with a short, tubular urogenital papilla; female with an enlarged bilobed fleshy, urogenital papilla.</paragraph>
</subSubSection>
</treatment>
</document>

View file

@ -0,0 +1,474 @@
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<mods:title id="B2F0A2DC14DF0670DA90CF7EA2316954">A new ricefish of genus Oryzias (Teleostei: Adrianichthyidae) from western Taiwan</mods:title>
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<mods:affiliation id="1E174787A33D0C5A3B1E17D693065965">Institute of Marine Biology, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C. &amp; Center of Excellence for the Oceans, National Taiwan Ocean University, Keelung, 202301, Taiwan, R. O. C.</mods:affiliation>
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<taxonomicName id="4C6D14C50B1FFFF2B993E003FE55BBEB" authorityName="Chen &amp; Lai" authorityYear="2024" box="[151,437,870,897]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1FFFF2B993E003FE55BBEB" bold="true" box="[151,437,870,897]" italics="true" pageId="1" pageNumber="321">Oryzias chenglongensis</emphasis>
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<paragraph id="8BD26F460B1FFFF2B993E0EFFEF2BBCF" blockId="1.[151,588,870,933]" box="[151,274,907,933]" pageId="1" pageNumber="321">
(
<figureCitation id="135673C30B1FFFF2B99BE0EFFEEABBCF" box="[159,266,907,933]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1944,1968]" captionStartId-1="2.[151,250,1169,1193]" captionStartId-2="4.[151,250,1588,1612]" captionStartId-3="5.[151,250,1548,1572]" captionTargetBox-0="[151,1436,957,1921]" captionTargetBox-1="[151,1436,181,1145]" captionTargetBox-2="[151,1436,742,1565]" captionTargetBox-3="[151,1436,181,1524]" captionTargetId-0="figure-226@1.[151,1436,957,1921]" captionTargetId-1="figure-19@2.[151,1436,181,1145]" captionTargetId-2="figure-95@4.[151,1436,728,1565]" captionTargetId-3="figure-25@5.[151,1436,179,872]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Oryzias chenglongensis n. sp., male, NTOUP-2018-06-310; 24.8 mm SL, Chenglong wetland, Yunlin County, Taiwan." captionText-1="FIGURE 2. Oryzias chenglongensis n. sp., female, NTOUP-2018 -06-311, 26.6 mm SL, Chenglong wetland, Yunlin County, Taiwan." captionText-2="FIGURE 3. Head profile and pigmentation pattern of Oryzias chenglongensis, holotype." captionText-3="FIGURE 4. Dorsal-fin pigmentation pattern of Oryzias chenglongensis, holotype (above) and Oryzias curvinotus (below)." figureDoi-0="http://doi.org/10.5281/zenodo.14389335" figureDoi-1="http://doi.org/10.5281/zenodo.14389337" figureDoi-2="http://doi.org/10.5281/zenodo.14389343" figureDoi-3="http://doi.org/10.5281/zenodo.14389345" httpUri-0="https://zenodo.org/record/14389335/files/figure.png" httpUri-1="https://zenodo.org/record/14389337/files/figure.png" httpUri-2="https://zenodo.org/record/14389343/files/figure.png" httpUri-3="https://zenodo.org/record/14389345/files/figure.png" pageId="1" pageNumber="321">Figs. 14</figureCitation>
)
</paragraph>
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<paragraph id="8BD26F460B1FFFF2B993E4FCFF09BFBE" blockId="1.[151,1437,1944,2004]" pageId="1" pageNumber="321">
<emphasis id="B919B3540B1FFFF2B993E4FCFEF6BFDA" bold="true" box="[151,278,1944,1968]" pageId="1" pageNumber="321">FIGURE 1.</emphasis>
<taxonomicName id="4C6D14C50B1FFFF2B81BE4FDFDF3BFDA" authorityName="Chen &amp; Lai" authorityYear="2024" box="[287,531,1944,1968]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="1" pageNumber="321" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1FFFF2B81BE4FDFDF3BFDA" box="[287,531,1944,1968]" italics="true" pageId="1" pageNumber="321">Oryzias chenglongensis</emphasis>
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<taxonomicNameLabel id="A22A0E2F0B1FFFF2BB18E4FCFDB8BFDA" box="[540,600,1944,1968]" pageId="1" pageNumber="321" rank="species">n. sp.</taxonomicNameLabel>
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, male, NTOUP-2018-06-310; 24.8 mm SL, Chenglong wetland, Yunlin County, Taiwan.
</paragraph>
</caption>
<caption id="DF123FCE0B1CFFF1B993E7F5FF09BCA7" ID-DOI="http://doi.org/10.5281/zenodo.14389337" ID-Zenodo-Dep="14389337" httpUri="https://zenodo.org/record/14389337/files/figure.png" pageId="2" pageNumber="322" startId="2.[151,250,1169,1193]" targetBox="[151,1436,181,1145]" targetPageId="2" targetType="figure">
<paragraph id="8BD26F460B1CFFF1B993E7F5FF09BCA7" blockId="2.[151,1437,1169,1229]" pageId="2" pageNumber="322">
<emphasis id="B919B3540B1CFFF1B993E7F5FEF4BCC3" bold="true" box="[151,276,1169,1193]" pageId="2" pageNumber="322">FIGURE 2.</emphasis>
<taxonomicName id="4C6D14C50B1CFFF1B81FE7F5FDEDBCC3" authorityName="Chen &amp; Lai" authorityYear="2024" box="[283,525,1169,1193]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="2" pageNumber="322" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1CFFF1B81FE7F5FDEDBCC3" box="[283,525,1169,1193]" italics="true" pageId="2" pageNumber="322">Oryzias chenglongensis</emphasis>
</taxonomicName>
<emphasis id="B919B3540B1CFFF1BB11E7F5FDAFBCC3" bold="true" box="[533,591,1169,1193]" pageId="2" pageNumber="322">
<taxonomicNameLabel id="A22A0E2F0B1CFFF1BB11E7F5FDAFBCC3" box="[533,591,1169,1193]" pageId="2" pageNumber="322" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, female, NTOUP-2018 -06-311, 26.6 mm SL, Chenglong wetland, Yunlin County, Taiwan.
</paragraph>
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<typeStatus id="54D6D1E40B1CFFF1B9C3E67BFED4BD53" box="[199,308,1311,1337]" pageId="2" pageNumber="322" type="holotype">Holotype</typeStatus>
</emphasis>
.—NTOUP-2018-06-310;
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SL, coll. I-S.
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wetland,
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,
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, ROC.
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<materialsCitation id="3B05651B0B1CFFF1B9C3E603FE04BDCF" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="3" stateProvince="Yunlin" typeStatus="paratype">
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<typeStatus id="54D6D1E40B1CFFF1B9C3E603FEDFBDEB" box="[199,319,1383,1409]" pageId="2" pageNumber="322" type="paratype">Paratypes</typeStatus>
</emphasis>
.—NTOUP-2018-06-311;
<specimenCount id="9D6BA4CF0B1CFFF1BB6FE603FD1EBDEB" box="[619,766,1383,1409]" count="3" pageId="2" pageNumber="322" type="generic">3 specimens</specimenCount>
,
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SL, coll. I-S.
<location id="8EB2399D0B1CFFF1BD63E603FB46BDEB" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF5B993E003FD84BC33:8EB2399D0B1CFFF1BD63E603FB46BDEB" box="[1127,1190,1383,1409]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BDB2E603FAD1BDEB" box="[1206,1329,1383,1409]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1B993E6EFFEDDBDCF" box="[151,317,1419,1445]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1B843E6EFFE7CBDCF" box="[327,412,1419,1445]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC
</materialsCitation>
.
<materialsCitation id="3B05651B0B1CFFF1B8EAE6EFFDAABDA3" country="Taiwan" location="Chen" municipality="Chenglong" pageId="2" pageNumber="322" specimenCount="20" stateProvince="Yunlin" typeStatus="paratype">
NTOUP-2018-02-312,
<quantity id="4C95C2A30B1CFFF1BBF7E6EFFC75BDCC" box="[755,917,1419,1446]" metricMagnitude="-2" metricUnit="m" metricValue="1.985" metricValueMax="2.45" metricValueMin="1.52" pageId="2" pageNumber="322" unit="mm" value="19.85" valueMax="24.5" valueMin="15.2">15.224.5 mm</quantity>
SL,
<specimenCount id="9D6BA4CF0B1CFFF1BACEE6EFFB85BDCF" box="[970,1125,1419,1445]" count="20" pageId="2" pageNumber="322" type="generic">20 specimens</specimenCount>
, coll. I-S.
<location id="8EB2399D0B1CFFF1BDDCE6EFFAF7BDCF" LSID="urn:lsid:plazi:treatment:03C4DE500B1FFFF5B993E003FD84BC33:8EB2399D0B1CFFF1BDDCE6EFFAF7BDCF" box="[1240,1303,1419,1445]" country="Taiwan" municipality="Chenglong" name="Chen" pageId="2" pageNumber="322" stateProvince="Yunlin">Chen</location>
;
<collectingMunicipality id="6BB6F53C0B1CFFF1BC25E6EFFA7CBDCF" box="[1313,1436,1419,1445]" pageId="2" pageNumber="322">Chenglong</collectingMunicipality>
wetland,
<collectingRegion id="49A9A1A40B1CFFF1B9F9E6CBFE42BDA3" box="[253,418,1455,1481]" country="Taiwan" name="Yunlin" pageId="2" pageNumber="322">Yunlin County</collectingRegion>
,
<collectingCountry id="F37A2FD60B1CFFF1B8A9E6CBFDE2BDA3" box="[429,514,1455,1481]" name="Taiwan" pageId="2" pageNumber="322">Taiwan</collectingCountry>
, ROC
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1CFFF1B993E57FFCF8BFF7" pageId="2" pageNumber="322" type="diagnosis">
<paragraph id="8BD26F460B1CFFF1B993E57FFEF0BE5F" blockId="2.[151,272,1563,1589]" box="[151,272,1563,1589]" pageId="2" pageNumber="322">
<emphasis id="B919B3540B1CFFF1B993E57FFEF0BE5F" bold="true" box="[151,272,1563,1589]" pageId="2" pageNumber="322">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BD26F460B1CFFF1B993E507FCF8BFF7" blockId="2.[151,1437,1635,1950]" pageId="2" pageNumber="322">
The new species,
<taxonomicName id="4C6D14C50B1CFFF1B864E507FD8BBE17" authorityName="Chen &amp; Lai" authorityYear="2024" box="[352,619,1635,1661]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="2" pageNumber="322" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1CFFF1B864E507FD8BBE17" box="[352,619,1635,1661]" italics="true" pageId="2" pageNumber="322">Oryzias chenglongensis</emphasis>
</taxonomicName>
can be well distinguished from other congeners by the following unique combination of features: (1) fin-ray counts: dorsal-fin rays 6 (6 seen in
<typeStatus id="54D6D1E40B1CFFF1BAB2E5E3FBFDBECB" box="[950,1053,1671,1697]" pageId="2" pageNumber="322" type="holotype">holotype</typeStatus>
); anal-fin rays 1920 (19); pelvic-fin rays 6 (6); pectoral-fin rays 10 (10); principle caudal-fin rays i,4/5,i (i,4/5,i); (2) body compressed laterally with body depth at pelvic origin 24.024.3 (24.3), body depth at anal fin origin 15.520.4 (20.4); (3) the dorsal-fin origin inserted almost at the vertical of 14
<superScript id="7C18C20E0B1CFFF1BB29E597FDDABF6B" attach="left" box="[557,570,1779,1793]" fontSize="6" pageId="2" pageNumber="322">th</superScript>
anal-fin rays, membrane between dorsal-fin 5
<superScript id="7C18C20E0B1CFFF1BD4EE597FBB7BF6B" attach="left" box="[1098,1111,1779,1793]" fontSize="6" pageId="2" pageNumber="322">th</superScript>
and 6
<superScript id="7C18C20E0B1CFFF1BD9AE597FB4BBF6B" attach="left" box="[1182,1195,1779,1793]" fontSize="6" pageId="2" pageNumber="322">th</superScript>
rays without distinct notch in adult male; and (4) body translucent, light yellowish green or brown, scattered with minute melanophores, a narrowly dotted, longitudinal black line along the middle forward to vertical above pelvic fin; T-shape black mark of caudal fin base in male; operculum and pre-pectoral region slivery, no any silvery scales on lateral body, lip color grayish or creamy yellow and eye bluish silvery dorsally.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B1DFFF6B993E3F3FAD1BF02" lastPageId="5" lastPageNumber="325" pageId="3" pageNumber="323" type="description">
<paragraph id="8BD26F460B1DFFF0B993E3F3FEC8B8DB" blockId="3.[151,296,151,177]" box="[151,296,151,177]" pageId="3" pageNumber="323">
<emphasis id="B919B3540B1DFFF0B993E3F3FEC8B8DB" bold="true" box="[151,296,151,177]" pageId="3" pageNumber="323">Description</emphasis>
</paragraph>
<paragraph id="8BD26F460B1DFFF0B993E3BBFE35B92B" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Body proportions as their detailed morphometry were summarized in
<tableCitation id="C6EF5AFD0B1DFFF0BA9DE3BBFC0CB893" box="[921,1004,223,249]" captionStart="TABLE 1" captionStartId="3.[152,244,1125,1149]" captionTargetPageId="3" captionText="TABLE 1. Morphometry of Oryzias chenglongensis n. sp. from Taiwan." httpUri="http://table.plazi.org/id/DF123FCE0B1DFFF0B99CE701FC44BC17" pageId="3" pageNumber="323" tableUuid="DF123FCE0B1DFFF0B99CE701FC44BC17">Table 1</tableCitation>
. Dorsal-fin rays 6 (6 seen in
<typeStatus id="54D6D1E40B1DFFF0BC2DE3BBFA70B893" box="[1321,1424,223,249]" pageId="3" pageNumber="323" type="holotype">holotype</typeStatus>
); anal-fin rays 1920 (19); pelvic-fin rays 6 (6); pectoral-fin rays 10 (10); principle caudal-fin rays i,4/5,i (i,4/5,i), and vertebral counts 2930 (30).
