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<document id="CA5700436ECCF5487648222DF6475209" ID-DOI="10.11646/zootaxa.5501.3.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="13628197" ID-ZooBank="F6A55A77-C4C8-4BDD-8D36-482395A01311" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="karina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="karina" checkinTime="1725278975848" checkinUser="plazi" docAuthor="Nagel, Peter" docDate="2024" docId="03F01B1CFFF56D6FA0EE5FE413C8FC55" docLanguage="en" docName="zootaxa.5501.3.3.pdf" docOrigin="Zootaxa 5501 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5501.3.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paussus haucki Bednarik &amp; Bocak 2023" docType="treatment" docVersion="3" lastPageNumber="453" masterDocId="FFC96364FFF36D69A0795F731654FFC5" masterDocTitle="The Paussus (Scaphipaussus) lanxangensis species group of Indomalayan ant nest beetles (Coleoptera, Carabidae, Paussinae, Paussini) with description of a new species" masterLastPageNumber="457" masterPageNumber="447" pageNumber="453" updateTime="1725501495294" updateUser="karina" zenodo-license-document="CLOSED">
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<mods:title id="73D80E9FC44E85F6C2CCAB9920EE91BE">The Paussus (Scaphipaussus) lanxangensis species group of Indomalayan ant nest beetles (Coleoptera, Carabidae, Paussinae, Paussini) with description of a new species</mods:title>
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<treatment id="03F01B1CFFF56D6FA0EE5FE413C8FC55" LSID="urn:lsid:plazi:treatment:03F01B1CFFF56D6FA0EE5FE413C8FC55" httpUri="http://treatment.plazi.org/id/03F01B1CFFF56D6FA0EE5FE413C8FC55" lastPageNumber="453" pageId="6" pageNumber="453">
<subSubSection id="C343F981FFF56D6FA0EE5FE41521FF74" box="[151,885,151,178]" pageId="6" pageNumber="453" type="nomenclature">
<paragraph id="8BE6AA0AFFF56D6FA0EE5FE41521FF74" blockId="6.[151,885,151,178]" box="[151,885,151,178]" pageId="6" pageNumber="453">
<heading id="D0AE1D66FFF56D6FA0EE5FE41521FF74" bold="true" box="[151,885,151,178]" fontSize="11" level="1" pageId="6" pageNumber="453" reason="1">
<emphasis id="B92D7618FFF56D6FA0EE5FE41521FF74" bold="true" box="[151,885,151,178]" pageId="6" pageNumber="453">
<taxonomicName id="4C59D189FFF56D6FA0EE5FE414C5FF74" authority="Bednarik &amp; Bocak, 2023" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[151,657,151,178]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA0EE5FE41701FF77" bold="true" box="[151,341,151,178]" italics="true" pageId="6" pageNumber="453">Paussus haucki</emphasis>
<bibRefCitation id="EFC8D7FBFFF56D6FA1255FE414C5FF74" author="Bednarik, M. &amp; Bocak, L." box="[348,657,151,177]" pageId="6" pageNumber="453" pagination="1 - 30" refId="ref6548" refString="Bednarik, M. &amp; Bocak, L. (2023) New species of Paussus, Subgenus Scaphipaussus (Coleoptera: Carabidae: Paussinae), from Southeast Asia Reveal Ambiguities in Species Group Limits and High Species Diversity in the Oriental Region. Insects, 14 (947), 1 - 30. https: // doi. org / 10.3390 / insects 14120947" type="journal article" year="2023">Bednařik &amp; Bocak, 2023</bibRefCitation>
</taxonomicName>
[DP
<date id="FFE78CCAFFF56D6FA2B75FE4153AFF74" box="[718,878,151,177]" pageId="6" pageNumber="453" value="2023-12-14">14 Dec. 2023</date>
]
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C343F981FFF56D6FA0EE5FAF12FCFF36" box="[151,1192,220,244]" pageId="6" pageNumber="453" type="description">
<paragraph id="8BE6AA0AFFF56D6FA0EE5FAF12FCFF36" blockId="6.[151,1315,220,336]" box="[151,1192,220,244]" pageId="6" pageNumber="453">
Female
<typeStatus id="54E214A8FFF56D6FA0915FAF1715FF31" box="[232,321,220,244]" pageId="6" pageNumber="453" type="holotype">holotype</typeStatus>
specimen only. The original authors classify the species as part of Sg.
