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<mods:title id="660F11DEF23AB6CAF55DA026E6573B54">Early Coal Swamp Vegetation From The Serpukhovian Lower Clackmannan Group Of Scotland</mods:title>
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<mods:affiliation id="C7C5D019061C7E0246E6FE52E299384F">Department of Earth Sciences, University of Bristol, Bristol, BS 8 1 TQ, United Kingdom; email: christopher. cleal @ bristol. ac. uk.</mods:affiliation>
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<mods:namePart id="07CA342E96DCCA6EA9F375858EBA7DB4">Strullu-Derrien, Christine</mods:namePart>
<mods:affiliation id="C1ED12248CC68B59A868DA7655D56586">Institut de Systématique, Évolution, Biodiversité (UMR 7205), Muséum National dHistoire Naturelle, CNRS, Sorbonne Université, Paris, France. &amp; Science Group, The Natural History Museum, London, United Kingdom.</mods:affiliation>
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<mods:namePart id="92D56BD0E41CAF432C1DE6DF48509E57">Spencer, Alan R. T.</mods:namePart>
<mods:affiliation id="D18C84D65ED5F39F9CE1BFE787F38AAB">Department of Earth Sciences, The Natural History Museum, London, United Kingdom. &amp; Department of Earth Science and Engineering, Imperial College London, London, United Kingdom.</mods:affiliation>
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<taxonomicName id="4C084D29B033FFAA77F6FB455476FB08" authority="ETTINGSH. ex HELMHACKER, 1874" authorityName="ETTINGSH. ex HELMHACKER" authorityYear="1874" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="tenerrimum">
<emphasis id="B97CEAB8B033FFAA77F6FB455476FB08" bold="true" pageId="10" pageNumber="45">
<emphasis id="B97CEAB8B033FFAA77F6FB45546EFB69" bold="true" box="[235,545,1170,1194]" italics="true" pageId="10" pageNumber="45">Sphenophyllum tenerrimum</emphasis>
<smallCapsWord id="8D51A076B033FFAA7535FB4554C1FB6A" baselines="1188,1189" box="[552,654,1170,1194]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Ettingsh" pageId="10" pageNumber="45">ETTINGSH</smallCapsWord>
. ex
<bibRefCitation id="EF994B5BB033FFAA767BFB6457B8FB0A" author="Helmhacker, R." box="[358,503,1203,1227]" pageId="10" pageNumber="45" pagination="23 - 97" refId="ref19864" refString="Helmhacker, R. (1874): Einige Beitrag zur Kenntnis der Flora des Sudrandes der oberschlesisch-polnischen Karbons. - Berg-und Huttenmannisches Jahrbuch der k. k. Bergakademien zu Leoben und Pribram, und der koniglich ungarischen Bergakademie zu Schemitz, 22: 23 - 97." type="journal article" year="1874">
<smallCapsWord id="8D51A076B033FFAA767BFB6457B8FB0A" baselines="1221,1221" box="[358,503,1203,1227]" lowerCaseFontSize="7" mainFontSize="10" normCase="title" normString="Helmhacker" pageId="10" pageNumber="45">HELMHACKER</smallCapsWord>
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, 1874
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</paragraph>
<paragraph id="8BB736AAB033FFAA765FFB045471FB28" blockId="10.[322,574,1235,1292]" box="[322,574,1235,1259]" pageId="10" pageNumber="45">
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emend. by
<bibRefCitation id="EF994B5BB033FFAA76A3FB045471FB28" author="Stur, D." box="[446,574,1235,1259]" pageId="10" pageNumber="45" pagination="107 - 472" refId="ref24329" refString="Stur, D. (1877): Die Culm-Flora der Ostrauer und Waldenburger Schichten. - Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 8 (2): 107 - 472." type="journal article" year="1877">Stur (1877)</bibRefCitation>
</emphasis>