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E22FFEE8BAEF" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Small species, with maximum size of available specimens examined as
<quantity id="4C95C2A30B1DFFF0BD02E22FFB8CB90C" box="[1030,1132,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="2.66" pageId="3" pageNumber="323" unit="mm" value="26.6">26.6 mm</quantity>
SL. Head moderate large, with head length 22.824.5 (22.8) depressed dorsoventrally; eye very large, eye diameter 34.841.3 (39.9); snout very short, snout length 22.825.9 (22.8). Body compressed laterally, robust anteriorly and slender posteriorly, body depth at pelvic origin 24.024.3 (24.3), body depth at anal fin origin 15.520.4 (20.4). No greatly pronounced abdominal concavity between pelvic fins and anterior part of anal fin in female. Mouth terminal, up-turned; tip of upper lip about the horizontal line crossing the upper margin of pupil, lower jaw projecting forward beyond upper jaw, each jaws in having minute conical teeth; lips slightly fleshy. Dorsal body profile relatively straight from head to mid-dorsum and curving from mid-dorsum to dorsal-fin origin; ventral profile as a gentle arch from head to anal-fin origin.
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E1EBFAADBAA4" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">Dorsal profile of head concave. Cheek very limited by large eye. Orbit not projecting toward the dorsal profile. Gill slit opening to superior margin of pectoral-fin base; isthmus located inferior to posterior margin of eye.</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E1B3FEACBB53" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">Scales relatively large and deciduous, cycloid. No scales on rostral zone. Cephalic lateral-line system absent on dorsal, lateral, jugular of head. Nostril opens upper jaw posterosuperior, and lachrymal in front of eye opens near anterior margin.</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E027FD4CBC7B" blockId="3.[151,1437,223,1078]" pageId="3" pageNumber="323">
Dorsal fin situated for posteriorly, the dorsal-fin origin inserted almost at the vertical of 14
<superScript id="7C18C20E0B1DFFF0BDEAE027FB1BBB3B" attach="left" box="[1262,1275,835,849]" fontSize="6" pageId="3" pageNumber="323">th</superScript>
anal-fin rays. Membrane between dorsal-fin 5
<superScript id="7C18C20E0B1DFFF0BB00E003FDF1BB1F" attach="left" box="[516,529,871,885]" fontSize="6" pageId="3" pageNumber="323">th</superScript>
and 6
<superScript id="7C18C20E0B1DFFF0BB5EE003FD87BB1F" attach="left" box="[602,615,871,885]" fontSize="6" pageId="3" pageNumber="323">th</superScript>
rays without distinct notch in adult male. Dorsal fin rays may elongate with growth in male. Pelvic fin abdominal. Anal fin base rather long, 28.330.1 (29.8); membrane margin between each anal-fin ray indented in male, somewhat and slightly concave in female. Pectoral fin rays expended medially. Last pelvic-fin ray connected along less than one-half its length to body forming connecting membrane. Caudal fin truncate, median to posterior 1/3 part branched.
</paragraph>
<paragraph id="8BD26F460B1DFFF0B9C3E77FFABBBC5F" blockId="3.[151,1437,223,1078]" box="[199,1371,1051,1078]" pageId="3" pageNumber="323">Males with a short, tubular urogenital papilla; female with an enlarged bilobed fleshy, urogenital papilla.</paragraph>
<caption id="DF123FCE0B1DFFF0B99CE701FC44BC17" ID-Table-UUID="DF123FCE0B1DFFF0B99CE701FC44BC17" box="[152,932,1125,1149]" httpUri="http://table.plazi.org/id/DF123FCE0B1DFFF0B99CE701FC44BC17" pageId="3" pageNumber="323" startId="3.[152,244,1125,1149]" targetBox="[163,1310,1165,1954]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="8BD26F460B1DFFF0B99CE701FC44BC17" blockId="3.[152,932,1125,1149]" box="[152,932,1125,1149]" pageId="3" pageNumber="323">
<emphasis id="B919B3540B1DFFF0B99CE701FEEFBC17" bold="true" box="[152,271,1125,1149]" pageId="3" pageNumber="323">TABLE 1.</emphasis>
Morphometry of
<taxonomicName id="4C6D14C50B1DFFF0B8C9E701FD2BBC17" authorityName="Chen &amp; Lai" authorityYear="2024" box="[461,715,1125,1149]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="3" pageNumber="323" phylum="Chordata" rank="species" species="chenglongensis" status="sp. nov.">
<emphasis id="B919B3540B1DFFF0B8C9E701FD2BBC17" box="[461,715,1125,1149]" italics="true" pageId="3" pageNumber="323">Oryzias chenglongensis</emphasis>
</taxonomicName>
<emphasis id="B919B3540B1DFFF0BBD6E701FCEEBC17" bold="true" box="[722,782,1125,1149]" pageId="3" pageNumber="323">
<taxonomicNameLabel id="A22A0E2F0B1DFFF0BBD6E701FCEEBC17" box="[722,782,1125,1149]" pageId="3" pageNumber="323" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Taiwan.
</paragraph>
</caption>
<paragraph id="8BD26F460B1DFFF0BBFCE7E9FAE6BFC8" pageId="3" pageNumber="323">
<table id="F96D9DE60B1D000CB9A7E7E9FAFEBFC8" box="[163,1310,1165,1954]" colsContinueIn="4.[163,1310,202,669]" gridcols="4" gridrows="20" pageId="3" pageNumber="323">
<tr id="355D6D040B1D000CB9A7E7E9FAFEBCCC" box="[163,1310,1165,1190]" gridrow="0" pageId="3" pageNumber="323" rowspan-0="1">
<th id="768C04780B1D000CBBFCE7E9FCB6BCCC" box="[760,854,1165,1190]" gridcol="1" gridrow="0" pageId="3" pageNumber="323">Holotype</th>
<th id="768C04780B1D000CBADAE7E9FBD7BCCC" box="[990,1079,1165,1190]" gridcol="2" gridrow="0" pageId="3" pageNumber="323">Paratype</th>
<th id="768C04780B1D000CBDC1E7E9FAFEBCCC" box="[1221,1310,1165,1190]" gridcol="3" gridrow="0" pageId="3" pageNumber="323">Paratype</th>
</tr>
<tr id="355D6D040B1D000CB9A7E7D1FAFEBCA7" box="[163,1310,1205,1229]" gridrow="1" pageId="3" pageNumber="323" rowspan-0="1">
<td id="768C04780B1D000CBBFCE7D1FCB6BCA7" box="[760,854,1205,1229]" gridcol="1" gridrow="1" pageId="3" pageNumber="323">Male</td>
<td id="768C04780B1D000CBADAE7D1FBD7BCA7" box="[990,1079,1205,1229]" gridcol="2" gridrow="1" pageId="3" pageNumber="323">Female</td>
<td id="768C04780B1D000CBDC1E7D1FAFEBCA7" box="[1221,1310,1205,1229]" gridcol="3" gridrow="1" pageId="3" pageNumber="323">Female</td>
</tr>
<tr id="355D6D040B1D000CB9A7E7BAFAFEBC9C" box="[163,1310,1246,1270]" gridrow="2" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E7BAFD70BC9C" box="[163,656,1246,1270]" gridcol="0" gridrow="2" pageId="3" pageNumber="323">Standard length (mm)</th>
<td id="768C04780B1D000CBBFCE7BAFCB6BC9C" box="[760,854,1246,1270]" gridcol="1" gridrow="2" pageId="3" pageNumber="323">24.8</td>
<td id="768C04780B1D000CBADAE7BAFBD7BC9C" box="[990,1079,1246,1270]" gridcol="2" gridrow="2" pageId="3" pageNumber="323">26.6</td>
<td id="768C04780B1D000CBDC1E7BAFAFEBC9C" box="[1221,1310,1246,1270]" gridcol="3" gridrow="2" pageId="3" pageNumber="323">23.8</td>
</tr>
<tr id="355D6D040B1D000CB9A7E662FAFEBD74" box="[163,1310,1286,1310]" gridrow="3" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E662FAFEBD74" box="[163,1310,1286,1310]" colspan="4" colspanRight="3" gridcol="0" gridrow="3" pageId="3" pageNumber="323">% in Standard length</th>
</tr>
<tr id="355D6D040B1D000CB9A7E64AFAFEBD2C" box="[163,1310,1326,1350]" gridrow="4" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E64AFD70BD2C" box="[163,656,1326,1350]" gridcol="0" gridrow="4" pageId="3" pageNumber="323">Body depth at pelvic fin origin</th>
<td id="768C04780B1D000CBBFCE64AFCB6BD2C" box="[760,854,1326,1350]" gridcol="1" gridrow="4" pageId="3" pageNumber="323">24.3%</td>
<td id="768C04780B1D000CBADAE64AFBD7BD2C" box="[990,1079,1326,1350]" gridcol="2" gridrow="4" pageId="3" pageNumber="323">24.0%</td>
<td id="768C04780B1D000CBDC1E64AFAFEBD2C" box="[1221,1310,1326,1350]" gridcol="3" gridrow="4" pageId="3" pageNumber="323">24.2%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E632FAFEBD04" box="[163,1310,1366,1390]" gridrow="5" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E632FD70BD04" box="[163,656,1366,1390]" gridcol="0" gridrow="5" pageId="3" pageNumber="323">Body depth at anal fin origin</th>
<td id="768C04780B1D000CBBFCE632FCB6BD04" box="[760,854,1366,1390]" gridcol="1" gridrow="5" pageId="3" pageNumber="323">20.4%</td>
<td id="768C04780B1D000CBADAE632FBD7BD04" box="[990,1079,1366,1390]" gridcol="2" gridrow="5" pageId="3" pageNumber="323">19.2%</td>
<td id="768C04780B1D000CBDC1E632FAFEBD04" box="[1221,1310,1366,1390]" gridcol="3" gridrow="5" pageId="3" pageNumber="323">15.5%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E61AFAFEBDFD" box="[163,1310,1406,1431]" gridrow="6" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E61AFD70BDFD" box="[163,656,1406,1431]" gridcol="0" gridrow="6" pageId="3" pageNumber="323">Body width</th>
<td id="768C04780B1D000CBBFCE61AFCB6BDFD" box="[760,854,1406,1431]" gridcol="1" gridrow="6" pageId="3" pageNumber="323">16.2%</td>
<td id="768C04780B1D000CBADAE61AFBD7BDFD" box="[990,1079,1406,1431]" gridcol="2" gridrow="6" pageId="3" pageNumber="323">16.6%</td>
<td id="768C04780B1D000CBDC1E61AFAFEBDFD" box="[1221,1310,1406,1431]" gridcol="3" gridrow="6" pageId="3" pageNumber="323">18.3%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E6C3FAFEBDD5" box="[163,1310,1447,1471]" gridrow="7" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E6C3FD70BDD5" box="[163,656,1447,1471]" gridcol="0" gridrow="7" pageId="3" pageNumber="323">Body depth</th>
<td id="768C04780B1D000CBBFCE6C3FCB6BDD5" box="[760,854,1447,1471]" gridcol="1" gridrow="7" pageId="3" pageNumber="323">22.8%</td>
<td id="768C04780B1D000CBADAE6C3FBD7BDD5" box="[990,1079,1447,1471]" gridcol="2" gridrow="7" pageId="3" pageNumber="323">24.5%</td>
<td id="768C04780B1D000CBDC1E6C3FAFEBDD5" box="[1221,1310,1447,1471]" gridcol="3" gridrow="7" pageId="3" pageNumber="323">23.8</td>
</tr>
<tr id="355D6D040B1D000CB9A7E6ABFAFEBD8D" box="[163,1310,1487,1511]" gridrow="8" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E6ABFD70BD8D" box="[163,656,1487,1511]" gridcol="0" gridrow="8" pageId="3" pageNumber="323">Caudal peduncle depth</th>
<td id="768C04780B1D000CBBFCE6ABFCB6BD8D" box="[760,854,1487,1511]" gridcol="1" gridrow="8" pageId="3" pageNumber="323">9.1%</td>
<td id="768C04780B1D000CBADAE6ABFBD7BD8D" box="[990,1079,1487,1511]" gridcol="2" gridrow="8" pageId="3" pageNumber="323">8.8%</td>
<td id="768C04780B1D000CBDC1E6ABFAFEBD8D" box="[1221,1310,1487,1511]" gridcol="3" gridrow="8" pageId="3" pageNumber="323">8.2%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E693FAFEBE65" box="[163,1310,1527,1551]" gridrow="9" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E693FD70BE65" box="[163,656,1527,1551]" gridcol="0" gridrow="9" pageId="3" pageNumber="323">Caudal peduncle length</th>
<td id="768C04780B1D000CBBFCE693FCB6BE65" box="[760,854,1527,1551]" gridcol="1" gridrow="9" pageId="3" pageNumber="323">13.9%</td>