<taxonomicName id="4C59D189FFF56D6FA4745FAF12F6FF31" authorityName="Fowler" authorityYear="1912" box="[1037,1186,220,244]" pageId="6" pageNumber="456" rank="subGenus" subGenus="Scaphipaussus">
<emphasis id="B92D7618FFF56D6FA4745FAF12F6FF31" box="[1037,1186,220,244]" italics="true" pageId="6" pageNumber="453">Scaphipaussus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C343F981FFF56D6FA0EE5F8813C8FC55" pageId="6" pageNumber="453" type="materials_examined">
<paragraph id="8BE6AA0AFFF56D6FA0EE5F881701FEF4" blockId="6.[151,1315,220,336]" pageId="6" pageNumber="453">
<materialsCitation id="3B31A057FFF56D6FA0EE5F881705FEF4" collectingDate="1999-04-06" collectingDateMax="1999-04-09" collectingDateMin="1999-04-06" collectionCode="MBC" collectorName="Phu Ruea N. P. &amp; D. Hauck" country="Thailand" elevation="1100" latitude="17.5" location="Loei prov." longLatPrecision="1278" longitude="101.35" pageId="6" pageNumber="453" specimenCount="1" stateProvince="Loei prov.">
Type locality: NE
<collectingCountry id="F34EEA9AFFF56D6FA1285F8917FAFED7" box="[337,430,250,274]" name="Thailand" pageId="6" pageNumber="453">Thailand</collectingCountry>
,
<collectingRegion id="499D64E8FFF56D6FA1C15F891477FED7" box="[440,547,250,274]" country="Thailand" name="Loei" pageId="6" pageNumber="453">Loei prov.</collectingRegion>
,
<collectorName id="26ACCFDCFFF56D6FA2555F891495FED7" box="[556,705,250,274]" pageId="6" pageNumber="453">Phu Ruea N.P.</collectorName>
,
<geoCoordinate id="EE6DCCCDFFF56D6FA2B25F891577FED7" box="[715,803,250,274]" degrees="17" direction="north" minutes="30" orientation="latitude" pageId="6" pageNumber="453" precision="925" value="17.5">17°30N</geoCoordinate>
<geoCoordinate id="EE6DCCCDFFF56D6FA3505F8915D8FED7" box="[809,908,250,274]" degrees="101" direction="east" minutes="21" orientation="longitude" pageId="6" pageNumber="453" precision="925" value="101.35">101°21E</geoCoordinate>
,
<quantity id="4CA107EFFFF56D6FA3EE5F8915B1FED7" box="[919,997,250,274]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="453" unit="m" value="1100.0">
<elevation id="00744D39FFF56D6FA3EE5F8915B1FED7" box="[919,997,250,274]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="6" pageNumber="453" unit="m" value="1100.0">1100 m</elevation>
</quantity>
,
<date id="FFE78CCAFFF56D6FA3965F8912DFFED7" box="[1007,1163,250,274]" pageId="6" pageNumber="453" value="1999-04-06" valueMax="1999-04-09" valueMin="1999-04-06">
<collectingDate id="EFA37522FFF56D6FA3965F8912DFFED7" box="[1007,1163,250,274]" pageId="6" pageNumber="453" value="1999-04-06" valueMax="1999-04-09" valueMin="1999-04-06">69 April 1999</collectingDate>
</date>
,
<collectorName id="26ACCFDCFFF56D6FA4EF5F8812A2FED7" box="[1174,1270,250,274]" pageId="6" pageNumber="453">D. Hauck</collectorName>
leg. Depository:
<collectionCode id="ED4832CFFFF56D6FA16C5E6A1705FEF4" box="[277,337,281,305]" pageId="6" pageNumber="453">MBC</collectionCode>
</materialsCitation>
.
</paragraph>
<paragraph id="8BE6AA0AFFF56D6FA0EE5E4B1746FE8A" blockId="6.[151,1315,220,336]" box="[151,274,312,336]" pageId="6" pageNumber="453">
BL
<quantity id="4CA107EFFFF56D6FA0C45E4B175AFE8A" box="[189,270,312,336]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="6" pageNumber="453" unit="mm" value="4.2">4.2 mm</quantity>
.