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<figureCitation id="13332A2FB033FFAA769EFB2357B2FACF" box="[387,509,1268,1292]" captionStart="Text-fig" captionStartId="11.[125,201,1966,1987]" captionTargetBox="[125,1440,151,1950]" captionTargetId="figure-6@11.[123,783,624,1098]" captionTargetPageId="11" captionText="Text-fig.6.a:SphenophyllumtenerrimumETTINGSH. exHELMHACKER;RobroystonColliery,2.8kmSEofBishopbriggs,Lanarkshire; ca.10m below Orchard Limestone, Upper Limestone Formation; BGS Kidst.156. b, c: Pecopteris aspera BRONGN.; Cuthill Shore (near highwater mark) between Musselburgh and Prestonpans, Haddingtonshire; between Nos 4 and 5 Limestones, Upper Limestone Formation; BGS Kidst.5415 (b), BGS Kidst.4517 (c). d: Zeilleria moravica (ETTINGSH.) BUREAU; Thorneycroft No. 1 Borehole, 0.5 km NE of Cumbernauld Railway Station, Dumbartonshire; 389 m depth, Limestone Coal Formation; BGS Kidst.2420. Scale bars = 5 mm (a, b), 10 mm (c, d)." figureDoi="http://doi.org/10.5281/zenodo.14722403" httpUri="https://zenodo.org/record/14722403/files/figure.png" pageId="10" pageNumber="45">Text-fig. 6a</figureCitation>
</paragraph>
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1874
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<emphasis id="B97CEAB8B033FFAA77D1FAF55790FAFB" box="[204,479,1314,1336]" italics="true" pageId="10" pageNumber="45">Sphenophyllum tenerrimum</emphasis>
<smallCapsWord id="8D51A076B033FFAA76F3FAF4540AFAF5" baselines="1331,1330" box="[494,581,1315,1336]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Ettingsh" pageId="10" pageNumber="45">ETTINGSH</smallCapsWord>
. ex
<smallCapsWord id="8D51A076B033FFAA7562FAF454B5FAF5" baselines="1331,1330" box="[639,762,1315,1336]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Helmhacker" pageId="10" pageNumber="45">HELMHACKER</smallCapsWord>
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, p. 50, pl. 3, figs 516.
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1877
<taxonomicName id="4C084D29B033FFAA77D1FA8954B6FAB2" authority="ETTINGSH. ex HELMHACKER" authorityName="ETTINGSH. ex HELMHACKER" authorityYear="1874" box="[204,761,1374,1396]" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="tenerrimum">
<emphasis id="B97CEAB8B033FFAA77D1FA895790FAB7" box="[204,479,1374,1396]" italics="true" pageId="10" pageNumber="45">Sphenophyllum tenerrimum</emphasis>
<smallCapsWord id="8D51A076B033FFAA76F3FA89540BFAB2" baselines="1390,1389" box="[494,580,1374,1395]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Ettingsh" pageId="10" pageNumber="45">ETTINGSH</smallCapsWord>
. ex
<smallCapsWord id="8D51A076B033FFAA7562FA8954B6FAB2" baselines="1390,1389" box="[639,761,1374,1395]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Helmhacker" pageId="10" pageNumber="45">HELMHACKER</smallCapsWord>
</taxonomicName>
; Stur, p. 108, pl. 7, figs 114. (?Ettingshausens
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)
</paragraph>
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1969
<taxonomicName id="4C084D29B033FFAA77D1FA4E54B6FA6E" authority="ETTINGSH. ex HELMHACKER" authorityName="ETTINGSH. ex HELMHACKER" authorityYear="1874" box="[204,761,1433,1455]" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="tenerrimum">
<emphasis id="B97CEAB8B033FFAA77D1FA4E5790FA6C" box="[204,479,1433,1455]" italics="true" pageId="10" pageNumber="45">Sphenophyllum tenerrimum</emphasis>
<smallCapsWord id="8D51A076B033FFAA76F3FA4D540BFA6E" baselines="1450,1449" box="[494,580,1434,1455]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Ettingsh" pageId="10" pageNumber="45">ETTINGSH</smallCapsWord>
. ex
<smallCapsWord id="8D51A076B033FFAA7562FA4D54B6FA6E" baselines="1450,1449" box="[639,761,1434,1455]" lowerCaseFontSize="6" mainFontSize="9" normCase="title" normString="Helmhacker" pageId="10" pageNumber="45">HELMHACKER</smallCapsWord>
</taxonomicName>
; Crookall, p. 602, pl. 108, fig. 6.