<td id="768C04780B1D000CBADAE693FBD7BE65" box="[990,1079,1527,1551]" gridcol="2" gridrow="9" pageId="3" pageNumber="323">13.5%</td>
<td id="768C04780B1D000CBDC1E693FAFEBE65" box="[1221,1310,1527,1551]" gridcol="3" gridrow="9" pageId="3" pageNumber="323">13.8%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E57BFAFEBE52" box="[163,1310,1567,1592]" gridrow="10" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E57BFD70BE52" box="[163,656,1567,1592]" gridcol="0" gridrow="10" pageId="3" pageNumber="323">Height of dorsal fin</th>
<td id="768C04780B1D000CBBFCE57BFCB6BE52" box="[760,854,1567,1592]" gridcol="1" gridrow="10" pageId="3" pageNumber="323">18.8%</td>
<td id="768C04780B1D000CBADAE57BFBD7BE52" box="[990,1079,1567,1592]" gridcol="2" gridrow="10" pageId="3" pageNumber="323">12.7%</td>
<td id="768C04780B1D000CBDC1E57BFAFEBE52" box="[1221,1310,1567,1592]" gridcol="3" gridrow="10" pageId="3" pageNumber="323">15.6%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E52CFAFEBE0A" box="[163,1310,1608,1632]" gridrow="11" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E52CFD70BE0A" box="[163,656,1608,1632]" gridcol="0" gridrow="11" pageId="3" pageNumber="323">Dorsal-fin base</th>
<td id="768C04780B1D000CBBFCE52CFCB6BE0A" box="[760,854,1608,1632]" gridcol="1" gridrow="11" pageId="3" pageNumber="323">8.6%</td>
<td id="768C04780B1D000CBADAE52CFBD7BE0A" box="[990,1079,1608,1632]" gridcol="2" gridrow="11" pageId="3" pageNumber="323">7.6%</td>
<td id="768C04780B1D000CBDC1E52CFAFEBE0A" box="[1221,1310,1608,1632]" gridcol="3" gridrow="11" pageId="3" pageNumber="323">8.7%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E514FAFEBEE2" box="[163,1310,1648,1672]" gridrow="12" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E514FD70BEE2" box="[163,656,1648,1672]" gridcol="0" gridrow="12" pageId="3" pageNumber="323">Height of anal fin</th>
<td id="768C04780B1D000CBBFCE514FCB6BEE2" box="[760,854,1648,1672]" gridcol="1" gridrow="12" pageId="3" pageNumber="323">14.6%</td>
<td id="768C04780B1D000CBADAE514FBD7BEE2" box="[990,1079,1648,1672]" gridcol="2" gridrow="12" pageId="3" pageNumber="323">13.3%</td>
<td id="768C04780B1D000CBDC1E514FAFEBEE2" box="[1221,1310,1648,1672]" gridcol="3" gridrow="12" pageId="3" pageNumber="323">14.3%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E5FCFAFEBEDA" box="[163,1310,1688,1712]" gridrow="13" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E5FCFD70BEDA" box="[163,656,1688,1712]" gridcol="0" gridrow="13" pageId="3" pageNumber="323">Anal-fin base</th>
<td id="768C04780B1D000CBBFCE5FCFCB6BEDA" box="[760,854,1688,1712]" gridcol="1" gridrow="13" pageId="3" pageNumber="323">29.8%</td>
<td id="768C04780B1D000CBADAE5FCFBD7BEDA" box="[990,1079,1688,1712]" gridcol="2" gridrow="13" pageId="3" pageNumber="323">28.3%</td>
<td id="768C04780B1D000CBDC1E5FCFAFEBEDA" box="[1221,1310,1688,1712]" gridcol="3" gridrow="13" pageId="3" pageNumber="323">30.1%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E5A4FAFEBEB3" box="[163,1310,1728,1753]" gridrow="14" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E5A4FD70BEB3" box="[163,656,1728,1753]" gridcol="0" gridrow="14" pageId="3" pageNumber="323">Pelvic fin length</th>
<td id="768C04780B1D000CBBFCE5A4FCB6BEB3" box="[760,854,1728,1753]" gridcol="1" gridrow="14" pageId="3" pageNumber="323">11.9%</td>
<td id="768C04780B1D000CBADAE5A4FBD7BEB3" box="[990,1079,1728,1753]" gridcol="2" gridrow="14" pageId="3" pageNumber="323">13.7%</td>
<td id="768C04780B1D000CBDC1E5A4FAFEBEB3" box="[1221,1310,1728,1753]" gridcol="3" gridrow="14" pageId="3" pageNumber="323">14.2%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E58DFAFEBF6B" box="[163,1310,1769,1793]" gridrow="15" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E58DFD70BF6B" box="[163,656,1769,1793]" gridcol="0" gridrow="15" pageId="3" pageNumber="323">Pectoral-fin lrngth</th>
<td id="768C04780B1D000CBBFCE58DFCB6BF6B" box="[760,854,1769,1793]" gridcol="1" gridrow="15" pageId="3" pageNumber="323">18.8%</td>
<td id="768C04780B1D000CBADAE58DFBD7BF6B" box="[990,1079,1769,1793]" gridcol="2" gridrow="15" pageId="3" pageNumber="323">17.5%</td>
<td id="768C04780B1D000CBDC1E58DFAFEBF6B" box="[1221,1310,1769,1793]" gridcol="3" gridrow="15" pageId="3" pageNumber="323">17.7%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E475FAFEBF43" box="[163,1310,1809,1833]" gridrow="16" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E475FD70BF43" box="[163,656,1809,1833]" gridcol="0" gridrow="16" pageId="3" pageNumber="323">Predorsal length</th>
<td id="768C04780B1D000CBBFCE475FCB6BF43" box="[760,854,1809,1833]" gridcol="1" gridrow="16" pageId="3" pageNumber="323">79.2%</td>
<td id="768C04780B1D000CBADAE475FBD7BF43" box="[990,1079,1809,1833]" gridcol="2" gridrow="16" pageId="3" pageNumber="323">79.2%</td>
<td id="768C04780B1D000CBDC1E475FAFEBF43" box="[1221,1310,1809,1833]" gridcol="3" gridrow="16" pageId="3" pageNumber="323">77.0%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E45DFAFEBF3B" box="[163,1310,1849,1873]" gridrow="17" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E45DFD70BF3B" box="[163,656,1849,1873]" gridcol="0" gridrow="17" pageId="3" pageNumber="323">Preanal length</th>
<td id="768C04780B1D000CBBFCE45DFCB6BF3B" box="[760,854,1849,1873]" gridcol="1" gridrow="17" pageId="3" pageNumber="323">59.0%</td>
<td id="768C04780B1D000CBADAE45DFBD7BF3B" box="[990,1079,1849,1873]" gridcol="2" gridrow="17" pageId="3" pageNumber="323">60.3%</td>
<td id="768C04780B1D000CBDC1E45DFAFEBF3B" box="[1221,1310,1849,1873]" gridcol="3" gridrow="17" pageId="3" pageNumber="323">59.5%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E405FAFEBF10" box="[163,1310,1889,1914]" gridrow="18" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E405FD70BF10" box="[163,656,1889,1914]" gridcol="0" gridrow="18" pageId="3" pageNumber="323">Prepelvic length</th>
<td id="768C04780B1D000CBBFCE405FCB6BF10" box="[760,854,1889,1914]" gridcol="1" gridrow="18" pageId="3" pageNumber="323">44.9%</td>
<td id="768C04780B1D000CBADAE405FBD7BF10" box="[990,1079,1889,1914]" gridcol="2" gridrow="18" pageId="3" pageNumber="323">46.8%</td>
<td id="768C04780B1D000CBDC1E405FAFEBF10" box="[1221,1310,1889,1914]" gridcol="3" gridrow="18" pageId="3" pageNumber="323">45.5%</td>
</tr>
<tr id="355D6D040B1D000CB9A7E4EEFAFEBFC8" box="[163,1310,1930,1954]" gridrow="19" pageId="3" pageNumber="323">
<th id="768C04780B1D000CB9A7E4EEFD70BFC8" box="[163,656,1930,1954]" gridcol="0" gridrow="19" pageId="3" pageNumber="323">Length from Pectoral fin basee to pelvic fin base</th>
<td id="768C04780B1D000CBBFCE4EEFCB6BFC8" box="[760,854,1930,1954]" gridcol="1" gridrow="19" pageId="3" pageNumber="323">16.8%</td>
<td id="768C04780B1D000CBADAE4EEFBD7BFC8" box="[990,1079,1930,1954]" gridcol="2" gridrow="19" pageId="3" pageNumber="323">17.2%</td>
<td id="768C04780B1D000CBDC1E4EEFAFEBFC8" box="[1221,1310,1930,1954]" gridcol="3" gridrow="19" pageId="3" pageNumber="323">16.6%</td>
</tr>
</table>
</paragraph>
<paragraph id="8BD26F460B1DFFF0BD5DE4D7FA70BFA0" blockId="3.[1113,1424,1971,1995]" box="[1113,1424,1971,1995]" pageId="3" pageNumber="323">
<tableNote id="768B6EC80B1DFFF0BD5DE4D7FA70BFA0" box="[1113,1424,1971,1995]" pageId="3" pageNumber="323" targetBox="[163,1310,1165,1954]" targetPageId="3">......continued on the next page</tableNote>
</paragraph>
<caption id="DF123FCE0B1AFFF7B9A7E3C6FE71B8D0" box="[163,401,162,186]" isContinuationCaption="true" pageId="4" pageNumber="324" startId="4.[163,247,162,186]" targetBox="[163,1310,202,669]" targetIsTable="true" targetPageId="4" targetType="table">
<paragraph id="8BD26F460B1AFFF7B9A7E3C6FE71B8D0" blockId="4.[163,401,162,186]" box="[163,401,162,186]" pageId="4" pageNumber="324">
<emphasis id="B919B3540B1AFFF7B9A7E3C6FEF0B8D0" bold="true" box="[163,272,162,186]" pageId="4" pageNumber="324">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BD26F460B1AFFF7BBFCE3AEFAE6BAF7" pageId="4" pageNumber="324">
<table id="F96D9DE60B1A000CB9A7E3AEFAFEBAF7" box="[163,1310,202,669]" colsContinueFrom="3.[163,1310,1165,1954]" gridcols="4" gridrows="5" pageId="4" pageNumber="324">
<tr id="355D6D040B1A000CB9A7E3AEFAFEB889" box="[163,1310,202,227]" gridrow="0" pageId="4" pageNumber="324" rowspan-0="1">
<th id="768C04780B1A000CBBFCE3AEFCB6B889" box="[760,854,202,227]" gridcol="1" gridrow="0" pageId="4" pageNumber="324">Holotype</th>
<th id="768C04780B1A000CBADAE3AEFBD7B889" box="[990,1079,202,227]" gridcol="2" gridrow="0" pageId="4" pageNumber="324">Paratype</th>
<th id="768C04780B1A000CBDC1E3AEFAFEB889" box="[1221,1310,202,227]" gridcol="3" gridrow="0" pageId="4" pageNumber="324">Paratype</th>
</tr>
<tr id="355D6D040B1A000CB9A7E396FAFEB960" box="[163,1310,242,266]" gridrow="1" pageId="4" pageNumber="324" rowspan-0="1">
<td id="768C04780B1A000CBBFCE396FCB6B960" box="[760,854,242,266]" gridcol="1" gridrow="1" pageId="4" pageNumber="324">Male</td>
<td id="768C04780B1A000CBADAE396FBD7B960" box="[990,1079,242,266]" gridcol="2" gridrow="1" pageId="4" pageNumber="324">Female</td>
<td id="768C04780B1A000CBDC1E396FAFEB960" box="[1221,1310,242,266]" gridcol="3" gridrow="1" pageId="4" pageNumber="324">Female</td>
</tr>
<tr id="355D6D040B1A000CB9A7E27FFAFEB959" box="[163,1310,283,307]" gridrow="2" pageId="4" pageNumber="324">
<th id="768C04780B1A000CB9A7E27FFD60B959" box="[163,640,283,307]" gridcol="0" gridrow="2" pageId="4" pageNumber="324">Length from dorsal fin basee to caudal fin base</th>
<td id="768C04780B1A000CBBFCE27FFCB6B959" box="[760,854,283,307]" gridcol="1" gridrow="2" pageId="4" pageNumber="324">14.3%</td>
<td id="768C04780B1A000CBADAE27FFBD7B959" box="[990,1079,283,307]" gridcol="2" gridrow="2" pageId="4" pageNumber="324">15.8%</td>
<td id="768C04780B1A000CBDC1E27FFAFEB959" box="[1221,1310,283,307]" gridcol="3" gridrow="2" pageId="4" pageNumber="324">16.9%</td>
</tr>
<tr id="355D6D040B1A000CB9A7E208FAFEBA27" box="[163,1310,364,589]" gridrow="3" pageId="4" pageNumber="324">
<th id="768C04780B1A000CB9A7E208FD60BA27" box="[163,640,364,589]" gridcol="0" gridrow="3" pageId="4" pageNumber="324">% in Head length Head width Eye diameter Postorbital length Snout length Interorbital width</th>
<td id="768C04780B1A000CBBFCE208FCB6BA27" box="[760,854,364,589]" gridcol="1" gridrow="3" pageId="4" pageNumber="324">66.6% 36.9% 39.9% 22.8% 33.1%</td>
<td id="768C04780B1A000CBADAE208FBD7BA27" box="[990,1079,364,589]" gridcol="2" gridrow="3" pageId="4" pageNumber="324">54.8% 35.8% 41.3% 23.9% 35.2%</td>
<td id="768C04780B1A000CBDC1E208FAFEBA27" box="[1221,1310,364,589]" gridcol="3" gridrow="3" pageId="4" pageNumber="324">70.2% 34.8% 40.0% 25.9% 39.1%</td>
</tr>