</paragraph>
<paragraph id="8BE6AA0AFFF56D6FA0EE5E0D174EFDC1" blockId="6.[151,1437,382,912]" pageId="6" pageNumber="453">
Although there is no specimen of either
<taxonomicName id="4C59D189FFF56D6FA20F5E0C155AFE5D" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[630,782,382,408]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">
<emphasis id="B92D7618FFF56D6FA20F5E0C155AFE5D" box="[630,782,382,408]" italics="true" pageId="6" pageNumber="453">P. wakaharai</emphasis>
</taxonomicName>
or
<taxonomicName id="4C59D189FFF56D6FA3425E0C15F2FE5D" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[827,934,382,408]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA3425E0C15F2FE5D" box="[827,934,382,408]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
available to the present study, the original descriptions and illustrations reveal just weak and possibly non-essential differences between the
<specimenCount id="9D5F6183FFF56D6FA57F5ED1175BFE25" count="2" pageId="6" pageNumber="453" type="generic" typeStatus="holotype">two holotype specimens</specimenCount>
.
<taxonomicName id="4C59D189FFF56D6FA1625EB417A3FE25" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[283,503,454,480]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">
<emphasis id="B92D7618FFF56D6FA1625EB417A3FE25" box="[283,503,454,480]" italics="true" pageId="6" pageNumber="453">Paussus wakaharai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C59D189FFF56D6FA2565EB414C3FE25" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[559,663,454,480]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA2565EB414C3FE25" box="[559,663,454,480]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
were published almost simultaneously with the latter published half a year later.
</paragraph>
<paragraph id="8BE6AA0AFFF56D6FA0BE5D7D1211FD89" blockId="6.[151,1437,382,912]" pageId="6" pageNumber="453">
The original description of
<taxonomicName id="4C59D189FFF56D6FA2785D7C143DFDED" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[513,617,526,552]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA2785D7C143DFDED" box="[513,617,526,552]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
does not include information on the mouthparts. The description of the pygidium indicates that the marginal semicircular trichome is most probably absent.
</paragraph>
<paragraph id="8BE6AA0AFFF56D6FA0BE5D251407FCA9" blockId="6.[151,1437,382,912]" pageId="6" pageNumber="453">
According to the original descriptions there seem to be possibly gradual differences between
<taxonomicName id="4C59D189FFF56D6FA4A35D241339FDB5" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[1242,1389,598,624]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">
<emphasis id="B92D7618FFF56D6FA4A35D241339FDB5" box="[1242,1389,598,624]" italics="true" pageId="6" pageNumber="453">P. wakaharai</emphasis>
</taxonomicName>
and
<taxonomicName id="4C59D189FFF56D6FA0EE5D0816ABFD51" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[151,255,634,660]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA0EE5D0816ABFD51" box="[151,255,634,660]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
: Antennal club with posterior excavation absent in
<taxonomicName id="4C59D189FFF56D6FA3375D0815B7FD51" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[846,995,634,660]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">
<emphasis id="B92D7618FFF56D6FA3375D0815B7FD51" box="[846,995,634,660]" italics="true" pageId="6" pageNumber="453">P. wakaharai</emphasis>
</taxonomicName>
and vertex with large medial tubercle. In
<taxonomicName id="4C59D189FFF56D6FA0CE5DEC174BFD7D" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[183,287,670,696]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA0CE5DEC174BFD7D" box="[183,287,670,696]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
the antennal club shows a “posterolateral cavity” and vertex “with pair of curved ... vertical shields” (
<bibRefCitation id="EFC8D7FBFFF56D6FA0E65DB117EEFD19" author="Bednarik, M. &amp; Bocak, L." box="[159,442,706,732]" pageId="6" pageNumber="453" pagination="1 - 30" refId="ref6548" refString="Bednarik, M. &amp; Bocak, L. (2023) New species of Paussus, Subgenus Scaphipaussus (Coleoptera: Carabidae: Paussinae), from Southeast Asia Reveal Ambiguities in Species Group Limits and High Species Diversity in the Oriental Region. Insects, 14 (947), 1 - 30. https: // doi. org / 10.3390 / insects 14120947" type="journal article" year="2023">Bednařik &amp; Bocak 2023</bibRefCitation>