</paragraph>
<paragraph id="8BB736AAB033FFAA77BBFA3254A3F903" blockId="10.[129,767,1509,1728]" pageId="10" pageNumber="45">
D e s c r i p t i o n. Ribbed stems with whorls of 912 leaves borne at about right-angles in proximal part of stem, but more obliquely in more distal positions. Leaves
<quantity id="4CF09B4FB033FFAA779CF99156ACF99D" box="[129,227,1606,1630]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="10" pageNumber="45" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">310 mm</quantity>
long, deeply incised into 24 narrow-linear, splayed segments, the incisions usually extending about a half of the leaf length, each segment with a blunt apex. Occasionally the leaf segments divide for a second time. Leaf surface striate.
</paragraph>
<paragraph id="8BB736AAB033FFAA77BBF90F52CCFE93" blockId="10.[129,768,1751,2100]" lastBlockId="10.[843,1482,152,624]" pageId="10" pageNumber="45">
R e m a r k s. This is the most commonly-reported
<taxonomicName id="4C084D29B033FFAA779CF92F573BF8D3" box="[129,372,1784,1808]" pageId="10" pageNumber="45">
<emphasis id="B97CEAB8B033FFAA779CF92F576BF8D3" box="[129,292,1784,1808]" italics="true" pageId="10" pageNumber="45">Sphenophyllum</emphasis>
species
</taxonomicName>
found in lower Namurian macrofloras and is easily recognised by its whorls of deeply-incised leaves with splayed segments (e.g.,
<bibRefCitation id="EF994B5BB033FFAA750CF8EE54D7F892" author="Storch, D." box="[529,664,1849,1873]" pageId="10" pageNumber="45" pagination="171 - 273" refId="ref24072" refString="Storch, D. (1980): Sphenophyllum - Arten aus drei intramontanen Karbonbecken - pflanzengeographische Beson- derheiten im mitteleuropaischen Karbon. - Schriftenreihe der geologische Wissenschaften, 16: 171 - 273." type="journal article" year="1980">Storch 1980</bibRefCitation>
,
<bibRefCitation id="EF994B5BB033FFAA75BAF8EE56F7F8B2" author="Havlena, V." pageId="10" pageNumber="45" pagination="1 - 22" refId="ref19777" refString="Havlena, V. (1982): Sphenophyllum tenerrimum and other species of Sphenophyllum in the Namurian of the Ostrava-Karvina coalfield (Upper Silesian basin). - Palaeon- tographica, Abt. B, 184: 1 - 22." type="journal article" year="1982">Havlena 1982</bibRefCitation>
,
<bibRefCitation id="EF994B5BB033FFAA77DBF88D5456F8B2" author="Terreaux de Felice, H. &amp; Decombeix, A. - L. &amp; Galtier, J." box="[198,537,1881,1905]" pageId="10" pageNumber="45" pagination="587 - 599" refId="ref24361" refString="Terreaux de Felice, H., Decombeix, A. - L., Galtier, J. (2019): Anatomy, affinities, and evolutionary implications of new silicified stems of Sphenophyllum Brongniart, 1828 from the early Carboniferous (Mississippian) of France and Germany. - Geodiversitas, 41: 587 - 599. https: // doi. org / 10.5252 / geodiversitas 2019 v 41 a 14" type="journal article" year="2019">Terreaux de Felice et al. 2019</bibRefCitation>
). However, there are nomenclatural issues with the species epithet. According to
<bibRefCitation id="EF994B5BB033FFAA779CF84D56B5F871" author="Stur, D." box="[129,250,1946,1970]" pageId="10" pageNumber="45" pagination="107 - 472" refId="ref24329" refString="Stur, D. (1877): Die Culm-Flora der Ostrauer und Waldenburger Schichten. - Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 8 (2): 107 - 472." type="journal article" year="1877">Stur (1877)</bibRefCitation>