<tr id="355D6D040B1A000CB9A7E1E1FAFEBAF7" box="[163,1310,645,669]" gridrow="4" pageId="4" pageNumber="324">
<th id="768C04780B1A000CB9A7E1E1FD60BAF7" box="[163,640,645,669]" gridcol="0" gridrow="4" pageId="4" pageNumber="324">Caudal peduncle depth/length</th>
<td id="768C04780B1A000CBBFCE1E1FCB6BAF7" box="[760,854,645,669]" gridcol="1" gridrow="4" pageId="4" pageNumber="324">65.5%</td>
<td id="768C04780B1A000CBADAE1E1FBD7BAF7" box="[990,1079,645,669]" gridcol="2" gridrow="4" pageId="4" pageNumber="324">65.5%</td>
<td id="768C04780B1A000CBDC1E1E1FAFEBAF7" box="[1221,1310,645,669]" gridcol="3" gridrow="4" pageId="4" pageNumber="324">59.4%</td>
</tr>
</table>
</paragraph>
<caption id="DF123FCE0B1AFFF7B993E550FBFBBE26" ID-DOI="http://doi.org/10.5281/zenodo.14389343" ID-Zenodo-Dep="14389343" box="[151,1051,1588,1613]" httpUri="https://zenodo.org/record/14389343/files/figure.png" pageId="4" pageNumber="324" startId="4.[151,250,1588,1612]" targetBox="[151,1436,742,1565]" targetPageId="4" targetType="figure">
<paragraph id="8BD26F460B1AFFF7B993E550FBFBBE26" blockId="4.[151,1051,1588,1613]" box="[151,1051,1588,1613]" pageId="4" pageNumber="324">
<emphasis id="B919B3540B1AFFF7B993E550FEF3BE26" bold="true" box="[151,275,1588,1612]" pageId="4" pageNumber="324">FIGURE 3.</emphasis>
Head profile and pigmentation pattern of
<taxonomicName id="4C6D14C50B1AFFF7BBBAE551FC4FBE27" authorityName="Chen &amp; Lai" authorityYear="2024" box="[702,943,1589,1613]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="4" pageNumber="324" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1AFFF7BBBAE551FC4FBE27" box="[702,943,1589,1613]" italics="true" pageId="4" pageNumber="324">Oryzias chenglongensis</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
<paragraph id="8BD26F460B1AFFF7B993E5FBFE8DBED3" blockId="4.[151,365,1695,1721]" box="[151,365,1695,1721]" pageId="4" pageNumber="324">
<heading id="D09AD82A0B1AFFF7B993E5FBFE8DBED3" bold="true" box="[151,365,1695,1721]" fontSize="11" level="1" pageId="4" pageNumber="324" reason="1">
<emphasis id="B919B3540B1AFFF7B993E5FBFE8DBED3" bold="true" box="[151,365,1695,1721]" pageId="4" pageNumber="324">Coloration in life</emphasis>
</heading>
</paragraph>
<paragraph id="8BD26F460B1AFFF7B993E583FAFBBFFB" blockId="4.[151,1436,1767,1937]" pageId="4" pageNumber="324">Body translucent, light yellowish green or brown, scattered with minute melanophores can be seen alive and preservative, a conspicuous, narrowly dotted, longitudinal black line along the middle forward to vertical above pelvic fin. T-shape black mark of caudal fin base in male. Belly wall, peritoneum slivery, somewhat subrectangular in both sexes. Operculum and pre-pectoral region slivery. No any silvery scales on lateral body. Lip color grayish or somewhat creamy yellow. Eye bluish silvery dorsally, silvery ventrally with some brilliant yellow dots.</paragraph>
<caption id="DF123FCE0B1BFFF6B993E568FA99BE4E" ID-DOI="http://doi.org/10.5281/zenodo.14389345" ID-Zenodo-Dep="14389345" box="[151,1401,1548,1572]" httpUri="https://zenodo.org/record/14389345/files/figure.png" pageId="5" pageNumber="325" startId="5.[151,250,1548,1572]" targetBox="[151,1436,181,1524]" targetPageId="5" targetType="figure">
<paragraph id="8BD26F460B1BFFF6B993E568FA99BE4E" blockId="5.[151,1401,1548,1572]" box="[151,1401,1548,1572]" pageId="5" pageNumber="325">
<emphasis id="B919B3540B1BFFF6B993E568FEF3BE4E" bold="true" box="[151,275,1548,1572]" pageId="5" pageNumber="325">FIGURE 4.</emphasis>
Dorsal-fin pigmentation pattern of
<taxonomicName id="4C6D14C50B1BFFF6BB78E569FC8DBE4E" authorityName="Chen &amp; Lai" authorityYear="2024" box="[636,877,1548,1572]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="5" pageNumber="325" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B1BFFF6BB78E569FC8DBE4E" box="[636,877,1548,1572]" italics="true" pageId="5" pageNumber="325">Oryzias chenglongensis</emphasis>
</taxonomicName>
, holotype (above) and
<taxonomicName id="4C6D14C50B1BFFF6BD5AE569FAFEBE4E" baseAuthorityName="Nichols &amp; Pope" baseAuthorityYear="1927" box="[1118,1310,1549,1572]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="5" pageNumber="325" phylum="Chordata" rank="species" species="curvinotus">
<emphasis id="B919B3540B1BFFF6BD5AE569FAFEBE4E" box="[1118,1310,1549,1572]" italics="true" pageId="5" pageNumber="325">Oryzias curvinotus</emphasis>
</taxonomicName>
(below).
</paragraph>
</caption>
<paragraph id="8BD26F460B1BFFF6B9C3E512FE96BF4A" blockId="5.[151,1437,1654,1897]" pageId="5" pageNumber="325">Generally, all fins and belly colored pale white to pale yellow. Caudal fin with yellowish dorsal and ventral submarginal stripes and the stripes darker with melanophores as well as distal yellowish margin in male. Dorsal edge of dorsal and pectoral fins, distal region of pelvic and anal fins with yellow bands. Entire pelvic fin yellowish in male. Pelvic and anal fin with widely scattered tiny melanophores. Caudal fin membrane with about 4 hairline longitudinal stripes.</paragraph>
<paragraph id="8BD26F460B1BFFF6B9C3E44EFAD1BF02" blockId="5.[151,1437,1654,1897]" pageId="5" pageNumber="325">Coloration while specimens in preservative similar above and all brilliant coloration and translucent part entirely faded, body generally light yellowish brown except the melanophores and some larger deep black marks.</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B18FFF5B993E3F3FEA5BA73" pageId="6" pageNumber="326" type="distribution">
<paragraph id="8BD26F460B18FFF5B993E3F3FE33B8DB" blockId="6.[151,467,151,177]" box="[151,467,151,177]" pageId="6" pageNumber="326">
<emphasis id="B919B3540B18FFF5B993E3F3FE33B8DB" bold="true" box="[151,467,151,177]" pageId="6" pageNumber="326">Distribution and habitats</emphasis>
</paragraph>
<paragraph id="8BD26F460B18FFF5B993E3BBFAFAB92B" blockId="6.[151,1437,223,537]" pageId="6" pageNumber="326">
Although the
<typeStatus id="54D6D1E40B18FFF5B834E384FE80B890" box="[304,352,224,250]" pageId="6" pageNumber="326">type</typeStatus>
specimens of
<taxonomicName id="4C6D14C50B18FFF5B8FAE384FCE9B893" authorityName="Chen &amp; Lai" authorityYear="2024" box="[510,777,223,250]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="6" pageNumber="326" phylum="Chordata" rank="species" species="chenglongensis">
<emphasis id="B919B3540B18FFF5B8FAE384FCE9B893" box="[510,777,223,250]" italics="true" pageId="6" pageNumber="326">Oryzias chenglongensis</emphasis>
</taxonomicName>
eventually described and collected from the
<collectingRegion id="49A9A1A40B18FFF5BDFDE3BBFA7CB893" box="[1273,1436,223,249]" country="Taiwan" name="Yunlin" pageId="6" pageNumber="326">Yunlin County</collectingRegion>
of western
<collectingCountry id="F37A2FD60B18FFF5B812E267FE8BB977" box="[278,363,259,285]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
, it can be also found from Chairyi and
<collectingRegion id="49A9A1A40B18FFF5BA28E267FC98B977" box="[812,888,259,285]" country="Taiwan" name="Tainan" pageId="6" pageNumber="326">Tainan</collectingRegion>
by some photo image evidences of local people without well preserved specimens. It is highly possible to be an endemic species around
<collectingCountry id="F37A2FD60B18FFF5BD72E243FB28B92B" box="[1142,1224,295,321]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
island.
</paragraph>
<paragraph id="8BD26F460B18FFF5B9C3E228FEA5BA73" blockId="6.[151,1437,223,537]" pageId="6" pageNumber="326">
It can be considered that the brackish ricefish might be existed in the wetlands of western
<collectingCountry id="F37A2FD60B18FFF5BDE2E22FFAD8B90F" box="[1254,1336,331,357]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
(at least ranging from
<collectingRegion id="49A9A1A40B18FFF5B831E20BFE60B9E3" box="[309,384,367,393]" country="Taiwan" name="Yunlin" pageId="6" pageNumber="326">Yunlin</collectingRegion>
southward to Taiwan) although the current population may be very low and rare. This new species merely collected from brackish habitat around 2030 for salinity. The recorded localities from both Chiaryi and
<collectingRegion id="49A9A1A40B18FFF5B9C3E2D3FEF3B9BB" box="[199,275,439,465]" country="Taiwan" name="Tainan" pageId="6" pageNumber="326">Tainan</collectingRegion>
are also the brackish habitats of wetlands and estuaries. In our breeding experiment, this fish can also survive well in fresh water, it might be more widely distributed around the lowland freshwater to brackish region around
<collectingCountry id="F37A2FD60B18FFF5B9EFE29BFEA1BA73" box="[235,321,511,537]" name="Taiwan" pageId="6" pageNumber="326">Taiwan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3773CCD0B18FFF5B993E10FFEAEBB7F" pageId="6" pageNumber="326" type="etymology">
<paragraph id="8BD26F460B18FFF5B993E10FFEFEBAEF" blockId="6.[151,286,619,645]" box="[151,286,619,645]" pageId="6" pageNumber="326">
<emphasis id="B919B3540B18FFF5B993E10FFEFEBAEF" bold="true" box="[151,286,619,645]" pageId="6" pageNumber="326">Etymology</emphasis>
</paragraph>
<paragraph id="8BD26F460B18FFF5B993E1D7FEAEBB7F" blockId="6.[151,1436,691,790]" pageId="6" pageNumber="326">
The specific name,
<taxonomicName id="4C6D14C50B18FFF5B882E1D7FDD3BAA7" authorityName="Chen &amp; Lai" authorityYear="2024" box="[390,563,691,717]" family="Adrianichthyidae" genus="Oryzias" kingdom="Animalia" order="Beloniformes" pageId="6" pageNumber="326" phylum="Chordata" rank="species" species="chenglongensis">
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, is referred to names of the fish
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locality collected “Chonglong wetland”, western
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although its available geographical range may from Yun-lin, Chiar-yi, to Tai-nan of western
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.
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<emphasis id="B919B3540B18FFF5B993E003FEDFBBEB" bold="true" box="[151,319,871,897]" pageId="6" pageNumber="326">Conservation</emphasis>
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Due the very rare of current new species in wild survey in western
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, we only kept very limited individuals into
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series. Its habitat has also faced the great ecological threat from at least two invasive species of American killifishes. Almost other fish individuals of new species have been kept for conservation reason and reproduce their further off spring and save its limited stock at NTOU,
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for avoiding great aquatic ecological threat from this endangered species becoming extinction.