, pp. 21, 22). The possible presence of the latter character in
<emphasis id="B92D7618FFF56D6FA40D5DB01371FD19" box="[1140,1317,706,732]" italics="true" pageId="6" pageNumber="453">
<taxonomicName id="4C59D189FFF56D6FA40D5DB0135DFD19" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[1140,1289,706,732]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">P. wakaharai</taxonomicName>
is
</emphasis>
suggested by the description and illustration (
<bibRefCitation id="EFC8D7FBFFF56D6FA24A5D9514A2FCC5" author="Maruyama, M. &amp; Nakase, Y." box="[563,758,742,768]" pageId="6" pageNumber="453" pagination="1 - 3" refId="ref7360" refString="Maruyama, M. &amp; Nakase, Y. (2023) A New Species of the Genus Paussus Linne, 1775 (Coleoptera: Carabidae: Paussinae) from Laos, Allied to P. lanxangensis Nagel, 2009. Japanese Journal of Systematic Entomology, 29 (1), 1 - 3." type="journal article" year="2023">Maruyama 2023</bibRefCitation>
). The probably shrunk antennal club of
<emphasis id="B92D7618FFF56D6FA4A95D94133BFCC5" box="[1232,1391,742,768]" italics="true" pageId="6" pageNumber="453">
<taxonomicName id="4C59D189FFF56D6FA4A95D94133FFCC5" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[1232,1387,742,768]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">P. wakaharai</taxonomicName>
,
</emphasis>
the non-teneral condition of
<taxonomicName id="4C59D189FFF56D6FA1D65C781443FCE1" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[431,535,778,804]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA1D65C781443FCE1" box="[431,535,778,804]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
and a possible sexual dimorphism (female
<taxonomicName id="4C59D189FFF56D6FA47B5C78123EFCE1" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[1026,1130,778,804]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA47B5C78123EFCE1" box="[1026,1130,778,804]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
versus male
<taxonomicName id="4C59D189FFF56D6FA4865C7813C7FCE1" authorityName="Maruyama &amp; Nakase" authorityYear="2023" box="[1279,1427,778,804]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">
<emphasis id="B92D7618FFF56D6FA4865C7813C7FCE1" box="[1279,1427,778,804]" italics="true" pageId="6" pageNumber="453">P. wakaharai</emphasis>
</taxonomicName>
) might sufficiently explain the differences. Consequently,
<taxonomicName id="4C59D189FFF56D6FA35B5C5C15DEFC8D" authorityName="Bednarik &amp; Bocak" authorityYear="2023" box="[802,906,814,840]" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="haucki">
<emphasis id="B92D7618FFF56D6FA35B5C5C15DEFC8D" box="[802,906,814,840]" italics="true" pageId="6" pageNumber="453">P. haucki</emphasis>
</taxonomicName>
might turn out to be a younger synonym of
<taxonomicName id="4C59D189FFF56D6FA5F15C5C141BFCA9" authority="Maruyama &amp; Nakase, 2023" authorityName="Maruyama &amp; Nakase" authorityYear="2023" class="Insecta" family="Carabidae" genus="Paussus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="453" phylum="Arthropoda" rank="species" species="wakaharai">
<emphasis id="B92D7618FFF56D6FA5F15C5C175BFCA9" italics="true" pageId="6" pageNumber="453">P. wakaharai</emphasis>
<bibRefCitation id="EFC8D7FBFFF56D6FA16F5C21141BFCA9" author="Maruyama, M. &amp; Nakase, Y." box="[278,591,850,876]" pageId="6" pageNumber="453" pagination="1 - 3" refId="ref7360" refString="Maruyama, M. &amp; Nakase, Y. (2023) A New Species of the Genus Paussus Linne, 1775 (Coleoptera: Carabidae: Paussinae) from Laos, Allied to P. lanxangensis Nagel, 2009. Japanese Journal of Systematic Entomology, 29 (1), 1 - 3." type="journal article" year="2023">Maruyama &amp; Nakase, 2023</bibRefCitation>
</taxonomicName>
.
</paragraph>
<paragraph id="8BE6AA0AFFF56D6FA0BE5C0513C8FC55" blockId="6.[151,1437,382,912]" box="[199,1436,886,912]" pageId="6" pageNumber="453">Future findings of non-teneral specimens each with the alternative sex will help to clarify the factual situation.</paragraph>
</subSubSection>
</treatment>
</document>