it was first used in manuscript but not published by Ettingshausen in 1853, and was mentioned by Stur (in
<bibRefCitation id="EF994B5BB033FFAA779CF80C570EF830" author="Foetterle, F." box="[129,321,2011,2035]" pageId="10" pageNumber="45" pagination="51 - 54" refId="ref18582" refString="Foetterle, F. (1868): Die Lagerungs Verhaltnisse der Steinkohlen-flotze in der Ostrauer Steinkohlenmulde. - Ver- handlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 1868: 51 - 54." type="journal article" year="1868">Foetterle 1868: 50</bibRefCitation>
) in a species list but without description or illustration. The name
<taxonomicName id="4C084D29B033FFAA7676F82B57B0F7D0" authorityName="ETTINGSH. ex HELMHACKER" authorityYear="1874" box="[363,511,2044,2067]" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="tenerrimum">
<emphasis id="B97CEAB8B033FFAA7676F82B57B0F7D0" box="[363,511,2044,2067]" italics="true" pageId="10" pageNumber="45">S. tenerrimum</emphasis>
</taxonomicName>
was first validly used by
<bibRefCitation id="EF994B5BB033FFAA779CF7CB5702F7F7" author="Helmhacker, R." box="[129,333,2076,2100]" pageId="10" pageNumber="45" pagination="23 - 97" refId="ref19864" refString="Helmhacker, R. (1874): Einige Beitrag zur Kenntnis der Flora des Sudrandes der oberschlesisch-polnischen Karbons. - Berg-und Huttenmannisches Jahrbuch der k. k. Bergakademien zu Leoben und Pribram, und der koniglich ungarischen Bergakademie zu Schemitz, 22: 23 - 97." type="journal article" year="1874">Helmhacker (1874)</bibRefCitation>
who included a description and illustrated several specimens, although most of those specimens are now reported lost (
<bibRefCitation id="EF994B5BB033FFAA74F5FF6F52D8FF13" author="Jongmans, W. J." box="[1000,1175,184,208]" pageId="10" pageNumber="45" refId="ref20564" refString="Jongmans, W. J. (1911): Anleitung zur bestimmung der Karbonpflanzen West-Europas, mit besonderer berucksichtigung der in den Niederlanden und den benachbarten Landern gefundenen oder noch zu erwartenden Arten. I Band. Thallophytae, Equisetales, Sphenophyllales (Mededeelingen van de Rijksopsporing van Delfstoffen, No. 3). - Kommissionsverlag von Craz &amp; Gerlach, Freiberg in Sachsen, 482 pp. https: // doi. org / 10.5962 / bhl. title. 83723" type="book" year="1911">Jongmans 1911</bibRefCitation>
). Nevertheless, Helmhacker made clear reference to Ettingshausens specimens, which were later illustrated by
<bibRefCitation id="EF994B5BB033FFAA7340FF2F5292FED3" author="Stur, D." box="[1117,1245,248,272]" pageId="10" pageNumber="45" pagination="107 - 472" refId="ref24329" refString="Stur, D. (1877): Die Culm-Flora der Ostrauer und Waldenburger Schichten. - Abhandlungen der Kaiserlich-Koniglichen Geologischen Reichsanstalt, 8 (2): 107 - 472." type="journal article" year="1877">Stur (1877)</bibRefCitation>
, and so these can be legitimately taken as
<typeStatus id="54B38808B033FFAA7328FECF52DAFEF3" box="[1077,1173,280,304]" pageId="10" pageNumber="45" type="syntype">syntypes</typeStatus>
; Helmhakers publication of the name was therefore valid.