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<taxonomicName id="AEC14D03FFD4FF801AA8FEBA75134910" ID-CoL="6R3YW" authority="La Cepede, 1801: 416" authorityName="La Cepede" authorityPageNumber="416" authorityYear="1801" box="[654,1148,291,316]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="329" phylum="Chordata" rank="species" species="viridiauratus">
<emphasis id="5BB5EA92FFD4FF801AA8FEBA72F84917" box="[654,919,292,316]" italics="true" pageId="1" pageNumber="329">Macropodus viridiauratus</emphasis>
<bibRefCitation id="0D504B71FFD4FF801BBBFEBA75134910" author="La Cepede, E." box="[925,1148,291,316]" pageId="1" pageNumber="329" refId="ref2269" refString="La Cepede, E. (1801) Histoire naturelle des poisson s; tome 3. Plassan, Paris, 558 pp." type="book" year="1801">La Cepède, 1801: 416</bibRefCitation>
</taxonomicName>
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, 417, by monotypy). Gender masculine.
</paragraph>
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</document>

View file

@ -0,0 +1,476 @@
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<mods:title id="9228E891AF9B9A17C7E0F923AE4A5DBC">A new species of Macropodus (Teleotsei: Osphronemidae) from Fujian Province, southern China</mods:title>
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<taxonomicName id="AEC14D03FFD4FF8018B1FE3270BE49ED" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[151,465,428,455]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="329" phylum="Chordata" rank="species" species="minnanensis" status="sp. nov.">
<emphasis id="5BB5EA92FFD4FF8018B1FE3270BE49ED" bold="true" box="[151,465,428,455]" italics="true" pageId="1" pageNumber="329">Macropodus minnanensis</emphasis>
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<taxonomicNameLabel id="408657E9FFD4FF8019FEFE33737449EC" box="[472,539,429,455]" pageId="1" pageNumber="329" rank="species">n. sp.</taxonomicNameLabel>
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<subSubSection id="21DB650BFFD4FF8018B1FE4E707D49C0" box="[151,274,464,491]" pageId="1" pageNumber="329" type="description">
<paragraph id="697E3680FFD4FF8018B1FE4E707D49C0" blockId="1.[151,539,428,491]" box="[151,274,464,491]" pageId="1" pageNumber="329">
(
<figureCitation id="F1FA2A05FFD4FF8018B9FE4E706649C0" box="[159,265,464,491]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="1.[152,255,1837,1861]" captionStartId-1="2.[152,255,796,820]" captionTargetBox-0="[151,1436,1306,1784]" captionTargetBox-1="[152,1436,198,772]" captionTargetId-0="figure-214@1.[151,1436,1306,1813]" captionTargetId-1="figure-19@2.[151,1436,181,772]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionText-0="FIGURE 1. Macropodus minnanensis, paratype, male, 45.7 mm SL, Pinhe, Joulongjiang basin, Jangjou, Fujian Province, China." captionText-1="FIGURE 2. Macropodus minnanensis, holotype, female, mm SL, female, 34.7 mm SL, Jangjiang basin, Jangjou, Fujian Province, China." figureDoi-0="http://doi.org/10.5281/zenodo.14389378" figureDoi-1="http://doi.org/10.5281/zenodo.14389380" httpUri-0="https://zenodo.org/record/14389378/files/figure.png" httpUri-1="https://zenodo.org/record/14389380/files/figure.png" pageId="1" pageNumber="329">Figs. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="21DB650BFFD4FF8018B1FD8670CD4BB1" pageId="1" pageNumber="329" type="materials_examined">
<paragraph id="697E3680FFD4FF8018B1FD8670FF4A19" blockId="1.[151,400,536,562]" box="[151,400,536,562]" pageId="1" pageNumber="329">
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<emphasis id="5BB5EA92FFD4FF8018B1FD8670FF4A19" bold="true" box="[151,400,536,562]" pageId="1" pageNumber="329">Materials examined</emphasis>
</heading>
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<paragraph id="697E3680FFD4FF8018B1FDFF70CD4AB5" blockId="1.[151,1436,608,670]" pageId="1" pageNumber="329">
<emphasis id="5BB5EA92FFD4FF8018B1FDFF704F4A50" bold="true" box="[151,288,609,635]" pageId="1" pageNumber="329">
<typeStatus id="B67A8822FFD4FF8018B1FDFF706B4A50" box="[151,260,609,635]" pageId="1" pageNumber="329" type="holotype">Holotype</typeStatus>
</emphasis>
NTOUP-2023-12-255,
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SL, coll. K.Y. Chen
<emphasis id="5BB5EA92FFD4FF801B53FDFF72C14A51" box="[885,942,608,634]" italics="true" pageId="1" pageNumber="329">et al.</emphasis>
, 23 Spt. 2023, Janjiang basin, Jangjou City,
<collectingRegion id="AB05F862FFD4FF8018B1FD1A70234AB5" box="[151,332,644,670]" country="China" name="Fujian" pageId="1" pageNumber="329">Fujian Province</collectingRegion>
,
<collectingCountry id="11D67610FFD4FF801971FD1A70F14AB5" box="[343,414,644,670]" name="China" pageId="1" pageNumber="329">China</collectingCountry>
.
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<typeStatus id="B67A8822FFD4FF8018B1FD6F707B4B20" box="[151,276,753,779]" pageId="1" pageNumber="329" type="paratype">Paratypes</typeStatus>
</emphasis>
</heading>
</paragraph>
<paragraph id="697E3680FFD4FF8018B1FCA675584B79" blockId="1.[151,1436,824,922]" box="[151,1079,824,851]" pageId="1" pageNumber="329">
NTOUP-2023-12-256,
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SL, the date and collection data same as
<typeStatus id="B67A8822FFD4FF801BEBFCA6755C4B79" box="[973,1075,824,850]" pageId="1" pageNumber="329" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="697E3680FFD4FF8018E1FCC270CD4BB1" blockId="1.[151,1436,824,922]" pageId="1" pageNumber="329">
NTOUP-2015-03-201,
<quantity id="AE399B65FFD4FF8019E9FCC273594B5C" box="[463,566,860,887]" metricMagnitude="-2" metricUnit="m" metricValue="4.57" pageId="1" pageNumber="329" unit="mm" value="45.7">45.7 mm</quantity>
SL, coll. I-S. Chen
<emphasis id="5BB5EA92FFD4FF801B00FCC3720E4B5D" box="[806,865,860,886]" italics="true" pageId="1" pageNumber="329">et al.</emphasis>
,
<date id="1D7F1040FFD4FF801B54FCC3757F4B5C" box="[882,1040,860,887]" pageId="1" pageNumber="329" value="2015-03-11">11 Mar. 2015</date>
, Joulongjiang basin, Jangjou City,
<collectingRegion id="AB05F862FFD4FF8018B1FC1E70234BB1" box="[151,332,896,922]" country="China" name="Fujian" pageId="1" pageNumber="329">Fujian Province</collectingRegion>
,
<collectingCountry id="11D67610FFD4FF801971FC1E70F14BB1" box="[343,414,896,922]" name="China" pageId="1" pageNumber="329">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="21DB650BFFD4FF8018B1FC7270114D29" pageId="1" pageNumber="329" type="diagnosis">
<paragraph id="697E3680FFD4FF8018B1FC72707F4C2D" blockId="1.[151,272,1004,1030]" box="[151,272,1004,1030]" pageId="1" pageNumber="329">
<emphasis id="5BB5EA92FFD4FF8018B1FC72707F4C2D" bold="true" box="[151,272,1004,1030]" pageId="1" pageNumber="329">Diagnosis</emphasis>
</paragraph>
<paragraph id="697E3680FFD4FF8018B1FBAA70114D29" blockId="1.[151,1437,1076,1283]" pageId="1" pageNumber="329">
The new species
<taxonomicName id="AEC14D03FFD4FF801971FBAA73194C65" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[343,630,1076,1102]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="329" phylum="Chordata" rank="species" species="minnanensis">
<emphasis id="5BB5EA92FFD4FF801971FBAA73194C65" box="[343,630,1076,1102]" italics="true" pageId="1" pageNumber="329">Macropodus minnanensis</emphasis>
</taxonomicName>
, from
<collectingRegion id="AB05F862FFD4FF801AE7FBAA72194C65" box="[705,886,1076,1102]" country="China" name="Fujian" pageId="1" pageNumber="329">Fujian Province</collectingRegion>
,
<collectingCountry id="11D67610FFD4FF801BA7FBAA72AB4C65" box="[897,964,1076,1102]" name="China" pageId="1" pageNumber="329">China</collectingCountry>
can be well distinguished from congeners by the following combination of features: (1) dorsal fin with 13 spines and 7 soft rays; (2) anal fin with 20 spines and modally 10 soft rays; (3) vertical insertion of dorsal fin origin at second to third soft rays of anal fin; (4) longitudinal scale rows in midline modally 28; (5) transverse scale rows 14; (6) vertebrae 27 and (7) specific coloration: body tawny to blackish brown with posteriorly black margin on scale pockets; eye greyish; a large opecular, deep black ocellus spot in male.
</paragraph>
</subSubSection>
<caption id="3DBE6608FFD4FF8018BEF8B371B44F42" ID-DOI="http://doi.org/10.5281/zenodo.14389378" ID-Zenodo-Dep="14389378" httpUri="https://zenodo.org/record/14389378/files/figure.png" pageId="1" pageNumber="329" startId="1.[152,255,1837,1861]" targetBox="[151,1436,1306,1784]" targetPageId="1" targetType="figure">
<paragraph id="697E3680FFD4FF8018BEF8B371B44F42" blockId="1.[151,1437,1837,1897]" pageId="1" pageNumber="329">
<emphasis id="5BB5EA92FFD4FF8018BEF8B3707A4F6E" bold="true" box="[152,277,1837,1861]" pageId="1" pageNumber="329">FIGURE 1</emphasis>
.
<taxonomicName id="AEC14D03FFD4FF801903F8B373424F6E" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[293,557,1837,1861]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="329" phylum="Chordata" rank="species" species="minnanensis">
<emphasis id="5BB5EA92FFD4FF801903F8B373424F6E" box="[293,557,1837,1861]" italics="true" pageId="1" pageNumber="329">Macropodus minnanensis</emphasis>
</taxonomicName>
, paratype, male, 45.7 mm SL, Pinhe, Joulongjiang basin, Jangjou, Fujian Province, China.
</paragraph>
</caption>
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<paragraph id="697E3680FFD7FF8318BEFC82702D4B73" blockId="2.[151,1436,796,856]" pageId="2" pageNumber="330">
<emphasis id="5BB5EA92FFD7FF8318BEFC8270794B1F" bold="true" box="[152,278,796,820]" pageId="2" pageNumber="330">FIGURE 2</emphasis>
.
<taxonomicName id="AEC14D03FFD7FF831901FC8273404B1F" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[295,559,796,820]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="330" phylum="Chordata" rank="species" species="minnanensis">
<emphasis id="5BB5EA92FFD7FF831901FC8273404B1F" box="[295,559,796,820]" italics="true" pageId="2" pageNumber="330">Macropodus minnanensis</emphasis>
</taxonomicName>
, holotype, female, mm SL, female, 34.7 mm SL, Jangjiang basin, Jangjou, Fujian Province, China.
</paragraph>
</caption>
<subSubSection id="21DB650BFFD7FF8318B1FC34737A4F47" pageId="2" pageNumber="330" type="description">
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<emphasis id="5BB5EA92FFD7FF8318B1FC3470474BEF" bold="true" box="[151,296,938,964]" pageId="2" pageNumber="330">Description</emphasis>
</paragraph>
<paragraph id="697E3680FFD7FF8318B1FC6C75594CB7" blockId="2.[151,1437,1010,1433]" pageId="2" pageNumber="330">
The all relevant morphometric data of
<specimenCount id="7FC7FD09FFD7FF831A6DFC6C73AD4C27" box="[587,706,1010,1036]" count="1" pageId="2" pageNumber="330" type="generic" typeStatus="holotype">1 holotype</specimenCount>
and
<specimenCount id="7FC7FD09FFD7FF831ADCFC6C72154C26" box="[762,890,1010,1037]" count="2" pageId="2" pageNumber="330" type="generic" typeStatus="paratype">2 paratypes</specimenCount>
see in
<tableCitation id="2443033BFFD7FF831BECFC6C75704C27" box="[970,1055,1010,1036]" captionStart="TABLE 1" captionStartId="3.[152,245,152,176]" captionTargetPageId="3" captionText="TABLE 1. Morphometric data of Macropodus minnanensis n. sp. from Fujian, China." pageId="2" pageNumber="330">Table 1</tableCitation>
. Medium size osphronemids with deep, elongate and strong laterally compressed body. Higher body depth at anal fin origin. Snout short but laterally compressed. Body completely covered ctenoid scales. Lateral line absent. Lateral midline scales 2829 (modally 28); 14 transverse scales. 1217 gill rackers on ceratobranchial of first arch. 23 rows of teeth on upper jaws, 34 rows of lower jaws. Larger recorded specimen as
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SL so far. Vertebrae 27.
</paragraph>
<paragraph id="697E3680FFD7FF8318E1FB38743A4DB3" blockId="2.[151,1437,1010,1433]" pageId="2" pageNumber="330">Dorsal fin with 13 spines and 7 branched and unbranched soft rays (totally 20 spines and rays). Bases of unpaired fins scaled. Length of anterior 34 rays increasing, then decreasing to the last ray. Anal fin with 20 spines and 10 branched and unbranched soft rays (totally 30 spines and rays). Length of anterior 78 soft rays increasing, then decreasing to last rays. Soft dorsal rays 34 and soft anal rays 78 out margin pointed. Dorsal and anal fin pointed posteriorly, extending far beyond the origin of caudal fin base. Caudal fin emarginate in female to deeply forked in male, with 1213 branched rays. Pectoral fin 1011 rays with rounded margin, its rear tip reaching vertical of 35 dorsal spines. Pelvic fin pointed with 1 spine and 5 soft rays. First of soft pelvic-fin rays filamentous.</paragraph>
<paragraph id="697E3680FFD7FF8318B1FA7473644E2F" blockId="2.[151,523,1514,1540]" box="[151,523,1514,1540]" pageId="2" pageNumber="330">
<heading id="323681ECFFD7FF8318B1FA7473644E2F" bold="true" box="[151,523,1514,1540]" fontSize="11" level="1" pageId="2" pageNumber="330" reason="1">
<emphasis id="5BB5EA92FFD7FF8318B1FA7473644E2F" bold="true" box="[151,523,1514,1540]" pageId="2" pageNumber="330">Coloration in fresh specimens</emphasis>
</heading>
</paragraph>
<paragraph id="697E3680FFD7FF8318B1F9AC75694E93" blockId="2.[151,1437,1586,1973]" pageId="2" pageNumber="330">Body and head with tawny to blackish brown background. Lateral body scales with net-like greyish to black margin. Most of poster, outer margin with a black spot.A large deep black, round ocellus spot with a very narrow, posteriorly bright orange to golden rim. A greyish black bar in between eye and the large spot. Two greyish black blotches above the bar. Eye greyish outside pupil. Snout with 23 greyish black stripes.</paragraph>
<paragraph id="697E3680FFD7FF8318E1F95C737A4F47" blockId="2.[151,1437,1586,1973]" pageId="2" pageNumber="330">Dorsal fin pale brown and 23 rows of deep brown to blackish brown spots on spinous region and 68 rows of deep brown to blackish brown spots on soft ray region.Anal fin pale brown with 12 rows of deep brown to blackish brown spots on spinous region and 35 rows of deep brown to blackish brown spots on soft rays region. Caudal fin pale brown with 915 rows of deep brown or blackish brown spots. Pectoral fin pale white and spotless. Pelvic fin pale white to light creamy yellow.</paragraph>
</subSubSection>
<subSubSection id="21DB650BFFD7FF8218E1F8E875A24F0C" lastPageId="3" lastPageNumber="331" pageId="2" pageNumber="330" type="etymology">
<paragraph id="697E3680FFD7FF8318E1F8E872724F9F" blockId="2.[151,1437,1586,1973]" pageId="2" pageNumber="330">
<emphasis id="5BB5EA92FFD7FF8318E1F8E870064FBB" bold="true" box="[199,361,1910,1936]" pageId="2" pageNumber="330">Etymology.—</emphasis>
The specific name,
<taxonomicName id="AEC14D03FFD7FF831A69F8E973B44FBB" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[591,731,1911,1936]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="330" phylum="Chordata" rank="species" species="minnanensis">
<emphasis id="5BB5EA92FFD7FF831A69F8E973B44FBB" box="[591,731,1911,1936]" italics="true" pageId="2" pageNumber="330">minnanensis</emphasis>
</taxonomicName>
, is referred to its own
<typeStatus id="B67A8822FFD7FF831BCDF8E975744FBA" box="[1003,1051,1911,1937]" pageId="2" pageNumber="330">type</typeStatus>
locality in Mandarin: “Min-nan” which means the southern part of
<collectingRegion id="AB05F862FFD7FF831A35F80473A84F9F" box="[531,711,1946,1972]" country="China" name="Fujian" pageId="2" pageNumber="330">Fujian Province</collectingRegion>
,
<collectingCountry id="11D67610FFD7FF831AF5F80472754F9F" box="[723,794,1946,1972]" name="China" pageId="2" pageNumber="330">China</collectingCountry>
.
</paragraph>
<caption id="3DBE6608FFD6FF8218BEFF06755A489B" box="[152,1077,152,177]" pageId="3" pageNumber="331" startId="3.[152,245,152,176]" targetBox="[163,1329,193,1223]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="697E3680FFD6FF8218BEFF06755A489B" blockId="3.[152,1077,152,177]" box="[152,1077,152,177]" pageId="3" pageNumber="331">
<emphasis id="5BB5EA92FFD6FF8218BEFF067060489B" bold="true" box="[152,271,152,176]" pageId="3" pageNumber="331">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="AEC14D03FFD6FF821A20FF067277489B" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[518,792,152,176]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="331" phylum="Chordata" rank="species" species="minnanensis" status="sp. nov.">
<emphasis id="5BB5EA92FFD6FF821A20FF067277489B" box="[518,792,152,176]" italics="true" pageId="3" pageNumber="331">Macropodus minnanensis</emphasis>
</taxonomicName>
<emphasis id="5BB5EA92FFD6FF821B39FF077234489A" bold="true" box="[799,859,153,177]" pageId="3" pageNumber="331">
<taxonomicNameLabel id="408657E9FFD6FF821B39FF077234489A" box="[799,859,153,177]" pageId="3" pageNumber="331" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
from Fujian, China.
</paragraph>
</caption>
<paragraph id="697E3680FFD6FF821AF5FF5F745E4CEC" pageId="3" pageNumber="331">
<table id="1BC1C420FFD6007E1885FF5F745E4CEC" box="[163,1329,193,1223]" gridcols="4" gridrows="26" pageId="3" pageNumber="331">
<tr id="D7F134C2FFD6007E1885FF5F745E48F2" box="[163,1329,193,217]" gridrow="0" pageId="3" pageNumber="331" rowspan-0="1">
<th id="94205DBEFFD6007E1AEAFF5F726148F2" box="[716,782,193,217]" gridcol="1" gridrow="0" pageId="3" pageNumber="331">Male</th>
<th id="94205DBEFFD6007E1BF2FF5F757048F2" box="[980,1055,193,217]" gridcol="2" gridrow="0" pageId="3" pageNumber="331">Female</th>
<th id="94205DBEFFD6007E1CC9FF5F745E48F2" box="[1263,1329,193,217]" gridcol="3" gridrow="0" pageId="3" pageNumber="331">Male</th>
</tr>
<tr id="D7F134C2FFD6007E1885FF77745E492A" box="[163,1329,233,257]" gridrow="1" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FF777097492A" box="[163,504,233,257]" gridcol="0" gridrow="1" pageId="3" pageNumber="331">SL(mm)</th>
<td id="94205DBEFFD6007E1AEAFF777261492A" box="[716,782,233,257]" gridcol="1" gridrow="1" pageId="3" pageNumber="331">45.7</td>
<td id="94205DBEFFD6007E1BF2FF777570492A" box="[980,1055,233,257]" gridcol="2" gridrow="1" pageId="3" pageNumber="331">34.7</td>
<td id="94205DBEFFD6007E1CC9FF77745E492A" box="[1263,1329,233,257]" gridcol="3" gridrow="1" pageId="3" pageNumber="331">38.4</td>
</tr>
<tr id="D7F134C2FFD6007E1885FE8F745E4902" box="[163,1329,273,297]" gridrow="2" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FE8F745E4902" box="[163,1329,273,297]" colspan="4" colspanRight="3" gridcol="0" gridrow="2" pageId="3" pageNumber="331">In % of SL</th>
</tr>
<tr id="D7F134C2FFD6007E1885FEA7745E4979" box="[163,1329,313,338]" gridrow="3" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FEA770974979" box="[163,504,313,338]" gridcol="0" gridrow="3" pageId="3" pageNumber="331">Head length (HL)</th>
<td id="94205DBEFFD6007E1AEAFEA772614979" box="[716,782,313,338]" gridcol="1" gridrow="3" pageId="3" pageNumber="331">27.8%</td>
<td id="94205DBEFFD6007E1BF2FEA775704979" box="[980,1055,313,338]" gridcol="2" gridrow="3" pageId="3" pageNumber="331">33.3%</td>
<td id="94205DBEFFD6007E1CC9FEA7745E4979" box="[1263,1329,313,338]" gridcol="3" gridrow="3" pageId="3" pageNumber="331">33.3%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FEFF745E4951" box="[163,1329,353,378]" gridrow="4" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FEFF70974951" box="[163,504,353,378]" gridcol="0" gridrow="4" pageId="3" pageNumber="331">Predorsal length</th>
<td id="94205DBEFFD6007E1AEAFEFF72614951" box="[716,782,353,378]" gridcol="1" gridrow="4" pageId="3" pageNumber="331">52.3%</td>
<td id="94205DBEFFD6007E1BF2FEFF75704951" box="[980,1055,353,378]" gridcol="2" gridrow="4" pageId="3" pageNumber="331">54.0%</td>
<td id="94205DBEFFD6007E1CC9FEFF745E4951" box="[1263,1329,353,378]" gridcol="3" gridrow="4" pageId="3" pageNumber="331">51.1%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FE14745E4989" box="[163,1329,394,418]" gridrow="5" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FE1470974989" box="[163,504,394,418]" gridcol="0" gridrow="5" pageId="3" pageNumber="331">Prepelvic length</th>
<td id="94205DBEFFD6007E1AEAFE1472614989" box="[716,782,394,418]" gridcol="1" gridrow="5" pageId="3" pageNumber="331">39.7%</td>
<td id="94205DBEFFD6007E1BF2FE1475704989" box="[980,1055,394,418]" gridcol="2" gridrow="5" pageId="3" pageNumber="331">38.9%</td>
<td id="94205DBEFFD6007E1CC9FE14745E4989" box="[1263,1329,394,418]" gridcol="3" gridrow="5" pageId="3" pageNumber="331">36.6%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FE2C745E49E1" box="[163,1329,434,458]" gridrow="6" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FE2C709749E1" box="[163,504,434,458]" gridcol="0" gridrow="6" pageId="3" pageNumber="331">Preanal length</th>
<td id="94205DBEFFD6007E1AEAFE2C726149E1" box="[716,782,434,458]" gridcol="1" gridrow="6" pageId="3" pageNumber="331">50.4%</td>
<td id="94205DBEFFD6007E1BF2FE2C757049E1" box="[980,1055,434,458]" gridcol="2" gridrow="6" pageId="3" pageNumber="331">49.2%</td>
<td id="94205DBEFFD6007E1CC9FE2C745E49E1" box="[1263,1329,434,458]" gridcol="3" gridrow="6" pageId="3" pageNumber="331">3.0%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FE44745E49D8" box="[163,1329,474,499]" gridrow="7" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FE44709749D8" box="[163,504,474,499]" gridcol="0" gridrow="7" pageId="3" pageNumber="331">Head depth in front of eye</th>
<td id="94205DBEFFD6007E1AEAFE44726149D8" box="[716,782,474,499]" gridcol="1" gridrow="7" pageId="3" pageNumber="331">14.0%</td>
<td id="94205DBEFFD6007E1BF2FE44757049D8" box="[980,1055,474,499]" gridcol="2" gridrow="7" pageId="3" pageNumber="331">13.9%</td>
<td id="94205DBEFFD6007E1CC9FE44745E49D8" box="[1263,1329,474,499]" gridcol="3" gridrow="7" pageId="3" pageNumber="331">44.9%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FD9D745E4A30" box="[163,1329,515,539]" gridrow="8" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FD9D70974A30" box="[163,504,515,539]" gridcol="0" gridrow="8" pageId="3" pageNumber="331">Head depth behind eye</th>
<td id="94205DBEFFD6007E1AEAFD9D72614A30" box="[716,782,515,539]" gridcol="1" gridrow="8" pageId="3" pageNumber="331">21.2%</td>
<td id="94205DBEFFD6007E1BF2FD9D75704A30" box="[980,1055,515,539]" gridcol="2" gridrow="8" pageId="3" pageNumber="331">23.0%</td>
<td id="94205DBEFFD6007E1CC9FD9D745E4A30" box="[1263,1329,515,539]" gridcol="3" gridrow="8" pageId="3" pageNumber="331">21.7%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FDB5745E4A68" box="[163,1329,555,579]" gridrow="9" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FDB570974A68" box="[163,504,555,579]" gridcol="0" gridrow="9" pageId="3" pageNumber="331">Head depth at nape</th>
<td id="94205DBEFFD6007E1AEAFDB572614A68" box="[716,782,555,579]" gridcol="1" gridrow="9" pageId="3" pageNumber="331">26.6%</td>
<td id="94205DBEFFD6007E1BF2FDB575704A68" box="[980,1055,555,579]" gridcol="2" gridrow="9" pageId="3" pageNumber="331">24.7%</td>