</paragraph>
<paragraph id="8BB736AAB033FFAA746CFE8F5226FE33" blockId="10.[843,1482,152,624]" pageId="10" pageNumber="45">
An added complication is that
<bibRefCitation id="EF994B5BB033FFAA73DBFE8F53D5FEB3" author="Helmhacker, R." box="[1222,1434,344,368]" pageId="10" pageNumber="45" pagination="43 - 44" refId="ref19819" refString="Helmhacker, R. (1873): O geologickem rozsireni rodu Sphenophyllum [On the geological distribution of the genus Sphenophyllum]. - Zpravy o zasedani Kralovske ceske spolecnosti nauk v Praze, 1872: 43 - 44. (in Czech)" type="journal article" year="1873">Helmhacker (1873)</bibRefCitation>
had earlier referred to the specimens he figured in his 1874a paper as
<taxonomicName id="4C084D29B033FFAA74B5FE4F53D6FE6C" authority="HELMACKER. However" authorityName="HELMACKER. However" box="[936,1433,408,432]" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="binatum">
<emphasis id="B97CEAB8B033FFAA74B5FE4F52E6FE73" box="[936,1193,408,432]" italics="true" pageId="10" pageNumber="45">Sphenophyllum binatum</emphasis>
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. However
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, this was a nomen nudum with no accompanying description or illustration, and so invalid.
</paragraph>
<paragraph id="8BB736AAB033FFAA746CFE2F53F8FDB3" blockId="10.[843,1482,152,624]" pageId="10" pageNumber="45">
This species is very similar to the upper Carboniferous
<taxonomicName id="4C084D29B033FFAA7456FDCF5299FDEE" authority="STUR" authorityName="STUR" box="[843,1238,536,560]" class="Polypodiopsida" family="Sphenophyllaceae" genus="Sphenophyllum" higherTaxonomySource="GBIF" kingdom="Plantae" order="Sphenophyllales" pageId="10" pageNumber="45" phylum="Tracheophyta" rank="species" species="trichomatosum">
<emphasis id="B97CEAB8B033FFAA7456FDCF52D6FDF3" box="[843,1177,536,560]" italics="true" pageId="10" pageNumber="45">Sphenophyllum trichomatosum</emphasis>
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, but the leaves of the latter have segments with a more acute apex and the leaf surface tends to be smooth (non-striate) (
<bibRefCitation id="EF994B5BB033FFAA721DFD8F53E4FDB3" author="Jongmans, W. J." box="[1280,1451,600,624]" pageId="10" pageNumber="45" refId="ref20564" refString="Jongmans, W. J. (1911): Anleitung zur bestimmung der Karbonpflanzen West-Europas, mit besonderer berucksichtigung der in den Niederlanden und den benachbarten Landern gefundenen oder noch zu erwartenden Arten. I Band. Thallophytae, Equisetales, Sphenophyllales (Mededeelingen van de Rijksopsporing van Delfstoffen, No. 3). - Kommissionsverlag von Craz &amp; Gerlach, Freiberg in Sachsen, 482 pp. https: // doi. org / 10.5962 / bhl. title. 83723" type="book" year="1911">Jongmans 1911</bibRefCitation>
).
</paragraph>
<paragraph id="8BB736AAB033FFAA746CFD50559DFD1C" blockId="10.[843,1482,647,735]" pageId="10" pageNumber="45">
O c c u r r e n c e. Ca.
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below Orchard Limestone, (Upper Limestone Formation), Robroyston Colliery, Lanarkshire.
</paragraph>
</subSubSection>
</treatment>
</document>