<td id="94205DBEFFD6007E1CC9FDB5745E4A68" box="[1263,1329,555,579]" gridcol="3" gridrow="9" pageId="3" pageNumber="331">24.7%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FDCD745E4A40" box="[163,1329,595,619]" gridrow="10" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FDCD70974A40" box="[163,504,595,619]" gridcol="0" gridrow="10" pageId="3" pageNumber="331">Head width at operculum</th>
<td id="94205DBEFFD6007E1AEAFDCD72614A40" box="[716,782,595,619]" gridcol="1" gridrow="10" pageId="3" pageNumber="331">20.5%</td>
<td id="94205DBEFFD6007E1BF2FDCD75704A40" box="[980,1055,595,619]" gridcol="2" gridrow="10" pageId="3" pageNumber="331">10.9%</td>
<td id="94205DBEFFD6007E1CC9FDCD745E4A40" box="[1263,1329,595,619]" gridcol="3" gridrow="10" pageId="3" pageNumber="331">18.1%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FDE5745E4AB8" box="[163,1329,635,659]" gridrow="11" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FDE570974AB8" box="[163,504,635,659]" gridcol="0" gridrow="11" pageId="3" pageNumber="331">Body depth at anal-fin origin</th>
<td id="94205DBEFFD6007E1AEAFDE572614AB8" box="[716,782,635,659]" gridcol="1" gridrow="11" pageId="3" pageNumber="331">37.6%</td>
<td id="94205DBEFFD6007E1BF2FDE575704AB8" box="[980,1055,635,659]" gridcol="2" gridrow="11" pageId="3" pageNumber="331">32.4%</td>
<td id="94205DBEFFD6007E1CC9FDE5745E4AB8" box="[1263,1329,635,659]" gridcol="3" gridrow="11" pageId="3" pageNumber="331">33.8%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FD3A745E4A97" box="[163,1329,676,700]" gridrow="12" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FD3A70974A97" box="[163,504,676,700]" gridcol="0" gridrow="12" pageId="3" pageNumber="331">Body width at dorsal-fin origin</th>
<td id="94205DBEFFD6007E1AEAFD3A72614A97" box="[716,782,676,700]" gridcol="1" gridrow="12" pageId="3" pageNumber="331">14.3%</td>
<td id="94205DBEFFD6007E1BF2FD3A75704A97" box="[980,1055,676,700]" gridcol="2" gridrow="12" pageId="3" pageNumber="331">13.3%</td>
<td id="94205DBEFFD6007E1CC9FD3A745E4A97" box="[1263,1329,676,700]" gridcol="3" gridrow="12" pageId="3" pageNumber="331">13.9%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FD52745E4ACF" box="[163,1329,716,740]" gridrow="13" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FD5270974ACF" box="[163,504,716,740]" gridcol="0" gridrow="13" pageId="3" pageNumber="331">Body depth at dorsal-fin base end</th>
<td id="94205DBEFFD6007E1AEAFD5272614ACF" box="[716,782,716,740]" gridcol="1" gridrow="13" pageId="3" pageNumber="331">25.8%</td>
<td id="94205DBEFFD6007E1BF2FD5275704ACF" box="[980,1055,716,740]" gridcol="2" gridrow="13" pageId="3" pageNumber="331">17.7%</td>
<td id="94205DBEFFD6007E1CC9FD52745E4ACF" box="[1263,1329,716,740]" gridcol="3" gridrow="13" pageId="3" pageNumber="331">20.7%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FD6A745E4B27" box="[163,1329,756,780]" gridrow="14" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FD6A70974B27" box="[163,504,756,780]" gridcol="0" gridrow="14" pageId="3" pageNumber="331">Postdorsal length</th>
<td id="94205DBEFFD6007E1AEAFD6A72614B27" box="[716,782,756,780]" gridcol="1" gridrow="14" pageId="3" pageNumber="331">51.9%</td>
<td id="94205DBEFFD6007E1BF2FD6A75704B27" box="[980,1055,756,780]" gridcol="2" gridrow="14" pageId="3" pageNumber="331">51.0%</td>
<td id="94205DBEFFD6007E1CC9FD6A745E4B27" box="[1263,1329,756,780]" gridcol="3" gridrow="14" pageId="3" pageNumber="331">49.4%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FC82745E4B1F" box="[163,1329,796,820]" gridrow="15" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FC8270974B1F" box="[163,504,796,820]" gridcol="0" gridrow="15" pageId="3" pageNumber="331">Length of dorsal fin base</th>
<td id="94205DBEFFD6007E1AEAFC8272614B1F" box="[716,782,796,820]" gridcol="1" gridrow="15" pageId="3" pageNumber="331">38.9%</td>
<td id="94205DBEFFD6007E1BF2FC8275704B1F" box="[980,1055,796,820]" gridcol="2" gridrow="15" pageId="3" pageNumber="331">39.1%</td>
<td id="94205DBEFFD6007E1CC9FC82745E4B1F" box="[1263,1329,796,820]" gridcol="3" gridrow="15" pageId="3" pageNumber="331">36.7%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FCDA745E4B76" box="[163,1329,836,861]" gridrow="16" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FCDA70974B76" box="[163,504,836,861]" gridcol="0" gridrow="16" pageId="3" pageNumber="331">Length of anal fin base</th>
<td id="94205DBEFFD6007E1AEAFCDA72614B76" box="[716,782,836,861]" gridcol="1" gridrow="16" pageId="3" pageNumber="331">51.1%</td>
<td id="94205DBEFFD6007E1BF2FCDA75704B76" box="[980,1055,836,861]" gridcol="2" gridrow="16" pageId="3" pageNumber="331">52.8%</td>
<td id="94205DBEFFD6007E1CC9FCDA745E4B76" box="[1263,1329,836,861]" gridcol="3" gridrow="16" pageId="3" pageNumber="331">47.0%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FCF3745E4BAE" box="[163,1329,877,901]" gridrow="17" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FCF370974BAE" box="[163,504,877,901]" gridcol="0" gridrow="17" pageId="3" pageNumber="331">Length of pectoral fin</th>
<td id="94205DBEFFD6007E1AEAFCF372614BAE" box="[716,782,877,901]" gridcol="1" gridrow="17" pageId="3" pageNumber="331">23.4%</td>
<td id="94205DBEFFD6007E1BF2FCF375704BAE" box="[980,1055,877,901]" gridcol="2" gridrow="17" pageId="3" pageNumber="331">26.9%</td>
<td id="94205DBEFFD6007E1CC9FCF3745E4BAE" box="[1263,1329,877,901]" gridcol="3" gridrow="17" pageId="3" pageNumber="331">25.8%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FC0B745E4B86" box="[163,1329,917,941]" gridrow="18" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FC0B70974B86" box="[163,504,917,941]" gridcol="0" gridrow="18" pageId="3" pageNumber="331">Length of pelvic spine</th>
<td id="94205DBEFFD6007E1AEAFC0B72614B86" box="[716,782,917,941]" gridcol="1" gridrow="18" pageId="3" pageNumber="331">11.7%</td>
<td id="94205DBEFFD6007E1BF2FC0B75704B86" box="[980,1055,917,941]" gridcol="2" gridrow="18" pageId="3" pageNumber="331">11.9%</td>
<td id="94205DBEFFD6007E1CC9FC0B745E4B86" box="[1263,1329,917,941]" gridcol="3" gridrow="18" pageId="3" pageNumber="331">13.5%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FC23745E4BFE" box="[163,1329,957,981]" gridrow="19" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FC2370974BFE" box="[163,504,957,981]" gridcol="0" gridrow="19" pageId="3" pageNumber="331">Length of pelvic filament</th>
<td id="94205DBEFFD6007E1AEAFC2372614BFE" box="[716,782,957,981]" gridcol="1" gridrow="19" pageId="3" pageNumber="331">30.3%</td>
<td id="94205DBEFFD6007E1BF2FC2375704BFE" box="[980,1055,957,981]" gridcol="2" gridrow="19" pageId="3" pageNumber="331">37.7%</td>
<td id="94205DBEFFD6007E1CC9FC23745E4BFE" box="[1263,1329,957,981]" gridcol="3" gridrow="19" pageId="3" pageNumber="331">37.2%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FC7B745E4BD5" box="[163,1329,997,1022]" gridrow="20" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FC7B70974BD5" box="[163,504,997,1022]" gridcol="0" gridrow="20" pageId="3" pageNumber="331">Length of middle caudal fin ray</th>
<td id="94205DBEFFD6007E1AEAFC7B72614BD5" box="[716,782,997,1022]" gridcol="1" gridrow="20" pageId="3" pageNumber="331">31.0%</td>
<td id="94205DBEFFD6007E1BF2FC7B75704BD5" box="[980,1055,997,1022]" gridcol="2" gridrow="20" pageId="3" pageNumber="331">37.4%</td>
<td id="94205DBEFFD6007E1CC9FC7B745E4BD5" box="[1263,1329,997,1022]" gridcol="3" gridrow="20" pageId="3" pageNumber="331">32.8%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FB90745E4C0D" box="[163,1329,1038,1062]" gridrow="21" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FB90745E4C0D" box="[163,1329,1038,1062]" colspan="4" colspanRight="3" gridcol="0" gridrow="21" pageId="3" pageNumber="331">In % of HL</th>
</tr>
<tr id="D7F134C2FFD6007E1885FBA8745E4C65" box="[163,1329,1078,1102]" gridrow="22" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FBA870974C65" box="[163,504,1078,1102]" gridcol="0" gridrow="22" pageId="3" pageNumber="331">Eye diameter</th>
<td id="94205DBEFFD6007E1AEAFBA872614C65" box="[716,782,1078,1102]" gridcol="1" gridrow="22" pageId="3" pageNumber="331">24.1%</td>
<td id="94205DBEFFD6007E1BF2FBA875704C65" box="[980,1055,1078,1102]" gridcol="2" gridrow="22" pageId="3" pageNumber="331">31.9%</td>
<td id="94205DBEFFD6007E1CC9FBA8745E4C65" box="[1263,1329,1078,1102]" gridcol="3" gridrow="22" pageId="3" pageNumber="331">25.3%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FBC0745E4C5D" box="[163,1329,1118,1142]" gridrow="23" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FBC070974C5D" box="[163,504,1118,1142]" gridcol="0" gridrow="23" pageId="3" pageNumber="331">Interorbital width</th>
<td id="94205DBEFFD6007E1AEAFBC072614C5D" box="[716,782,1118,1142]" gridcol="1" gridrow="23" pageId="3" pageNumber="331">32.2%</td>
<td id="94205DBEFFD6007E1BF2FBC075704C5D" box="[980,1055,1118,1142]" gridcol="2" gridrow="23" pageId="3" pageNumber="331">29.9%</td>
<td id="94205DBEFFD6007E1CC9FBC0745E4C5D" box="[1263,1329,1118,1142]" gridcol="3" gridrow="23" pageId="3" pageNumber="331">30.8%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FB18745E4CB4" box="[163,1329,1158,1183]" gridrow="24" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FB1870974CB4" box="[163,504,1158,1183]" gridcol="0" gridrow="24" pageId="3" pageNumber="331">Snout length</th>
<td id="94205DBEFFD6007E1AEAFB1872614CB4" box="[716,782,1158,1183]" gridcol="1" gridrow="24" pageId="3" pageNumber="331">22.7%</td>
<td id="94205DBEFFD6007E1BF2FB1875704CB4" box="[980,1055,1158,1183]" gridcol="2" gridrow="24" pageId="3" pageNumber="331">23.1%</td>
<td id="94205DBEFFD6007E1CC9FB18745E4CB4" box="[1263,1329,1158,1183]" gridcol="3" gridrow="24" pageId="3" pageNumber="331">24.1%</td>
</tr>
<tr id="D7F134C2FFD6007E1885FB31745E4CEC" box="[163,1329,1199,1223]" gridrow="25" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FB3170974CEC" box="[163,504,1199,1223]" gridcol="0" gridrow="25" pageId="3" pageNumber="331">Upper lip length</th>
<td id="94205DBEFFD6007E1AEAFB3172614CEC" box="[716,782,1199,1223]" gridcol="1" gridrow="25" pageId="3" pageNumber="331">23.9%</td>
<td id="94205DBEFFD6007E1BF2FB3175704CEC" box="[980,1055,1199,1223]" gridcol="2" gridrow="25" pageId="3" pageNumber="331">23.8%</td>
<td id="94205DBEFFD6007E1CC9FB31745E4CEC" box="[1263,1329,1199,1223]" gridcol="3" gridrow="25" pageId="3" pageNumber="331">23.3%</td>
</tr>
</table>
</paragraph>
<caption id="3DBE6608FFD6FF8218BEFB6F75A54D22" box="[152,1226,1265,1289]" pageId="3" pageNumber="331" startId="3.[152,245,1265,1289]" targetBox="[163,1337,1306,1692]" targetIsTable="true" targetPageId="3" targetType="table">
<paragraph id="697E3680FFD6FF8218BEFB6F75A54D22" blockId="3.[152,1226,1265,1289]" box="[152,1226,1265,1289]" pageId="3" pageNumber="331">
<emphasis id="5BB5EA92FFD6FF8218BEFB6F70604D22" bold="true" box="[152,271,1265,1289]" pageId="3" pageNumber="331">TABLE 2.</emphasis>
Meristic comparison of two species in
<taxonomicName id="AEC14D03FFD6FF821A95FB6C721A4D22" authorityName="Feryhof &amp; Herder" authorityYear="2002" box="[691,885,1265,1289]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="331" phylum="Chordata" rank="species" species="hongkongensis">
<emphasis id="5BB5EA92FFD6FF821A95FB6C721A4D22" box="[691,885,1265,1289]" italics="true" pageId="3" pageNumber="331">M. hongkongensis</emphasis>
</taxonomicName>
complex from mainland China.
</paragraph>
</caption>
<paragraph id="697E3680FFD6FF821885FA84759B4EB7" pageId="3" pageNumber="331">
<table id="1BC1C420FFD6007E1885FA8474564EB7" box="[163,1337,1306,1692]" gridcols="3" gridrows="10" pageId="3" pageNumber="331">
<tr id="D7F134C2FFD6007E1885FA8474564D19" box="[163,1337,1306,1330]" gridrow="0" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FA84737D4D19" box="[163,530,1306,1330]" gridcol="0" gridrow="0" pageId="3" pageNumber="331">Species</th>
<th id="94205DBEFFD6007E1AB5FA84728D4D19" box="[659,994,1306,1330]" gridcol="1" gridrow="0" pageId="3" pageNumber="331">
<taxonomicName id="AEC14D03FFD6FF821AECFA8472044D1A" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[714,875,1306,1329]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="331" phylum="Chordata" rank="species" species="minnanensis" status="n. sp.">
<emphasis id="5BB5EA92FFD6FF821AECFA8472044D1A" box="[714,875,1306,1329]" italics="true" pageId="3" pageNumber="331">M. minnanensis</emphasis>
</taxonomicName>
<emphasis id="5BB5EA92FFD6FF821B57FA8472C54D19" bold="true" box="[881,938,1306,1330]" pageId="3" pageNumber="331">
<taxonomicNameLabel id="408657E9FFD6FF821B57FA8472C54D19" box="[881,938,1306,1330]" pageId="3" pageNumber="331" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</th>
<th id="94205DBEFFD6007E1C56FA8474564D19" box="[1136,1337,1306,1330]" gridcol="2" gridrow="0" pageId="3" pageNumber="331">
<taxonomicName id="AEC14D03FFD6FF821C5EFA84745E4D19" authority="Fish" authorityName="Fish" box="[1144,1329,1306,1330]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="331" phylum="Chordata" rank="species" species="hongkongensis">
<emphasis id="5BB5EA92FFD6FF821C5EFA84745E4D19" box="[1144,1329,1306,1330]" italics="true" pageId="3" pageNumber="331">M. hongkongensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="D7F134C2FFD6007E1885FADC74564D71" box="[163,1337,1346,1370]" gridrow="1" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FADC737D4D71" box="[163,530,1346,1370]" gridcol="0" gridrow="1" pageId="3" pageNumber="331">Fish samples</th>
<td id="94205DBEFFD6007E1AB5FADC728D4D71" box="[659,994,1346,1370]" gridcol="1" gridrow="1" pageId="3" pageNumber="331">3</td>
<td id="94205DBEFFD6007E1C56FADC74564D71" box="[1136,1337,1346,1370]" gridcol="2" gridrow="1" pageId="3" pageNumber="331">5</td>
</tr>
<tr id="D7F134C2FFD6007E1885FAF474564DA9" box="[163,1337,1386,1410]" gridrow="2" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FAF4737D4DA9" box="[163,530,1386,1410]" gridcol="0" gridrow="2" pageId="3" pageNumber="331">Spines of dorsal fin</th>
<td id="94205DBEFFD6007E1AB5FAF4728D4DA9" box="[659,994,1386,1410]" gridcol="1" gridrow="2" pageId="3" pageNumber="331">13</td>
<td id="94205DBEFFD6007E1C56FAF474564DA9" box="[1136,1337,1386,1410]" gridcol="2" gridrow="2" pageId="3" pageNumber="331">1314 (modally 13)</td>
</tr>
<tr id="D7F134C2FFD6007E1885FA0C74564D80" box="[163,1337,1426,1451]" gridrow="3" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FA0C737D4D80" box="[163,530,1426,1451]" gridcol="0" gridrow="3" pageId="3" pageNumber="331">Soft rays of dorsal fin</th>
<td id="94205DBEFFD6007E1AB5FA0C728D4D80" box="[659,994,1426,1451]" gridcol="1" gridrow="3" pageId="3" pageNumber="331">7</td>
<td id="94205DBEFFD6007E1C56FA0C74564D80" box="[1136,1337,1426,1451]" gridcol="2" gridrow="3" pageId="3" pageNumber="331">78 (modally 7)</td>
</tr>
<tr id="D7F134C2FFD6007E1885FA2574564DF8" box="[163,1337,1467,1491]" gridrow="4" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FA25737D4DF8" box="[163,530,1467,1491]" gridcol="0" gridrow="4" pageId="3" pageNumber="331">Spines of anal fin</th>
<td id="94205DBEFFD6007E1AB5FA25728D4DF8" box="[659,994,1467,1491]" gridcol="1" gridrow="4" pageId="3" pageNumber="331">20</td>
<td id="94205DBEFFD6007E1C56FA2574564DF8" box="[1136,1337,1467,1491]" gridcol="2" gridrow="4" pageId="3" pageNumber="331">1719 (modally 18)</td>
</tr>
<tr id="D7F134C2FFD6007E1885FA7D74564DD0" box="[163,1337,1507,1531]" gridrow="5" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885FA7D737D4DD0" box="[163,530,1507,1531]" gridcol="0" gridrow="5" pageId="3" pageNumber="331">Soft rays of anal fin</th>
<td id="94205DBEFFD6007E1AB5FA7D728D4DD0" box="[659,994,1507,1531]" gridcol="1" gridrow="5" pageId="3" pageNumber="331">10</td>
<td id="94205DBEFFD6007E1C56FA7D74564DD0" box="[1136,1337,1507,1531]" gridcol="2" gridrow="5" pageId="3" pageNumber="331">1314 (modally 14)</td>
</tr>
<tr id="D7F134C2FFD6007E1885F99574564E08" box="[163,1337,1547,1571]" gridrow="6" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885F995737D4E08" box="[163,530,1547,1571]" gridcol="0" gridrow="6" pageId="3" pageNumber="331">Vertical position of dorsal fin origin</th>
<td id="94205DBEFFD6007E1AB5F995728D4E08" box="[659,994,1547,1571]" gridcol="1" gridrow="6" pageId="3" pageNumber="331">Second to third spines of anal fin</td>
<td id="94205DBEFFD6007E1C56F99574564E08" box="[1136,1337,1547,1571]" gridcol="2" gridrow="6" pageId="3" pageNumber="331">Anal fin origin</td>
</tr>
<tr id="D7F134C2FFD6007E1885F9AD74564E60" box="[163,1337,1587,1611]" gridrow="7" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885F9AD737D4E60" box="[163,530,1587,1611]" gridcol="0" gridrow="7" pageId="3" pageNumber="331">Lateral midline scale rows</th>
<td id="94205DBEFFD6007E1AB5F9AD728D4E60" box="[659,994,1587,1611]" gridcol="1" gridrow="7" pageId="3" pageNumber="331">2829 (Modally 28)</td>
<td id="94205DBEFFD6007E1C56F9AD74564E60" box="[1136,1337,1587,1611]" gridcol="2" gridrow="7" pageId="3" pageNumber="331">3032</td>
</tr>
<tr id="D7F134C2FFD6007E1885F9C274564E5F" box="[163,1337,1628,1652]" gridrow="8" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885F9C2737D4E5F" box="[163,530,1628,1652]" gridcol="0" gridrow="8" pageId="3" pageNumber="331">Transverse scale rows</th>
<td id="94205DBEFFD6007E1AB5F9C2728D4E5F" box="[659,994,1628,1652]" gridcol="1" gridrow="8" pageId="3" pageNumber="331">14</td>
<td id="94205DBEFFD6007E1C56F9C274564E5F" box="[1136,1337,1628,1652]" gridcol="2" gridrow="8" pageId="3" pageNumber="331">12</td>
</tr>
<tr id="D7F134C2FFD6007E1885F91A74564EB7" box="[163,1337,1668,1692]" gridrow="9" pageId="3" pageNumber="331">
<th id="94205DBEFFD6007E1885F91A737D4EB7" box="[163,530,1668,1692]" gridcol="0" gridrow="9" pageId="3" pageNumber="331">Vertebrae</th>
<td id="94205DBEFFD6007E1AB5F91A728D4EB7" box="[659,994,1668,1692]" gridcol="1" gridrow="9" pageId="3" pageNumber="331">27</td>
<td id="94205DBEFFD6007E1C56F91A74564EB7" box="[1136,1337,1668,1692]" gridcol="2" gridrow="9" pageId="3" pageNumber="331">2930</td>
</tr>
</table>
</paragraph>
<paragraph id="697E3680FFD6FF8218E1F97675A24F0C" blockId="3.[151,1436,1768,1831]" pageId="3" pageNumber="331">
<emphasis id="5BB5EA92FFD6FF8218E1F97670134F29" bold="true" box="[199,380,1768,1794]" pageId="3" pageNumber="331">Distribution.—</emphasis>
So far, the new species is merely found from the southern region of
<collectingRegion id="AB05F862FFD6FF821C58F977745D4F28" box="[1150,1330,1769,1795]" country="China" name="Fujian" pageId="3" pageNumber="331">Fujian province</collectingRegion>
,
<collectingCountry id="11D67610FFD6FF821D18F97774EE4F28" box="[1342,1409,1769,1795]" name="China" pageId="3" pageNumber="331">China</collectingCountry>
at least existing in the Joulongjiang basin and Jangjiang basin of
<collectingRegion id="AB05F862FFD6FF821B75F89375674F0C" box="[851,1032,1805,1831]" country="China" name="Fujian" pageId="3" pageNumber="331">Fujian Province</collectingRegion>
, mainland
<collectingCountry id="11D67610FFD6FF821CA4F89375A64F0C" box="[1154,1225,1805,1831]" name="China" pageId="3" pageNumber="331">China</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="21DB650BFFD6FF8518B1F8E7747C496A" lastPageId="4" lastPageNumber="332" pageId="3" pageNumber="331" type="discussion">
<paragraph id="697E3680FFD6FF8218B1F8E770664FB8" blockId="3.[151,265,1913,1939]" box="[151,265,1913,1939]" pageId="3" pageNumber="331">
<emphasis id="5BB5EA92FFD6FF8218B1F8E770664FB8" bold="true" box="[151,265,1913,1939]" pageId="3" pageNumber="331">Remarks</emphasis>
</paragraph>
<paragraph id="697E3680FFD6FF8518B1F824747C496A" blockId="3.[151,1437,1978,2041]" lastBlockId="4.[151,1437,151,321]" lastPageId="4" lastPageNumber="332" pageId="3" pageNumber="331">
The allopatric, new species
<taxonomicName id="AEC14D03FFD6FF8219E9F82473814FFF" authorityName="Chen &amp; Chiu" authorityYear="2024" box="[463,750,1978,2004]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="331" phylum="Chordata" rank="species" species="minnanensis">
<emphasis id="5BB5EA92FFD6FF8219E9F82473814FFF" box="[463,750,1978,2004]" italics="true" pageId="3" pageNumber="331">Macropodus minnanensis</emphasis>
</taxonomicName>
, from
<collectingRegion id="AB05F862FFD6FF821B1FF82472814FFF" box="[825,1006,1978,2004]" country="China" name="Fujian" pageId="3" pageNumber="331">Fujian Province</collectingRegion>
,
<collectingCountry id="11D67610FFD6FF821BDFF82475534FFF" box="[1017,1084,1978,2004]" name="China" pageId="3" pageNumber="331">China</collectingCountry>
can be well distinguished from its closely related species—
<taxonomicName id="AEC14D03FFD6FF8219F1F84173CA4FD3" box="[471,677,2014,2040]" family="Osphronemidae" genus="Macropodus" kingdom="Animalia" order="Perciformes" pageId="3" pageNumber="331" phylum="Chordata" rank="species" species="hongkongensis">
<emphasis id="5BB5EA92FFD6FF8219F1F84173CA4FD3" box="[471,677,2014,2040]" italics="true" pageId="3" pageNumber="331">M. hongkongensis</emphasis>
</taxonomicName>
from
<collectingCountry id="11D67610FFD6FF821AC9F84172044FD2" box="[751,875,2015,2041]" name="Hong Kong" pageId="3" pageNumber="331">HongKong</collectingCountry>
by the following features: (1) anal fin soft rays: modally 10 vs. modally 14; (3) vertical insertion of dorsal fin origin: second to third soft rays of anal fin vs. anal fin origin; (4) longitudinal scale rows in midline: modally 28 vs. 3032; (5) transverse scales: 14 vs. 12; (6) vertebral count: 27 vs. 2930; and (7) specific coloration: eye greyish in male vs red in male; opecular ocellus spot deep black in male vs. brownish green in male. Since the rare fish is not popular even in the
<typeStatus id="B67A8822FFD1FF851C28FE9D75514936" box="[1038,1086,259,285]" pageId="4" pageNumber="332">type</typeStatus>
locality, the detailed ecological preference and conservation issue would be very necessary to concerned and conducted in near future.
</paragraph>
</subSubSection>
</treatment>
</document>